Song Thrush
Encyclopedia
The Song Thrush is a thrush
Thrush (bird)
The thrushes, family Turdidae, are a group of passerine birds that occur worldwide.-Characteristics:Thrushes are plump, soft-plumaged, small to medium-sized birds, inhabiting wooded areas, and often feed on the ground or eat small fruit. The smallest thrush may be the Forest Rock-thrush, at and...

 that breeds across much of Eurasia
Eurasia
Eurasia is a continent or supercontinent comprising the traditional continents of Europe and Asia ; covering about 52,990,000 km2 or about 10.6% of the Earth's surface located primarily in the eastern and northern hemispheres...

. It is also known in English
English language
English is a West Germanic language that arose in the Anglo-Saxon kingdoms of England and spread into what was to become south-east Scotland under the influence of the Anglian medieval kingdom of Northumbria...

 dialect
Dialect
The term dialect is used in two distinct ways, even by linguists. One usage refers to a variety of a language that is a characteristic of a particular group of the language's speakers. The term is applied most often to regional speech patterns, but a dialect may also be defined by other factors,...

s as throstle or mavis. It has brown upperparts and black-spotted cream or buff underparts and has three recognised subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

. Its distinctive song
Bird song
Bird vocalization includes both bird calls and bird songs. In non-technical use, bird songs are the bird sounds that are melodious to the human ear. In ornithology and birding, songs are distinguished by function from calls.-Definition:The distinction between songs and calls is based upon...

, which has repeated musical phrases, has frequently been referred to in poetry.

The Song Thrush breeds in forest
Forest
A forest, also referred to as a wood or the woods, is an area with a high density of trees. As with cities, depending where you are in the world, what is considered a forest may vary significantly in size and have various classification according to how and what of the forest is composed...

s, gardens and parks, and is partially migratory
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 with many birds wintering in southern Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

 and the Middle East
Middle East
The Middle East is a region that encompasses Western Asia and Northern Africa. It is often used as a synonym for Near East, in opposition to Far East...

; it has also been introduced into New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

 and Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

. Although it is not threatened globally, there have been serious population declines in parts of Europe, possibly due to changes in farming practices.

The Song Thrush builds a neat mud-lined cup nest in a bush or tree and lays four or five dark-spotted blue eggs
Bird egg
Bird eggs are laid by females and incubated for a time that varies according to the species; a single young hatches from each egg. Average clutch sizes range from one to about 17...

. It is omnivorous and has the habit of using a favourite stone as an "anvil" on which to smash snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

s. Like other perching birds (passerine
Passerine
A passerine is a bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds or, less accurately, as songbirds, the passerines form one of the most diverse terrestrial vertebrate orders: with over 5,000 identified species, it has roughly...

s), it is affected by external and internal parasites and is vulnerable to predation by cat
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

s and birds of prey
Bird of prey
Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. They are defined as birds that primarily hunt vertebrates, including other birds. Their talons and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....

.

Name

The Song Thrush was described by German
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

 ornithologist
Ornithology
Ornithology is a branch of zoology that concerns the study of birds. Several aspects of ornithology differ from related disciplines, due partly to the high visibility and the aesthetic appeal of birds...

 Christian Ludwig Brehm
Christian Ludwig Brehm
Christian Ludwig Brehm was a German pastor and ornithologist. He was the father of Alfred Brehm.Brehm was born near Gotha, and studied at the University of Jena. In 1813 he became the minister at Renthendorf, a village sixty miles south of Leipzig, where he remained until his death...

 in 1831, and still bears its original scientific name, Turdus philomelos. The generic name, Turdus, is the Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 for thrush, and the specific epithet refers to a character in Greek mythology
Greek mythology
Greek mythology is the body of myths and legends belonging to the ancient Greeks, concerning their gods and heroes, the nature of the world, and the origins and significance of their own cult and ritual practices. They were a part of religion in ancient Greece...

, Philomela, who had her tongue cut out, but was changed into a singing bird. Her name is derived from the Ancient Greek
Ancient Greek
Ancient Greek is the stage of the Greek language in the periods spanning the times c. 9th–6th centuries BC, , c. 5th–4th centuries BC , and the c. 3rd century BC – 6th century AD of ancient Greece and the ancient world; being predated in the 2nd millennium BC by Mycenaean Greek...

