English Lowlands beech forests
Encyclopedia
The term English Lowlands beech forests refers to a terrestrial ecoregion
Ecoregion
An ecoregion , sometimes called a bioregion, is an ecologically and geographically defined area that is smaller than an ecozone and larger than an ecosystem. Ecoregions cover relatively large areas of land or water, and contain characteristic, geographically distinct assemblages of natural...

 in Northern Europe
Northern Europe
Northern Europe is the northern part or region of Europe. Northern Europe typically refers to the seven countries in the northern part of the European subcontinent which includes Denmark, Estonia, Latvia, Lithuania, Norway, Finland and Sweden...

, as defined by the World Wide Fund for Nature
World Wide Fund for Nature
The World Wide Fund for Nature is an international non-governmental organization working on issues regarding the conservation, research and restoration of the environment, formerly named the World Wildlife Fund, which remains its official name in Canada and the United States...

 (WWF) and the European Environment Agency
European Environment Agency
European Environment Agency is an agency of the European Union. Its task is to provide sound, independent information on the environment. It is a major information source for those involved in developing, adopting, implementing and evaluating environmental policy, and also the general public...

 (EEA). Part of the Temperate broadleaf and mixed forests
Temperate broadleaf and mixed forests
Mixed forests are a temperate and humid biome. The typical structure of these forests includes four layers. The uppermost layer is the canopy composed of tall mature trees ranging from 33 to 66 m high. Below the canopy is the three-layered, shade-tolerant understory that is roughly 9 to...

 biome
Biome
Biomes are climatically and geographically defined as similar climatic conditions on the Earth, such as communities of plants, animals, and soil organisms, and are often referred to as ecosystems. Some parts of the earth have more or less the same kind of abiotic and biotic factors spread over a...

 in the Palaearctic ecozone
Ecozone
An ecozone is the broadest biogeographic division of the Earth's land surface, based on distributional patterns of terrestrial organisms.Ecozones delineate large areas of the Earth's surface within which organisms have been evolving in relative isolation over long periods of time, separated from...

, it covers 45600 km² (17,606.3 sq mi) of Southern England
Southern England
Southern England, the South and the South of England are imprecise terms used to refer to the southern counties of England bordering the English Midlands. It has a number of different interpretations of its geographic extents. The South is considered by many to be a cultural region with a distinct...

, approximately as far as the border with Devon
Devon
Devon is a large county in southwestern England. The county is sometimes referred to as Devonshire, although the term is rarely used inside the county itself as the county has never been officially "shired", it often indicates a traditional or historical context.The county shares borders with...

 and South Wales
South Wales
South Wales is an area of Wales bordered by England and the Bristol Channel to the east and south, and Mid Wales and West Wales to the north and west. The most densely populated region in the south-west of the United Kingdom, it is home to around 2.1 million people and includes the capital city of...

 in the west, into the Severn
River Severn
The River Severn is the longest river in Great Britain, at about , but the second longest on the British Isles, behind the River Shannon. It rises at an altitude of on Plynlimon, Ceredigion near Llanidloes, Powys, in the Cambrian Mountains of mid Wales...

 valley in the north-west, into the East Midlands
East Midlands
The East Midlands is one of the regions of England, consisting of most of the eastern half of the traditional region of the Midlands. It encompasses the combined area of Nottinghamshire, Derbyshire, Leicestershire, Rutland, Northamptonshire and most of Lincolnshire...

 in the north, and up to the border of Norfolk
Norfolk
Norfolk is a low-lying county in the East of England. It has borders with Lincolnshire to the west, Cambridgeshire to the west and southwest and Suffolk to the south. Its northern and eastern boundaries are the North Sea coast and to the north-west the county is bordered by The Wash. The county...

 in the north-east of its range. The WWF code for this ecoregion is PA0421.

Ecoregional context

To the north, west and south-west lies the similar Celtic broadleaf forests ecoregion, which covers most of the rest of the British Isles
British Isles
The British Isles are a group of islands off the northwest coast of continental Europe that include the islands of Great Britain and Ireland and over six thousand smaller isles. There are two sovereign states located on the islands: the United Kingdom of Great Britain and Northern Ireland and...

. In addition, two further ecoregions are located in the south-western and north-western edges of Ireland
Ireland
Ireland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...

, and the north-western fringes of Scotland
Scotland
Scotland is a country that is part of the United Kingdom. Occupying the northern third of the island of Great Britain, it shares a border with England to the south and is bounded by the North Sea to the east, the Atlantic Ocean to the north and west, and the North Channel and Irish Sea to the...

 (North Atlantic moist mixed forests), and beyond the Scottish Highland Boundary Fault
Highland Boundary Fault
The Highland Boundary Fault is a geological fault that traverses Scotland from Arran and Helensburgh on the west coast to Stonehaven in the east...

