Invertebrate
Overview
An invertebrate is an animal
Animal
Animals are a major group of multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and...

 without a backbone. The group includes 97% of all animal species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 – all animals except those in the chordate
Chordate
Chordates are animals which are either vertebrates or one of several closely related invertebrates. They are united by having, for at least some period of their life cycle, a notochord, a hollow dorsal nerve cord, pharyngeal slits, an endostyle, and a post-anal tail...

 subphylum
Subphylum
In life, a subphylum is a taxonomic rank intermediate between phylum and superclass. The rank of subdivision in plants and fungi is equivalent to subphylum.Not all phyla are divided into subphyla...

 Vertebrata
Vertebrate
Vertebrates are animals that are members of the subphylum Vertebrata . Vertebrates are the largest group of chordates, with currently about 58,000 species described. Vertebrates include the jawless fishes, bony fishes, sharks and rays, amphibians, reptiles, mammals, and birds...

 (fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

, amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

s, reptile
Reptile
Reptiles are members of a class of air-breathing, ectothermic vertebrates which are characterized by laying shelled eggs , and having skin covered in scales and/or scutes. They are tetrapods, either having four limbs or being descended from four-limbed ancestors...

s, bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s, and mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

s).

Invertebrates form a paraphyletic group. Given a common multicellular, eukaryotic ancestor, all contained phyla are invertebrates along with two of the three subphyla
Subphylum
In life, a subphylum is a taxonomic rank intermediate between phylum and superclass. The rank of subdivision in plants and fungi is equivalent to subphylum.Not all phyla are divided into subphyla...

 in Phylum Chordata
Chordate
Chordates are animals which are either vertebrates or one of several closely related invertebrates. They are united by having, for at least some period of their life cycle, a notochord, a hollow dorsal nerve cord, pharyngeal slits, an endostyle, and a post-anal tail...

: Tunicata
Tunicate
Tunicates, also known as urochordates, are members of the subphylum Tunicata, previously known as Urochordata, a group of underwater saclike filter feeders with incurrent and excurrent siphons that is classified within the phylum Chordata. While most tunicates live on the ocean floor, others such...

 and Cephalochordata
Lancelet
The lancelets , also known as amphioxus, are the modern representatives of the subphylum Cephalochordata, formerly thought to be the sister group of the craniates. They are usually found buried in sand in shallow parts of temperate or tropical seas. In Asia, they are harvested commercially as food...

. These two, plus all the other known invertebrates, have only one cluster of Hox genes, while the vertebrates have duplicated their original cluster more than once.

Within palaeozoology
Paleozoology
Paleozoology, also spelled as palaeozoology , is the branch of paleontology or paleobiology dealing with the recovery and identification of multicellular animal remains from geological contexts, and the use of these fossils in the reconstruction of prehistoric environments and ancient...

 and palaeobiology
Paleobiology
Paleobiology is a growing and comparatively new discipline which combines the methods and findings of the natural science biology with the methods and findings of the earth science paleontology...

, invertebrates are often studied within the fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

 discipline called invertebrate palaeontology
Invertebrate paleontology
Invertebrate paleontology is sometimes described as Invertebrate paleozoology or Invertebrate paleobiology....

.
The word invertebrate comes from the word vertebrate, with the prefix in- attached to it.
 
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