 Φιλο philo- (loving), and μέλος melos (song). The dialect names throstle and mavis both mean thrush, being related to the German
German language
German is a West Germanic language, related to and classified alongside English and Dutch. With an estimated 90 – 98 million native speakers, German is one of the world's major languages and is the most widely-spoken first language in the European Union....

 drossel and French
French language
French is a Romance language spoken as a first language in France, the Romandy region in Switzerland, Wallonia and Brussels in Belgium, Monaco, the regions of Quebec and Acadia in Canada, and by various communities elsewhere. Second-language speakers of French are distributed throughout many parts...

 mauvis respectively. Throstle dates back to at least the fourteenth century and was used by Chaucer
Geoffrey Chaucer
Geoffrey Chaucer , known as the Father of English literature, is widely considered the greatest English poet of the Middle Ages and was the first poet to have been buried in Poet's Corner of Westminster Abbey...

 in the Parliament of Fowls. Mavis is derived via Middle English
Middle English
Middle English is the stage in the history of the English language during the High and Late Middle Ages, or roughly during the four centuries between the late 11th and the late 15th century....

 mavys and Old French
Old French
Old French was the Romance dialect continuum spoken in territories that span roughly the northern half of modern France and parts of modern Belgium and Switzerland from the 9th century to the 14th century...

 mauvis from Middle Breton milhuyt meaning "thrush." Mavis (Μαβής) can also mean "purple
Purple
Purple is a range of hues of color occurring between red and blue, and is classified as a secondary color as the colors are required to create the shade....

" in Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...

.

Classification

A recent molecular study indicates that the Song Thrush's closest relatives are the similarly plumaged Mistle Thrush
Mistle Thrush
The Mistle Thrush is a member of the thrush family Turdidae.It is found in open woods and cultivated land over all of Europe and much of Asia...

 (T. viscivorus) and the Chinese Thrush
Chinese Thrush
The Chinese Thrush is a species of bird in the Turdidae family.It is found in China and Vietnam.Its natural habitats are temperate forests and subtropical or tropical moist montanes....

 (T. mupinensis); these three species are early offshoots from the lineage of Turdus thrushes before they diversified and spread across the globe, and hence are less closely related to other European thrush species such as the Blackbird (T. merula).

The Song Thrush has three subspecies, with the nominate subspecies, T. p. philomelos, covering the majority of the species' range. T. p. hebridensis, described by British
United Kingdom
The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...

 ornithologist William Eagle Clarke
William Eagle Clarke
Dr William Eagle Clarke I.S.O LL.D. was a British ornithologist.Clarke was born in Leeds where his father William Clarke was a solicitor and educated at the Grammar School and at Yorkshire College, Leeds where he studied under Professor L C Miall. He was originally a civil engineer and surveyor,...

 in 1913, is a mainly sedentary (non-migratory) form found in the Outer Hebrides
Outer Hebrides
The Outer Hebrides also known as the Western Isles and the Long Island, is an island chain off the west coast of Scotland. The islands are geographically contiguous with Comhairle nan Eilean Siar, one of the 32 unitary council areas of Scotland...

 and Isle of Skye in Scotland
Scotland
Scotland is a country that is part of the United Kingdom. Occupying the northern third of the island of Great Britain, it shares a border with England to the south and is bounded by the North Sea to the east, the Atlantic Ocean to the north and west, and the North Channel and Irish Sea to the...

. It is the darkest subspecies, with a dark brown back, greyish rump, pale buff background colour to the underparts and grey-tinged flanks.

T. p. clarkei, described by German zoologist
Zoology
Zoology |zoölogy]]), is the branch of biology that relates to the animal kingdom, including the structure, embryology, evolution, classification, habits, and distribution of all animals, both living and extinct...

 Ernst Hartert
Ernst Hartert
Ernst Johann Otto Hartert was a German ornithologist. Hartert was born in Hamburg. He was employed by Lionel Walter Rothschild as ornithological curator of his private museum at Tring from 1892 to 1929....

 in 1909, and named for Eagle Clarke, breeds in the rest of Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

 and Ireland
Ireland
Ireland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...

 and on mainland Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 in France, Belgium
Belgium
Belgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...