 (Caledonian conifer forests
Caledonian Forest
The Caledonian Forest is the name of a type of woodland that once covered vast areas of Scotland. Today, however, only 1% of the original forest survives, covering in 84 locations. The forests are home to a wide variety of wildlife, much of which is not found elsewhere in the British...

). The whole of this Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 archipelago
Archipelago
An archipelago , sometimes called an island group, is a chain or cluster of islands. The word archipelago is derived from the Greek ἄρχι- – arkhi- and πέλαγος – pélagos through the Italian arcipelago...

 is thus considered as originally (or in some sense ideally) forested, with only the far mountainous north being primarily coniferous
Pinophyta
The conifers, division Pinophyta, also known as division Coniferophyta or Coniferae, are one of 13 or 14 division level taxa within the Kingdom Plantae. Pinophytes are gymnosperms. They are cone-bearing seed plants with vascular tissue; all extant conifers are woody plants, the great majority being...

. Across the English Channel
English Channel
The English Channel , often referred to simply as the Channel, is an arm of the Atlantic Ocean that separates southern England from northern France, and joins the North Sea to the Atlantic. It is about long and varies in width from at its widest to in the Strait of Dover...

 lies the Atlantic mixed forests ecoregion in northern France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

 and the Low Countries
Low Countries
The Low Countries are the historical lands around the low-lying delta of the Rhine, Scheldt, and Meuse rivers, including the modern countries of Belgium, the Netherlands, Luxembourg and parts of northern France and western Germany....

.

The difference between the English lowlands beech forests and the Celtic broadleaf forests lies in the fact that south-eastern England is comparatively drier and warmer in climate
Climate of the United Kingdom
The United Kingdom straddles the geographic mid-latitudes between 50-60 N from the equator. It is also positioned on the western seaboard of Eurasia, the world's largest land mass. These boundary conditions allow convergence between moist maritime air and dry continental air...

, and lower-lying in terms of topography. Geologically, something of the distinction can be found in the dominance of the Southern England Chalk Formation
Southern England Chalk Formation
The Chalk Formation of Southern England is a system of chalk downland in the south of England. The formation is perhaps best known for Salisbury Plain, the location of Stonehenge, the Isle of Wight and the twin ridgeways of the North Downs and South Downs....

 in this ecoregion, and the Tees-Exe line
Tees-Exe line
The Tees-Exe line is an imaginary line that can be drawn on a map of Great Britain which roughly divides the lowland and upland regions of the country....

, which divides the island of Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

 into a sedimentary
Sedimentary rock
Sedimentary rock are types of rock that are formed by the deposition of material at the Earth's surface and within bodies of water. Sedimentation is the collective name for processes that cause mineral and/or organic particles to settle and accumulate or minerals to precipitate from a solution....

 south-east, and a metamorphic
Metamorphic rock
Metamorphic rock is the transformation of an existing rock type, the protolith, in a process called metamorphism, which means "change in form". The protolith is subjected to heat and pressure causing profound physical and/or chemical change...

 and igneous
Igneous rock
Igneous rock is one of the three main rock types, the others being sedimentary and metamorphic rock. Igneous rock is formed through the cooling and solidification of magma or lava...

 north-west. However, the WWF division was preceded by that of the Hungarian
Hungary
Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...

 biologist Miklos Udvardy
Miklos Udvardy
Miklos Dezso Ferenc Udvardy was a biologist and biogeographer. He was born on March 23, 1919 in Debrecen, Hungary to Miklos Udvardy and Elizabeth Komlossy. Despite an early interest in birds, his father encouraged him to study law, but later went on to earn a doctorate in biology from the...

, who had considered the greater part of the British Isles as just one biogeographic province in the Palearctic Realm, which he termed British Islands.

Characteristics

Historically, much of this lowland and submontane region was covered with high-canopy
Canopy (forest)
In biology, the canopy is the aboveground portion of a plant community or crop, formed by plant crowns.For forests, canopy also refers to the upper layer or habitat zone, formed by mature tree crowns and including other biological organisms .Sometimes the term canopy is used to refer to the extent...

 forest
Forest
A forest, also referred to as a wood or the woods, is an area with a high density of trees. As with cities, depending where you are in the world, what is considered a forest may vary significantly in size and have various classification according to how and what of the forest is composed...

s dominated by European Beech
European Beech
Fagus sylvatica, the European Beech or Common Beech, is a deciduous tree belonging to the beech family Fagaceae.-Natural range:...

 (Fagus sylvatica), but also including other species of tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

, including oak, ash
European Ash
Fraxinus excelsior , is a species of Fraxinus native to most of Europe with the exception of northern Scandinavia and southern Iberia, and also southwestern Asia from northern Turkey east to the Caucasus and Alborz mountains...

, rowan
Rowan
The rowans or mountain-ashes are shrubs or small trees in genus Sorbus of family Rosaceae. They are native throughout the cool temperate regions of the Northern Hemisphere, with the highest species diversity in the mountains of western China and the Himalaya, where numerous apomictic microspecies...

 and yew
Taxus baccata
Taxus baccata is a conifer native to western, central and southern Europe, northwest Africa, northern Iran and southwest Asia. It is the tree originally known as yew, though with other related trees becoming known, it may be now known as the English yew, or European yew.-Description:It is a small-...