, the Netherlands
Netherlands
The Netherlands is a constituent country of the Kingdom of the Netherlands, located mainly in North-West Europe and with several islands in the Caribbean. Mainland Netherlands borders the North Sea to the north and west, Belgium to the south, and Germany to the east, and shares maritime borders...

 and possibly somewhat further east. It has brown upperparts which are warmer in tone than those of the nominate form, an olive-tinged rump and rich yellow background colour to the underparts. It is a partial migrant with some birds wintering in southern France and Iberia
Iberian Peninsula
The Iberian Peninsula , sometimes called Iberia, is located in the extreme southwest of Europe and includes the modern-day sovereign states of Spain, Portugal and Andorra, as well as the British Overseas Territory of Gibraltar...

. This form intergrades with the nominate subspecies in central Europe, and with T. p. hebridensis in the Inner Hebrides
Inner Hebrides
The Inner Hebrides is an archipelago off the west coast of Scotland, to the south east of the Outer Hebrides. Together these two island chains form the Hebrides, which enjoy a mild oceanic climate. There are 36 inhabited islands and a further 43 uninhabited Inner Hebrides with an area greater than...

 and western Scotland
Scotland
Scotland is a country that is part of the United Kingdom. Occupying the northern third of the island of Great Britain, it shares a border with England to the south and is bounded by the North Sea to the east, the Atlantic Ocean to the north and west, and the North Channel and Irish Sea to the...

, and in these areas birds show intermediate characteristics. Additional subspecies, such as T. p. nataliae of Siberia
Siberia
Siberia is an extensive region constituting almost all of Northern Asia. Comprising the central and eastern portion of the Russian Federation, it was part of the Soviet Union from its beginning, as its predecessor states, the Tsardom of Russia and the Russian Empire, conquered it during the 16th...

, proposed by the Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

n Sergei Buturlin
Sergei Aleksandrovich Buturlin
Sergei Aleksandrovich Buturlin was a Russian ornithologist.A scion of one of the oldest families of Russian nobility, Buturlin spent most his life in Russia. He went to school in Simbirsk and studied jurisprudence in St...

 in 1929, are not widely accepted.

Description

The Song Thrush (as represented by the nominate subspecies T. p. philomelos) is 20 to 23.5 centimetres (8 to 9.25 in) in length and weighs 50–107 gram
Gram
The gram is a metric system unit of mass....

mes (1.8 to 3.8 oz
Ounce
The ounce is a unit of mass with several definitions, the most commonly used of which are equal to approximately 28 grams. The ounce is used in a number of different systems, including various systems of mass that form part of the imperial and United States customary systems...

). The sexes are similar, with plain brown backs and neatly black-spotted cream or yellow-buff underparts, becoming paler on the belly. The underwing is warm yellow, the bill is yellowish and the legs and feet are pink. The upperparts of this species become colder in tone from west to east across the breeding range from Sweden
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

 to Siberia. The juvenile resembles the adult, but has buff or orange streaks on the back and wing coverts
Covert (feather)
A covert feather on a bird is one of a set of feathers, called coverts, which as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail.- Wing-coverts :...

.

The most similar European thrush species is the Redwing
Redwing
The Redwing is a bird in the thrush family Turdidae, native to Europe and Asia, slightly smaller than the related Song Thrush.-Taxonomy:...

 (T. iliacus), but that bird has a strong white supercilium
Supercilium
The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head. Also known as an "eyebrow", it is distinct from the eyestripe, which is a line which runs...

, red flanks, and shows a red underwing in flight. The Mistle Thrush
Mistle Thrush
The Mistle Thrush is a member of the thrush family Turdidae.It is found in open woods and cultivated land over all of Europe and much of Asia...

 (T. viscivorus) is much larger and has white tail corners, and the Chinese Thrush
Chinese Thrush
The Chinese Thrush is a species of bird in the Turdidae family.It is found in China and Vietnam.Its natural habitats are temperate forests and subtropical or tropical moist montanes....

 (T. mupinensis), although much more similar in plumage, has black face markings and does not overlap in range.

The Song Thrush has a short, sharp tsip call, replaced on migration by a thin high seep, similar to the Redwing's call but shorter. The alarm call is a chook-chook becoming shorter and more strident with increasing danger. The male's song, given from trees, rooftops or other elevated perches, is a loud clear run of musical phrases, repeated two to four times, filip filip filip codidio codidio quitquiquit tittit tittit tereret tereret tereret, and interspersed with grating notes and mimicry. It is given mainly from February to June by the Outer Hebridean race, but from November to July by the more widespread subspecies. For its weight, this species has one of the loudest bird calls.

An individual male may have a repertoire of more than 100 phrases, many copied from its parents and neighbouring birds. Mimicry may include the imitation of man-made items like telephones, and the Song Thrush will also repeat the calls of captive birds, including exotics such as the White-faced Whistling Duck
White-faced Whistling Duck
The White-faced Whistling Duck, Dendrocygna viduata, is a whistling duck which breeds in sub-Saharan Africa and much of South America.This species is gregarious, and at favoured sites, the flocks of a thousand or more birds arriving at dawn are an impressive sight...

.

Distribution and habitat

The Song Thrush breeds in most of Europe (although not in the greater part of Iberia
Iberian Peninsula
The Iberian Peninsula , sometimes called Iberia, is located in the extreme southwest of Europe and includes the modern-day sovereign states of Spain, Portugal and Andorra, as well as the British Overseas Territory of Gibraltar...

, lowland Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

 or southern Greece
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

), and across the Ukraine
Ukraine
Ukraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...

 and Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 almost to Lake Baikal
Lake Baikal
Lake Baikal is the world's oldest at 30 million years old and deepest lake with an average depth of 744.4 metres.Located in the south of the Russian region of Siberia, between Irkutsk Oblast to the northwest and the Buryat Republic to the southeast, it is the most voluminous freshwater lake in the...

. It reaches to 75oN in Norway, but only to about 60oN in Siberia. Birds from Scandinavia
Scandinavia
Scandinavia is a cultural, historical and ethno-linguistic region in northern Europe that includes the three kingdoms of Denmark, Norway and Sweden, characterized by their common ethno-cultural heritage and language. Modern Norway and Sweden proper are situated on the Scandinavian Peninsula,...

, Eastern Europe
Eastern Europe
Eastern Europe is the eastern part of Europe. The term has widely disparate geopolitical, geographical, cultural and socioeconomic readings, which makes it highly context-dependent and even volatile, and there are "almost as many definitions of Eastern Europe as there are scholars of the region"...

 and Russia winter around the Mediterranean, North Africa and the Middle East
Middle East
The Middle East is a region that encompasses Western Asia and Northern Africa. It is often used as a synonym for Near East, in opposition to Far East...

, but only some of the birds in the milder west of the breeding range leave their breeding areas.

Birds of the nominate subspecies were introduced to New Zealand and Australia by acclimatisation societies
Acclimatisation society
Acclimatisation societies were societies created in order to enrich the fauna of a region with animals and plants from around the world. The first such society was La Societé Zoologique d'Acclimatation founded in Paris in 1854 by Isidore Geoffroy Saint-Hilaire. Such societies spread quickly around...

 between 1860 and 1880, apparently for purely sentimental reasons. In New Zealand, where it was introduced on both the main islands, the Song Thrush quickly established itself and spread to surrounding islands such as the Kermadecs
Kermadec Islands
The Kermadec Islands are a subtropical island arc in the South Pacific Ocean northeast of New Zealand's North Island, and a similar distance southwest of Tonga...

, Chatham
Chatham Islands
The Chatham Islands are an archipelago and New Zealand territory in the Pacific Ocean consisting of about ten islands within a radius, the largest of which are Chatham Island and Pitt Island. Their name in the indigenous language, Moriori, means Misty Sun...

 and Auckland Islands
Auckland Islands
The Auckland Islands are an archipelago of the New Zealand Sub-Antarctic Islands and include Auckland Island, Adams Island, Enderby Island, Disappointment Island, Ewing Island, Rose Island, Dundas Island and Green Island, with a combined area of...

. Although it is common and widespread in New Zealand, in Australia only a small population survives around Melbourne
Melbourne
Melbourne is the capital and most populous city in the state of Victoria, and the second most populous city in Australia. The Melbourne City Centre is the hub of the greater metropolitan area and the Census statistical division—of which "Melbourne" is the common name. As of June 2009, the greater...

. In New Zealand, there appears to be a limited detrimental effect on some invertebrates due to predation by introduced bird species, and the Song Thrush also damages commercial fruit crops in that country. As an introduced species it has no legal protection in New Zealand, and can be killed at any time.