. In summer, the forests are generally cool and dark, because the beech produces a dense canopy, and thus restricts the growth of other species of tree and wild flowers. In the spring, however, thick carpets
Bluebell wood
A bluebell wood is a woodland that in spring-time has a carpet of bluebells underneath a newly forming leaf canopy...

 of bluebells
Common Bluebell
Hyacinthoides non-scripta, commonly known as the common bluebell, is a spring-flowering bulbous perennial plant. -Taxonomy:...

 can be found, flourishing before the beech leaf-cover becomes too thick.

The National Vegetation Classification
British National Vegetation Classification
The British National Vegetation Classification or NVC is a system of classifying natural habitat types in Great Britain according to the vegetation they contain....

 (NVC) plant communities associated with beech forests (together with their occurrence ratios in England as a whole) are:
  • W12 Fagus sylvatica - Mercurialis perennis (Dog's Mercury
    Dog's Mercury
    Mercurialis perennis, commonly known as dog's mercury, is a woodland plant found in much of Europe, but almost absent from Ireland, Orkney and Shetland. A member of the spurge family , it is a herbaceous, downy perennial with erect stems bearing simple, serrate leaves. The dioecious inflorescences...

    ) woodland (base-rich soils) - c. 40%
  • W14 Fagus sylvatica - Rubus fruticosus (Bramble
    Blackberry
    The blackberry is an edible fruit produced by any of several species in the Rubus genus of the Rosaceae family. The fruit is not a true berry; botanically it is termed an aggregate fruit, composed of small drupelets. The plants typically have biennial canes and perennial roots. Blackberries and...

    ) woodland (mesotrophic soils) - c. 45%
  • W15 Fagus sylvatica - Deschampsia flexuosa (Wavy Hair-grass
    Deschampsia
    Deschampsia is a genus of grasses in the family Poaceae, commonly known as hair grass or tussock grass. There are 30 to 40 species....

    ) woodland (acidic soils) - c. 15%


River systems, the most significant of which is the Thames
River Thames
The River Thames flows through southern England. It is the longest river entirely in England and the second longest in the United Kingdom. While it is best known because its lower reaches flow through central London, the river flows alongside several other towns and cities, including Oxford,...

, were historically host to lower-canopy riverine forests dominated by Black Alder, and this can still be encountered occasionally today. Also included in this ecoregion are the distinctive ecosystem
Ecosystem
An ecosystem is a biological environment consisting of all the organisms living in a particular area, as well as all the nonliving , physical components of the environment with which the organisms interact, such as air, soil, water and sunlight....

s associated with the rivers themselves, as well as their flood-meadow
Flood-meadow
A flood-meadow is an area of grassland or pasture beside a river, subject to seasonal flooding. Flood-meadows are distinct from water-meadows in that the latter are artificially created and maintained, with flooding controlled on a seasonal and even daily basis.- Examples :* Angel & Greyhound...

s and estuaries
Estuary
An estuary is a partly enclosed coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea....

. The soils are largely based on limestone, and the climate is temperate with steady amounts of rainfall. Temperatures can fall below freezing in the winter.

Nowadays, much of this ecoregion has been given over to agriculture - with the growing of wheat
Wheat
Wheat is a cereal grain, originally from the Levant region of the Near East, but now cultivated worldwide. In 2007 world production of wheat was 607 million tons, making it the third most-produced cereal after maize and rice...

, barley
Barley
Barley is a major cereal grain, a member of the grass family. It serves as a major animal fodder, as a base malt for beer and certain distilled beverages, and as a component of various health foods...

 and rapeseed
Rapeseed
Rapeseed , also known as rape, oilseed rape, rapa, rappi, rapaseed is a bright yellow flowering member of the family Brassicaceae...

 particularly common - as well as to the raising of livestock, especially cattle
Cattle
Cattle are the most common type of large domesticated ungulates. They are a prominent modern member of the subfamily Bovinae, are the most widespread species of the genus Bos, and are most commonly classified collectively as Bos primigenius...

 and sheep
Domestic sheep
Sheep are quadrupedal, ruminant mammals typically kept as livestock. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Although the name "sheep" applies to many species in the genus Ovis, in everyday usage it almost always refers to Ovis aries...

. In places it is very heavily populated, with towns, suburbs and villages found nearly everywhere - although the plateau of Salisbury Plain
Salisbury Plain
Salisbury Plain is a chalk plateau in central southern England covering . It is part of the Southern England Chalk Formation and largely lies within the county of Wiltshire, with a little in Hampshire. The plain is famous for its rich archaeology, including Stonehenge, one of England's best known...

 remains largely wild. The most significant centre of population is London
London
London is the capital city of :England and the :United Kingdom, the largest metropolitan area in the United Kingdom, and the largest urban zone in the European Union by most measures. Located on the River Thames, London has been a major settlement for two millennia, its history going back to its...