The Song Thrush typically nests in forest with good undergrowth and nearby more open areas, and in western Europe also uses gardens and parks. It breeds up to the tree-line, reaching 2,200 metres (7,250 ft) in Switzerland
Switzerland
Switzerland name of one of the Swiss cantons. ; ; ; or ), in its full name the Swiss Confederation , is a federal republic consisting of 26 cantons, with Bern as the seat of the federal authorities. The country is situated in Western Europe,Or Central Europe depending on the definition....

. The island subspecies T. p. hebridensis breeds in more open country, including heathland
Heath (habitat)
A heath or heathland is a dwarf-shrub habitat found on mainly low quality acidic soils, characterised by open, low growing woody vegetation, often dominated by plants of the Ericaceae. There are some clear differences between heath and moorland...

, and in the east of the Song Thrush's Eurasian range, the nominate subspecies is restricted to the edge of the dense conifer forests.

In intensively farmed areas where agricultural practices appear to have made cropped land unsuitable, gardens are an important breeding habitat. In one English
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

 study, only 3.5% of territories were found in farmland, whereas gardens held 71.5% of the territories, despite that habitat making up only 2% of the total area. The remaining nests were in woodlands (1% of total area).

The winter habitat is similar to that used for breeding, except that high ground and other exposed localities are avoided; however, the island subspecies T. p. hebridensis will frequent the seashore in winter.

Behaviour

The Song Thrush is not usually gregarious, although several birds may roost together in winter or be loosely associated in suitable feeding habitats, perhaps with other thrushes such as the Blackbird, Fieldfare
Fieldfare
The Fieldfare is a member of the thrush family Turdidae. It breeds in woodland and scrub in northern Europe and Asia. It is strongly migratory, with many northern birds moving south during the winter. It is a very rare breeder in Great Britain and Ireland, but winters in large numbers in these...

, Redwing and Dark-throated Thrush
Dark-throated Thrush
The Dark-throated Thrush is a passerine bird in the thrush family. It has two races: T. r. atrogularis, the Black-throated Thrush, and T. r. ruficollis, the Red-throated Thrush. These are sometimes considered different species...

. Unlike the more nomadic Fieldfare and Redwing, the Song Thrush tends to return regularly to the same wintering areas.

This is a monogamous
Monogamy
Monogamy /Gr. μονός+γάμος - one+marriage/ a form of marriage in which an individual has only one spouse at any one time. In current usage monogamy often refers to having one sexual partner irrespective of marriage or reproduction...

 territorial species, and in areas where it is fully migratory, the male re-establishes its breeding territory and starts singing as soon as he returns. In the milder areas where some birds stay year round, the resident male remains in his breeding territory, singing intermittently, but the female may establish a separate individual wintering range until pair formation begins in the early spring.

During migration, the Song Thrush travels mainly at night with a strong and direct flight action. It flies in loose flocks which cross the sea on a broad front rather than concentrating at short crossings (as occurs in the migration of large soaring birds), and calls frequently to maintain contact. Migration may start as early as late August in the most easterly and northerly parts of the range, but the majority of birds, with shorter distances to cover, head south from September to mid-December. However, hard weather may force further movement. Return migration varies between mid-February around the Mediterranean to May in northern Sweden and central Siberia. Vagrants
Vagrancy (biology)
Vagrancy is a phenomenon in biology whereby individual animals appear well outside their normal range; individual animals which exhibit vagrancy are known as vagrants. The term accidental is sometimes also used...

 have been recorded in Greenland
Greenland
Greenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...

, various Atlantic islands, and West Africa.