, at the head of the Thames estuary
Thames Estuary
The Thames Mouth is the estuary in which the River Thames meets the waters of the North Sea.It is not easy to define the limits of the estuary, although physically the head of Sea Reach, near Canvey Island on the Essex shore is probably the western boundary...

, one of the largest cities
World's largest cities
This article ranks the world's largest cities, in population or land area, using a variety of ranking methods.-Ambiguities in measuring the "size" of a city:...

 in the world. Due to this high population density, and to a certain amount of depredation caused by grey squirrels
Eastern Gray Squirrel
The eastern gray squirrel is a tree squirrel in the genus Sciurus native to the eastern and midwestern United States, and to the southerly portions of the eastern provinces of Canada...

, edible dormice
Edible dormouse
The edible dormouse or fat dormouse is a large dormouse and the only living species in the genus Glis.-Description:...

 (in the Chilterns
Chiltern Hills
The Chiltern Hills form a chalk escarpment in South East England. They are known locally as "the Chilterns". A large portion of the hills was designated officially as an Area of Outstanding Natural Beauty in 1965.-Location:...

) and deer
Deer
Deer are the ruminant mammals forming the family Cervidae. Species in the Cervidae family include white-tailed deer, elk, moose, red deer, reindeer, fallow deer, roe deer and chital. Male deer of all species and female reindeer grow and shed new antlers each year...

, this forest ecoregion is considered at high risk, with a Critical/Endangered
Ecoregion conservation status
Conservation status of the Global 200 ecoregions is used to classify ecoregions into one of three broad categories: "critical/endangered","vulnerable", or "relatively stable/relatively intact"....

 conservation status accorded it by the WWF. Air pollution
Air pollution
Air pollution is the introduction of chemicals, particulate matter, or biological materials that cause harm or discomfort to humans or other living organisms, or cause damage to the natural environment or built environment, into the atmosphere....

 may also be leading to a reduction in beech numbers, through increased susceptibility to disease
Beech Bark Disease
Beech bark disease is a disease that causes mortality and defects in beech trees in the eastern United States and Europe. In North America, the disease results when the beech scale insect, Cryptococcus fagisuga, attacks the bark, creating a wound. Later, two different fungi Beech bark disease is a...

.

Amongst fauna found in this ecoregion, the West European Hedgehog, Red Fox
Red Fox
The red fox is the largest of the true foxes, as well as being the most geographically spread member of the Carnivora, being distributed across the entire northern hemisphere from the Arctic Circle to North Africa, Central America, and the steppes of Asia...

, Eurasian Badger
Eurasian Badger
The European Badger is a species of badger of the genus Meles, native to almost all of Europe. It is classed as Least Concern for extinction by the IUCN, due to its wide distribution and large population....

, European Rabbit
European Rabbit
The European Rabbit or Common Rabbit is a species of rabbit native to south west Europe and north west Africa . It has been widely introduced elsewhere often with devastating effects on local biodiversity...

 and Wood Mouse
Wood mouse
The wood mouse is a common murid rodent from Europe and northwestern Africa. It is closely related to the yellow-necked mouse but differs in that it has no band of yellow fur around the neck, has slightly smaller ears, and is usually slightly smaller overall: around 90 mm in length...

 are relatively common, while the following are classed as Near Threatened
Near Threatened
Near Threatened is a conservation status assigned to species or lower taxa that may be considered threatened with extinction in the near future, although it does not currently qualify for the threatened status...

 on the IUCN Red List
IUCN Red List
The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

:
  • European Otter
    European Otter
    The European Otter , also known as the Eurasian otter, Eurasian river otter, common otter and Old World otter, is a European and Asian member of the Lutrinae or otter subfamily, and is typical of freshwater otters....

  • Red Squirrel
    Red Squirrel
    The red squirrel or Eurasian red squirrel is a species of tree squirrel in the genus Sciurus common throughout Eurasia...

  • Harvest Mouse
    Harvest Mouse
    The Harvest Mouse, Micromys minutus, is a small rodent native to Europe and Asia. It is typically found in fields of cereal crops such as wheat and oats, in reed beds and in other tall ground vegetation such as long grass and hedgerows. It has reddish-brown fur with white underparts and a naked,...

  • Hazel Dormouse
    Hazel Dormouse
    The Hazel Dormouse or Common Dormouse is a small mammal and the only living species in the genus Muscardinus....

  • Greater Horseshoe Bat
    Greater Horseshoe Bat
    The Greater Horseshoe Bat is a European bat of the Rhinolophus genus. Its distribution covers Europe, Africa, South Asia and Australia. It is the largest of the European Horseshoe Bats and is thus easily distinguished from other species...

  • Corn Crake
    Corn Crake
    The Corn Crake, Corncrake or Landrail is a bird in the rail family. It breeds in Europe and Asia as far east as western China, and migrates to Africa for the winter...



The Barbastelle
Barbastelle
The Barbastelle , also known as the Western Barbastelle, is a European bat. It has a short nose, small eyes and wide ears.It is rare throughout its range...

, as a Vulnerable species
Vulnerable species
On 30 January 2010, the IUCN Red List of Threatened Species identified 9694 Vulnerable species, subspecies and varieties, stocks and sub-populations.-References:...

 on the Red List, is in greater danger still.

Rare plants include the Red Helleborine
Red Helleborine
Red Helleborine is an orchid found in Europe, north Africa and parts of Asia. Although reasonably common in parts of its range, this helleborine has always been one of the rarest orchids in Britain.-Distribution and habitat:...

, Bird's-nest Orchid and Knothole Yoke-moss. Rare fungi include the Devil's Bolete and Hedgehog Mushroom
Hericium erinaceus
Hericium erinaceus is an edible mushroom and medicinal mushroom in the tooth fungus group...

.

History

At the end of the last glaciation, about 10,000 years ago, the area's ecosystem was characterised by a largely treeless tundra
Tundra
In physical geography, tundra is a biome where the tree growth is hindered by low temperatures and short growing seasons. The term tundra comes through Russian тундра from the Kildin Sami word tūndâr "uplands," "treeless mountain tract." There are three types of tundra: Arctic tundra, alpine...

. Pollen studies
Palynology
Palynology is the science that studies contemporary and fossil palynomorphs, including pollen, spores, orbicules, dinoflagellate cysts, acritarchs, chitinozoans and scolecodonts, together with particulate organic matter and kerogen found in sedimentary rocks and sediments...

 have shown that this was replaced by a taiga
Taiga
Taiga , also known as the boreal forest, is a biome characterized by coniferous forests.Taiga is the world's largest terrestrial biome. In North America it covers most of inland Canada and Alaska as well as parts of the extreme northern continental United States and is known as the Northwoods...

 of birch
Birch
Birch is a tree or shrub of the genus Betula , in the family Betulaceae, closely related to the beech/oak family, Fagaceae. The Betula genus contains 30–60 known taxa...

, and then pine
Pine
Pines are trees in the genus Pinus ,in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authorities accept between 105 and 125 species.-Etymology:...

, before their replacement in turn (c. 4500 BC
5th millennium BC
The 5th millennium BC saw the spread of agriculture from the Near East throughout southern and central Europe.Urban cultures in Mesopotamia and Anatolia flourished, developing the wheel. Copper ornaments became more common, marking the Chalcolithic. Animal husbandry spread throughout Eurasia,...

) by most of the species of tree encountered today - including, by 4000 BC
40th century BC
-Events:*c. 4000 BC, Liangzhu culture in China.*c. 4000 BC, More than 100 dwellings surrounding a community center, a cemetery and a kiln are built in Jiangzhai, near modern Xi'an, China.*Start of Naqada culture in Egypt....

, the beech, which seems to have been introduced from mainland Europe. This was used as a source of flour
Flour
Flour is a powder which is made by grinding cereal grains, other seeds or roots . It is the main ingredient of bread, which is a staple food for many cultures, making the availability of adequate supplies of flour a major economic and political issue at various times throughout history...

, ground from the triangular nutlets contained in the "mast", or fruit of the beech, after its tannin
Tannin
A tannin is an astringent, bitter plant polyphenolic compound that binds to and precipitates proteins and various other organic compounds including amino acids and alkaloids.The term tannin refers to the use of...

s had been leached out by soaking. Beechmast has also traditionally been fed to pig
Pig
A pig is any of the animals in the genus Sus, within the Suidae family of even-toed ungulates. Pigs include the domestic pig, its ancestor the wild boar, and several other wild relatives...

s.

However, by 4000 BC, as Oliver Rackham
Oliver Rackham
Oliver Rackham OBE is a Life Fellow of Corpus Christi College, Cambridge. He is also Keeper of theCollege Silver.An acknowledged authority on the British countryside, especially trees, woodlands and wood pasture, Rackham has written a number of well-known books, including The History of the...

 has indicated, the dominant tree species was not the beech, but the Small-leaved Lime
Tilia cordata
Tilia cordata is a species of Tilia native to much of Europe and western Asia, north to southern Great Britain , central Scandinavia, east to central Russia, and south to central Spain, Italy, Bulgaria, Turkey, and the Caucasus; in the south of its range it is restricted to...

, also known as the pry tree. The wildwood was made up of a patchwork of lime-wood areas and hazel
Hazel
The hazels are a genus of deciduous trees and large shrubs native to the temperate northern hemisphere. The genus is usually placed in the birch family Betulaceae, though some botanists split the hazels into a separate family Corylaceae.They have simple, rounded leaves with double-serrate margins...