Breeding and survival

The female Song Thrush builds a neat cup-shaped nest lined with mud and dry grass in a bush, tree or creeper, or, in the case of the Hebridean subspecies, on the ground. She lays four or five bright glossy blue eggs
Egg (biology)
An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

 which are lightly spotted with black or purple; they are typically 2.7 x 2.0 centimetres (0.79 x 1.06 in) in size and weigh 6.0 grammes (0.21 oz), of which 6% is shell. The female incubates the eggs alone for 10–17 days, and after hatching a similar time elapses until the young fledge. Two or three broods in a year is normal, although only one may be raised in the north of the range. On average, 54.6% of British juveniles survive the first year of life, and the adult annual survival rate is 62.2%. The typical lifespan is three years, but the maximum recorded age is 10 years 8 months.
The Song Thrush is occasionally a host of parasitic
Brood parasite
Brood parasites are organisms that use the strategy of brood parasitism, a kind of kleptoparasitism found among birds, fish or insects, involving the manipulation and use of host individuals either of the same or different species to raise the young of the brood-parasite...

 cuckoo
Cuckoo
The cuckoos are a family, Cuculidae, of near passerine birds. The order Cuculiformes, in addition to the cuckoos, also includes the turacos . Some zoologists and taxonomists have also included the unique Hoatzin in the Cuculiformes, but its taxonomy remains in dispute...

s, such as the Common Cuckoo
Common Cuckoo
The Common Cuckoo is a member of the cuckoo order of birds, Cuculiformes, which includes the roadrunners, the anis and the coucals....

, but this is very rare because the thrush recognizes the cuckoo's non-mimetic eggs. However, the Song Thrush does not demonstrate the same aggression toward the adult Cuckoo that is shown by the Blackbird. The introduced birds in New Zealand, where the cuckoo does not occur, have, over the past 130 years, retained the ability to recognise and reject non-mimetic eggs.

Adult birds may be killed by cat
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

s, Little Owl
Little Owl
The Little Owl is a bird which is resident in much of the temperate and warmer parts of Europe, Asia east to Korea, and north Africa. It is not native to Great Britain, but was first introduced in 1842, and is now naturalised there...

s and Sparrowhawks, and eggs and nestlings are taken by Magpies
European Magpie
The European Magpie, Eurasian Magpie, or Common Magpie, , is a resident breeding bird throughout Europe, much of Asia and northwest Africa. It is one of several birds in the crow family named as magpies, and belongs to the Holarctic radiation of "monochrome" magpies...

, Jays
Eurasian Jay
The Eurasian Jay is a species of bird occurring over a vast region from Western Europe and north-west Africa to the Indian Subcontinent and further to the eastern seaboard of Asia and down into south-east Asia...

, and, where present, Grey Squirrels
Eastern Gray Squirrel
The eastern gray squirrel is a tree squirrel in the genus Sciurus native to the eastern and midwestern United States, and to the southerly portions of the eastern provinces of Canada...

. As with other passerine birds, parasites are common, and include endoparasites, such as the nematode
Nematode
The nematodes or roundworms are the most diverse phylum of pseudocoelomates, and one of the most diverse of all animals. Nematode species are very difficult to distinguish; over 28,000 have been described, of which over 16,000 are parasitic. It has been estimated that the total number of nematode...

 Splendidofilaria (Avifilaria) mavis whose specific name mavis derives from this thrush. A Russian study of blood parasites showed that all the Fieldfares, Redwings and Song Thrushes sampled carried haematozoans, particularly Haemoproteus and Trypanosoma
Trypanosoma
Trypanosoma is a genus of kinetoplastids , a monophyletic group of unicellular parasitic flagellate protozoa. The name is derived from the Greek trypano and soma because of their corkscrew-like motion. All trypanosomes are heteroxenous and are transmitted via a vector...

. Ixodes
Ixodes
Ixodes is a genus of hard-bodied ticks . It includes important disease vectors of animals and humans . Some ticks in this genus may transmit the pathogenic bacterium Borrelia burgdorferi responsible for causing Lyme disease...

ticks are also common, and can carry pathogens, including tick-borne encephalitis
Encephalitis
Encephalitis is an acute inflammation of the brain. Encephalitis with meningitis is known as meningoencephalitis. Symptoms include headache, fever, confusion, drowsiness, and fatigue...

 in forested areas of central and eastern Europe and Russia, and, more widely, Borrelia
Borrelia
Borrelia is a genus of bacteria of the spirochete phylum. It causes borreliosis, a zoonotic, vector-borne disease transmitted primarily by ticks and some by lice, depending on the species...

bacteria
Bacteria
Bacteria are a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria have a wide range of shapes, ranging from spheres to rods and spirals...

. Some species of Borrelia cause Lyme disease
Lyme disease
Lyme disease, or Lyme borreliosis, is an emerging infectious disease caused by at least three species of bacteria belonging to the genus Borrelia. Borrelia burgdorferi sensu stricto is the main cause of Lyme disease in the United States, whereas Borrelia afzelii and Borrelia garinii cause most...