-wood areas, interspersed with oak and elm
Elm
Elms are deciduous and semi-deciduous trees comprising the genus Ulmus in the plant family Ulmaceae. The dozens of species are found in temperate and tropical-montane regions of North America and Eurasia, ranging southward into Indonesia. Elms are components of many kinds of natural forests...

 and other species. The pry seems to have become less abundant now because the climate has turned against it, making it difficult for it to grow from seed. Nevertheless, some remnants of ancient lime-wood still remain in south Suffolk
Suffolk
Suffolk is a non-metropolitan county of historic origin in East Anglia, England. It has borders with Norfolk to the north, Cambridgeshire to the west and Essex to the south. The North Sea lies to the east...

.

Clearance of forests began with the introduction of farming (c. 4500 BC), particularly in the higher-lying parts of the country, like the South Downs
South Downs
The South Downs is a range of chalk hills that extends for about across the south-eastern coastal counties of England from the Itchen Valley of Hampshire in the west to Beachy Head, near Eastbourne, East Sussex, in the east. It is bounded on its northern side by a steep escarpment, from whose...

. At this time, the whole region, apart from upland areas under plough, and marshy areas (e.g. Romney Marsh
Romney Marsh
Romney Marsh is a sparsely populated wetland area in the counties of Kent and East Sussex in the south-east of England. It covers about 100 mi ² .-Quotations:*“As Egypt was the gift of the Nile, this level tract .....

 in Kent
Kent
Kent is a county in southeast England, and is one of the home counties. It borders East Sussex, Surrey and Greater London and has a defined boundary with Essex in the middle of the Thames Estuary. The ceremonial county boundaries of Kent include the shire county of Kent and the unitary borough of...

 and much of Somerset
Somerset
The ceremonial and non-metropolitan county of Somerset in South West England borders Bristol and Gloucestershire to the north, Wiltshire to the east, Dorset to the south-east, and Devon to the south-west. It is partly bounded to the north and west by the Bristol Channel and the estuary of the...

), was heavily forested, with woodland stretching nearly everywhere.

Notable surviving examples include:
  • The Forest of Arden
    Arden, Warwickshire
    Arden is an area, mainly located in Warwickshire, England, traditionally regarded as stretching from the River Avon to the River Tame.-History:...

     (Warwickshire
    Warwickshire
    Warwickshire is a landlocked non-metropolitan county in the West Midlands region of England. The county town is Warwick, although the largest town is Nuneaton. The county is famous for being the birthplace of William Shakespeare...

    )
  • The Chilterns
    Chiltern Hills
    The Chiltern Hills form a chalk escarpment in South East England. They are known locally as "the Chilterns". A large portion of the hills was designated officially as an Area of Outstanding Natural Beauty in 1965.-Location:...

     (on the heights running from Oxfordshire
    Oxfordshire
    Oxfordshire is a county in the South East region of England, bordering on Warwickshire and Northamptonshire , Buckinghamshire , Berkshire , Wiltshire and Gloucestershire ....

     through Buckinghamshire
    Buckinghamshire
    Buckinghamshire is a ceremonial and non-metropolitan home county in South East England. The county town is Aylesbury, the largest town in the ceremonial county is Milton Keynes and largest town in the non-metropolitan county is High Wycombe....

     and Hertfordshire
    Hertfordshire
    Hertfordshire is a ceremonial and non-metropolitan county in the East region of England. The county town is Hertford.The county is one of the Home Counties and lies inland, bordered by Greater London , Buckinghamshire , Bedfordshire , Cambridgeshire and...

     to Bedfordshire
    Bedfordshire
    Bedfordshire is a ceremonial county of historic origin in England that forms part of the East of England region.It borders Cambridgeshire to the north-east, Northamptonshire to the north, Buckinghamshire to the west and Hertfordshire to the south-east....

    )
  • Epping Forest
    Epping Forest
    Epping Forest is an area of ancient woodland in south-east England, straddling the border between north-east Greater London and Essex. It is a former royal forest, and is managed by the City of London Corporation....

     (on the border of north-east Greater London
    Greater London
    Greater London is the top-level administrative division of England covering London. It was created in 1965 and spans the City of London, including Middle Temple and Inner Temple, and the 32 London boroughs. This territory is coterminate with the London Government Office Region and the London...

     and Essex
    Essex
    Essex is a ceremonial and non-metropolitan county in the East region of England, and one of the home counties. It is located to the northeast of Greater London. It borders with Cambridgeshire and Suffolk to the north, Hertfordshire to the west, Kent to the South and London to the south west...

    )
  • Kinver Edge
    Kinver Edge
    Kinver Edge is a high heath and woodland escarpment just west of Kinver, about four miles west of Stourbridge, and four miles north of Kidderminster, and is on the border between Worcestershire and Staffordshire, England. It is now owned by the National Trust....

     (a remnant of the Mercia
    Mercia
    Mercia was one of the kingdoms of the Anglo-Saxon Heptarchy. It was centred on the valley of the River Trent and its tributaries in the region now known as the English Midlands...

    n forest on the border of south Staffordshire
    Staffordshire
    Staffordshire is a landlocked county in the West Midlands region of England. For Eurostat purposes, the county is a NUTS 3 region and is one of four counties or unitary districts that comprise the "Shropshire and Staffordshire" NUTS 2 region. Part of the National Forest lies within its borders...

     and Worcestershire
    Worcestershire
    Worcestershire is a non-metropolitan county, established in antiquity, located in the West Midlands region of England. For Eurostat purposes it is a NUTS 3 region and is one of three counties that comprise the "Herefordshire, Worcestershire and Warwickshire" NUTS 2 region...

    )
  • Morfe Forest (south Shropshire
    Shropshire
    Shropshire is a county in the West Midlands region of England. For Eurostat purposes, the county is a NUTS 3 region and is one of four counties or unitary districts that comprise the "Shropshire and Staffordshire" NUTS 2 region. It borders Wales to the west...

    )
  • Savernake Forest
    Savernake Forest
    Savernake Forest is on a Cretaceous chalk plateau between Marlborough and Great Bedwyn in Wiltshire, England. Its area is approximately .It is privately owned by the Trustees of Savernake Estate, the Earl of Cardigan, and his family solicitor. Since 1939 the running of the forest has been...

     (Wiltshire
    Wiltshire
    Wiltshire is a ceremonial county in South West England. It is landlocked and borders the counties of Dorset, Somerset, Hampshire, Gloucestershire, Oxfordshire and Berkshire. It contains the unitary authority of Swindon and covers...

    )
  • Selwood Forest
    Selwood Forest
    Selwood Forest is an area of woodland on the borders between Somerset, Dorset and Wiltshire in south west England. In Anglo-Saxon times it was far more substantial and covered a much greater area forming a natural barrier between the Anglo-Saxons of Wessex and the Britons of Dumnonia and the Severn...

     (Somerset)
  • The Weald (Kent, East Sussex
    East Sussex
    East Sussex is a county in South East England. It is bordered by the counties of Kent, Surrey and West Sussex, and to the south by the English Channel.-History:...

    , West Sussex
    West Sussex
    West Sussex is a county in the south of England, bordering onto East Sussex , Hampshire and Surrey. The county of Sussex has been divided into East and West since the 12th century, and obtained separate county councils in 1888, but it remained a single ceremonial county until 1974 and the coming...

     and Surrey
    Surrey
    Surrey is a county in the South East of England and is one of the Home Counties. The county borders Greater London, Kent, East Sussex, West Sussex, Hampshire and Berkshire. The historic county town is Guildford. Surrey County Council sits at Kingston upon Thames, although this has been part of...

    )
  • Wychwood
    Wychwood
    The Wychwood, or Wychwood Forest, is an area now covering a small part of rural Oxfordshire. In past centuries the forest covered a much larger area, since cleared in favour of agriculture, villages and towns. However, the forest's area has fluctuated...

     (Oxfordshire)
  • Wyre Forest
    Wyre Forest
    Wyre Forest is a large, semi-natural woodland and forest which straddles the borders of Worcestershire and Shropshire, England.The Wyre Forest district of Worcestershire takes its name from the forest, despite the fact that much of the woodland does not lie within the district's boundaries, but...

     (on the border of Worcestershire and Shropshire)


All of these were once far more extensive than they are today. For example, according to a late 9th century writer, the Weald (from the Anglo-Saxon
Old English language
Old English or Anglo-Saxon is an early form of the English language that was spoken and written by the Anglo-Saxons and their descendants in parts of what are now England and southeastern Scotland between at least the mid-5th century and the mid-12th century...

 word weald = "forest") once stretched from Kent to Hampshire
Hampshire
Hampshire is a county on the southern coast of England in the United Kingdom. The county town of Hampshire is Winchester, a historic cathedral city that was once the capital of England. Hampshire is notable for housing the original birthplaces of the Royal Navy, British Army, and Royal Air Force...

, and was 120 miles (193.1 km) long by 30 broad. The New Forest
New Forest
The New Forest is an area of southern England which includes the largest remaining tracts of unenclosed pasture land, heathland and forest in the heavily-populated south east of England. It covers south-west Hampshire and extends into south-east Wiltshire....

 (in south-west Hampshire) remains the largest intact forested area in this ecoregion (at 571 km²), although the hedgerow system, which separates fields from lanes and also from other fields, is also extensive, and serves as an important habitat for otherwise displaced woodland fauna. Some species-rich
Species richness
Species richness is the number of different species in a given area. It is represented in equation form as S.Species richness is the fundamental unit in which to assess the homogeneity of an environment. Typically, species richness is used in conservation studies to determine the sensitivity of...

 hedgerows date back at least 700 years, if not 1,000. For many species of bird, significant estuarine habitats include the Thames and Severn estuaries, and the mid-Essex coast.