, and ground-feeding birds like the Song Thrush may act as a reservoir for the disease.

Feeding

The Song Thrush is omnivorous, eating a wide range of invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...

s, especially earthworm
Earthworm
Earthworm is the common name for the largest members of Oligochaeta in the phylum Annelida. In classical systems they were placed in the order Opisthopora, on the basis of the male pores opening posterior to the female pores, even though the internal male segments are anterior to the female...

s and snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

s, as well as soft fruit and berries
Berry
The botanical definition of a berry is a fleshy fruit produced from a single ovary. Grapes are an example. The berry is the most common type of fleshy fruit in which the entire ovary wall ripens into an edible pericarp. They may have one or more carpels with a thin covering and fleshy interiors....

. Like its relative, the Blackbird, the Song Thrush finds animal prey by sight, has a run-and-stop hunting technique on open ground, and will rummage through leaf-litter seeking potential food items.

Snails are especially important when drought or hard weather makes it difficult to find other food. The thrush often uses a favourite stone as an "anvil" on which to smash the snail before extracting the soft body and invariably wiping it on the ground before consumption. Young birds initially flick objects and attempt to play with them until they learn to use anvils as tools to smash snails. The nestlings are mainly fed on animal food such as worms, slug
Slug
Slug is a common name that is normally applied to any gastropod mollusc that lacks a shell, has a very reduced shell, or has a small internal shell...

s, snails and insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

 larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e.

The Grove Snail
Grove snail
The grove snail or brown-lipped snail is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc...

 (Cepaea nemoralis) is regularly eaten by the Song Thrush, and its polymorphic
Polymorphism (biology)
Polymorphism in biology occurs when two or more clearly different phenotypes exist in the same population of a species — in other words, the occurrence of more than one form or morph...

 shell patterns have been suggested as evolutionary responses to reduce predation; however, Song Thrushes may not be the only selective force involved.

Status

The Song Thrush has an extensive range, estimated at 10 million square kilometres (3.8 million square miles), and a large population, with an estimated 40 to 71 million individuals in Europe alone.

In the western Palaearctic, there is evidence of population decline, but at a level below the threshold required for global conservation concern (i.e., a reduction in numbers of more than 30% in ten years or three generations) and the IUCN Red List
IUCN Red List
The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

 categorises this species as of "Least Concern
Least Concern
Least Concern is an IUCN category assigned to extant taxon or lower taxa which have been evaluated but do not qualify for any other category. As such they do not qualify as threatened, Near Threatened, or Conservation Dependent...

". In Great Britain and the Netherlands, there has been a more than 50% decline in population and the Song Thrush is included in regional Red List
Regional Red List
A Regional Red List is a report of the threatened status of species within a certain country or region. It is based on the IUCN Red List of Threatened Species, an inventory of the conservation status of species on a global scale...

s. The decreases are greatest in farmlands (73% since the mid 1970s) and believed to be due to changes in agricultural practices in recent decades. The precise reasons for the decline are not known but may be related to the loss of hedgerows, a move to sowing crops in autumn rather than spring, and possibly the increased use of pesticides. These changes may have reduced the availability of food and of nest sites. In gardens, the use of poison bait to control slugs and snails may pose a threat and in urban areas, some thrushes are killed
Road Kill
Road Kill is a pair of live albums released by Celtic rock band Seven Nations in 1998. According to the band, the discs were meant to portray the band's live act realistically, and to preserve "the intensity and energy that make our concerts so much fun both for us and our audiences." The band's...

 while using the hard surface of roads to smash snails.

In culture

The Song Thrush's characteristic song, with melodic phrases repeated twice or more, is described by the nineteenth-century British poet Robert Browning
Robert Browning
Robert Browning was an English poet and playwright whose mastery of dramatic verse, especially dramatic monologues, made him one of the foremost Victorian poets.-Early years:...

 in his poem Home Thoughts, from Abroad:

That's the wise thrush; he sings each song twice over,

Lest you should think he never could recapture

The first fine careless rapture!