The Mesozoic
Mesozoic
The Mesozoic era is an interval of geological time from about 250 million years ago to about 65 million years ago. It is often referred to as the age of reptiles because reptiles, namely dinosaurs, were the dominant terrestrial and marine vertebrates of the time...

 history of the area can be seen in the Jurassic Coast
Jurassic Coast
The Jurassic Coast is a World Heritage Site on the English Channel coast of southern England. The site stretches from Orcombe Point near Exmouth in East Devon to Old Harry Rocks near Swanage in East Dorset, a distance of ....

 World Heritage Site
World Heritage Site
A UNESCO World Heritage Site is a place that is listed by the UNESCO as of special cultural or physical significance...

, where about 180 Ma of fossil-rich sedimentary deposits have been exposed along a 95 miles (152.9 km) stretch of the Dorset
Dorset
Dorset , is a county in South West England on the English Channel coast. The county town is Dorchester which is situated in the south. The Hampshire towns of Bournemouth and Christchurch joined the county with the reorganisation of local government in 1974...

 and East Devon coast. The science of palaeontology
Paleontology
Paleontology "old, ancient", ὄν, ὀντ- "being, creature", and λόγος "speech, thought") is the study of prehistoric life. It includes the study of fossils to determine organisms' evolution and interactions with each other and their environments...

 can be said to have started in large measure here, with the pioneering work of Mary Anning
Mary Anning
Mary Anning was a British fossil collector, dealer and palaeontologist who became known around the world for a number of important finds she made in the Jurassic age marine fossil beds at Lyme Regis where she lived...

.

The Great Storm of 1987
Great Storm of 1987
The Great Storm of 1987 occurred on the night of 15/16 October 1987, when an unusually strong weather system caused winds to hit much of southern England and northern France...

 was responsible for the uprooting
Windthrow
In forestry, windthrow refers to trees uprooted or broken by wind. Breakage of the tree bole instead of uprooting is sometimes called windsnap.- Causes :...

 of some 15 million trees in this area.

See also

  • Ancient woodland
    Ancient woodland
    Ancient woodland is a term used in the United Kingdom to refer specifically to woodland that has existed continuously since 1600 or before in England and Wales . Before those dates, planting of new woodland was uncommon, so a wood present in 1600 was likely to have developed naturally...

  • Biodiversity
    Biodiversity
    Biodiversity is the degree of variation of life forms within a given ecosystem, biome, or an entire planet. Biodiversity is a measure of the health of ecosystems. Biodiversity is in part a function of climate. In terrestrial habitats, tropical regions are typically rich whereas polar regions...

  • Biodiversity Action Plan
  • Bioregionalism
    Bioregionalism
    Bioregionalism is a political, cultural, and environmental system or set of views based on naturally defined areas called bioregions, similar to ecoregions. Bioregions are defined through physical and environmental features, including watershed boundaries and soil and terrain characteristics...

  • Community Forests in England
    Community Forests in England
    England's twelve community forests are afforestation-based regeneration projects which were established in the early 1990s. Each of them is a partnership between the Forestry Commission and the Countryside Agency, which are agencies of the British government, and the relevant local councils.Most of...

  • Conservation biology
    Conservation biology
    Conservation biology is the scientific study of the nature and status of Earth's biodiversity with the aim of protecting species, their habitats, and ecosystems from excessive rates of extinction...

  • Forestry Commission
    Forestry Commission
    The Forestry Commission is a non-ministerial government department responsible for forestry in Great Britain. Its mission is to protect and expand Britain's forests and woodlands and increase their value to society and the environment....

  • Geology of England
    Geology of England
    The Geology of England is mainly sedimentary. The youngest rocks are in the south east around London, progressing in age in a north westerly direction. The Tees-Exe line marks the division between younger, softer and low-lying rocks in the south east and older, harder, and generally a higher relief...

  • List of ecoregions
  • List of ecoregions in the United Kingdom (UK)
  • List of forests in the United Kingdom
  • National Nature Reserves in England
    National Nature Reserves in England
    National Nature Reserves in England are managed by Natural England and are key places for wildlife and natural features in England. They were established to protect the most important areas of habitat and of geological formations...

  • National parks of England and Wales (New Forest and South Downs)
    National parks of England and Wales
    The national parks of England and Wales are areas of relatively undeveloped and scenic landscape that are designated under the National Parks and Access to the Countryside Act 1949...

  • Protected areas of the United Kingdom
    Protected areas of the United Kingdom
    Protected areas of the United Kingdom are areas in the United Kingdom which need and /or receive protection because of their environmental, historical or cultural value to the nation. Methods and aims of protection vary depending on the nature and importance of the resource...

  • Royal forest
    Royal forest
    A royal forest is an area of land with different meanings in England, Wales and Scotland; the term forest does not mean forest as it is understood today, as an area of densely wooded land...

  • Trees of Britain and Ireland
    Trees of Britain and Ireland
    Many lists of Trees of Britain and Ireland have been written. There are a number of issues surrounding the inclusion of a species in such a list...


External links

The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
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