The song also inspired the nineteenth-century British writer Thomas Hardy
Thomas Hardy
Thomas Hardy, OM was an English novelist and poet. While his works typically belong to the Naturalism movement, several poems display elements of the previous Romantic and Enlightenment periods of literature, such as his fascination with the supernatural.While he regarded himself primarily as a...

, who spoke in Darkling Thrush of the bird's "full-hearted song evensong/Of joy illimited", but twentieth-century British poet Ted Hughes
Ted Hughes
Edward James Hughes OM , more commonly known as Ted Hughes, was an English poet and children's writer. Critics routinely rank him as one of the best poets of his generation. Hughes was British Poet Laureate from 1984 until his death.Hughes was married to American poet Sylvia Plath, from 1956 until...

 in Thrushes concentrated on its hunting prowess: "Nothing but bounce and/stab/and a ravening second". Nineteenth-century Welsh poet Edward Thomas
Edward Thomas (poet)
Philip Edward Thomas was an Anglo-Welsh writer of prose and poetry. He is commonly considered a war poet, although few of his poems deal directly with his war experiences. Already an accomplished writer, Thomas turned to poetry only in 1914...

 wrote 15 poems concerning Blackbirds or thrushes, including The Thrush:

I hear the thrush, and I see

Him alone at the end of the lane

Near the bare poplar's tip,

Singing continuously.

In The Tables Turned, Romantic
Romanticism
Romanticism was an artistic, literary and intellectual movement that originated in the second half of the 18th century in Europe, and gained strength in reaction to the Industrial Revolution...

 poet William Wordsworth
William Wordsworth
William Wordsworth was a major English Romantic poet who, with Samuel Taylor Coleridge, helped to launch the Romantic Age in English literature with the 1798 joint publication Lyrical Ballads....

 references the Song Thrush, writing

Hark, how blithe the throstle sings

And he is no mean preacher

Come forth into the light of things

Let Nature be your teacher


The Song Thrush is the emblem of West Bromwich Albion
West Bromwich Albion F.C.
West Bromwich Albion Football Club, also known as West Brom, The Baggies, The Throstles, Albion or WBA, are an English Premier League association football club based in West Bromwich in the West Midlands...

 Football Club, chosen because the public house
Public house
A public house, informally known as a pub, is a drinking establishment fundamental to the culture of Britain, Ireland, Australia and New Zealand. There are approximately 53,500 public houses in the United Kingdom. This number has been declining every year, so that nearly half of the smaller...

 in which the team used to change kept a pet thrush in a cage. It also gave rise to Albion's early nickname, The Throstles.

As food

Thrushes have been trapped for food from as far back as 12,000 years ago and an early reference is found in the Odyssey
Odyssey
The Odyssey is one of two major ancient Greek epic poems attributed to Homer. It is, in part, a sequel to the Iliad, the other work ascribed to Homer. The poem is fundamental to the modern Western canon, and is the second—the Iliad being the first—extant work of Western literature...

: "Then, as doves or thrushes beating their spread wings against some snare rigged up in thickets—flying in for a cozy nest but a grisly bed receives them." Hunting continues today around the Mediterranean, but is not believed to be a major factor in this species’ decline in parts of its range.

In Spain, this species is normally caught as it migrates through the country, often using birdlime
Birdlime
Birdlime is an adhesive substance used in trapping birds. It is spread on a branch or twig, upon which a bird may land and be caught. Its use is illegal in many jurisdictions....

 which, although banned by the European Union, is still tolerated and permitted in the Valencian Community
Valencian Community
The Valencian Community is an autonomous community of Spain located in central and south-eastern Iberian Peninsula. Its capital and largest city is Valencia...

. In 2003 and 2004 the EU
European Union
The European Union is an economic and political union of 27 independent member states which are located primarily in Europe. The EU traces its origins from the European Coal and Steel Community and the European Economic Community , formed by six countries in 1958...

 tried, but failed, to stop this practice in the Valencian region.

Pet

Up to at least the nineteenth century the Song Thrush was kept as a cage bird because of its melodious voice. As with hunting, there is little evidence that the taking of wild birds for aviculture
Aviculture
Aviculture is the practice of keeping and breeding birds and the culture that forms around it. Aviculture is generally focused on not only the raising and breeding of birds, but also on preserving avian habitat, and public awareness campaigns....

has had a significant effect on wild populations.

External links

The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
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