Goatfish
Encyclopedia
Goatfishes are tropical marine perciform fish
of the family Mullidae. Seldom found in brackish waters, goatfish are most associated with the reef
s of the Atlantic
, Indian
, and Pacific Ocean
s. The goatfish are sometimes called the red mullets as opposed to the Mugilidae, the grey mullets, though that name is usually reserved for the red mullet
s of the genus Mullus
of the Mediterranean
. Within the family are approximately six genera and 55 species.
. Rather, goatfish are valued food fish in many countries. The largest species, the dash-and-dot goatfish
(Parupeneus barberinus) grows to 55 centimetres in length; most species are less than half this size. Their bodies are deep and elongate with forked tail fins and widely separated dorsal fins.
Perhaps the goatfish's unpopularity among fishkeepers can be attributed to its feeding habits: Goatfish are tireless benthic feeders, using a pair of long chemosensory barbel
s ("whiskers") protruding from their chins to rifle through the sediments in search of a meal. Like goats, they seek anything edible; worm
s, crustaceans, molluscs and other small invertebrate
s are staples.
By day, many goatfish will form large inactive (non-feeding) schools: these aggregates may contain both conspecifics and heterospecifics. For example, the yellowfin goatfish (Mulloidichthys vanicolensis) of the Red Sea
and Hawaii
is often seen congregating with blue-striped snapper
s (Lutjanus kasmira). With such mixed company, the yellowfins will actually change their coloration to match that of the snappers.
By night the schools disperse and individual goatfish head their separate ways to loot the sands. Other nocturnal feeders will shadow the active goatfish, waiting patiently for any overlooked morsels. Goatfish stay within the shallows, going no deeper than about 110 metres. Some species, such as the freckled goatfish (Upeneus tragula) of East Africa
, have been known to enter estuaries
and river
s, although not to any great extent.
All goatfish have the ability to change their coloration depending on their current activity. One notable example, the diurnal goldsaddle goatfish
(Parupeneus cyclostomus) will change from a lemon-yellow to a pale cream whilst feeding. Diurnal species also tend to be solitary, but will school as juveniles.
Goatfish are pelagic spawners; that is, they release many buoyant eggs into the water which become part of the plankton
. The eggs float freely with the currents until hatching. The postlarva floats in surface waters until it reaches around 5 or 6 centimetres in length, when it takes on the adult, bottom-feeding, lifestyle.
In ancient Rome until the end of the second century A.D. two species of goatfish (Mullus barbatus
and Mullus surmuletus
) were highly sought-after and expensive, however not as a delicacy, but for aesthetic pleasure, since the fish assume a variety of colors and shades also during death (see also above). Therefore it was paramount to serve live fish and let it die before the eyes of the guests.
FishBase lists 67 species in six genera:
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
of the family Mullidae. Seldom found in brackish waters, goatfish are most associated with the reef
Reef
In nautical terminology, a reef is a rock, sandbar, or other feature lying beneath the surface of the water ....
s of the Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...
, Indian
Indian Ocean
The Indian Ocean is the third largest of the world's oceanic divisions, covering approximately 20% of the water on the Earth's surface. It is bounded on the north by the Indian Subcontinent and Arabian Peninsula ; on the west by eastern Africa; on the east by Indochina, the Sunda Islands, and...
, and Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...
s. The goatfish are sometimes called the red mullets as opposed to the Mugilidae, the grey mullets, though that name is usually reserved for the red mullet
Red mullet
The red mullets or surmullets are two species of goatfish, Mullus barbatus and Mullus surmuletus, found in the Mediterranean Sea, east North Atlantic Ocean, and the Black Sea. Both "red mullet" and "surmullet" can also refer to the Mullidae in general.Though they can easily be distinguished—M...
s of the genus Mullus
Mullus
Mullus known as Goatfishes or Mullets is subtropical marine genus of perciform fish of the family Mullidae. Occurs mainly in the Southwest Atlantic near the South American coast and in the Eastern Atlantic including the Mediterranean and the Black Sea. These fish are bottom dwelling over sands and...
of the Mediterranean
Mediterranean Sea
The Mediterranean Sea is a sea connected to the Atlantic Ocean surrounded by the Mediterranean region and almost completely enclosed by land: on the north by Anatolia and Europe, on the south by North Africa, and on the east by the Levant...
. Within the family are approximately six genera and 55 species.
Description and habits
Many species of goatfish are conspicuously coloured; however, they are not popular in aquariaAquarium
An aquarium is a vivarium consisting of at least one transparent side in which water-dwelling plants or animals are kept. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, marine mammals, turtles, and aquatic plants...
. Rather, goatfish are valued food fish in many countries. The largest species, the dash-and-dot goatfish
Dash-and-dot goatfish
The dash-and-dot goatfish, Parupeneus barberinus, is a goatfish of the family Mullidae.-Description:The dash-and-dot goatfish has the twin chin barbels typical of goatfishes....
(Parupeneus barberinus) grows to 55 centimetres in length; most species are less than half this size. Their bodies are deep and elongate with forked tail fins and widely separated dorsal fins.
Perhaps the goatfish's unpopularity among fishkeepers can be attributed to its feeding habits: Goatfish are tireless benthic feeders, using a pair of long chemosensory barbel
Barbel (anatomy)
A barbel on a fish is a slender, whiskerlike tactile organ near the mouth. Fish that have barbels include the catfish, the carp, the goatfish, sturgeon, the zebrafish and some species of shark...
s ("whiskers") protruding from their chins to rifle through the sediments in search of a meal. Like goats, they seek anything edible; worm
Worm
The term worm refers to an obsolete taxon used by Carolus Linnaeus and Jean-Baptiste Lamarck for all non-arthropod invertebrate animals, and stems from the Old English word wyrm. Currently it is used to describe many different distantly-related animals that typically have a long cylindrical...
s, crustaceans, molluscs and other small invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...
s are staples.
By day, many goatfish will form large inactive (non-feeding) schools: these aggregates may contain both conspecifics and heterospecifics. For example, the yellowfin goatfish (Mulloidichthys vanicolensis) of the Red Sea
Red Sea
The Red Sea is a seawater inlet of the Indian Ocean, lying between Africa and Asia. The connection to the ocean is in the south through the Bab el Mandeb strait and the Gulf of Aden. In the north, there is the Sinai Peninsula, the Gulf of Aqaba, and the Gulf of Suez...
and Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...
is often seen congregating with blue-striped snapper
Bluestripe snapper
The bluestripe snapper, Lutjanus kasmira, is a common species of marine fish of the snapper family Lutjanidae...
s (Lutjanus kasmira). With such mixed company, the yellowfins will actually change their coloration to match that of the snappers.
By night the schools disperse and individual goatfish head their separate ways to loot the sands. Other nocturnal feeders will shadow the active goatfish, waiting patiently for any overlooked morsels. Goatfish stay within the shallows, going no deeper than about 110 metres. Some species, such as the freckled goatfish (Upeneus tragula) of East Africa
East Africa
East Africa or Eastern Africa is the easterly region of the African continent, variably defined by geography or geopolitics. In the UN scheme of geographic regions, 19 territories constitute Eastern Africa:...
, have been known to enter estuaries
Estuary
An estuary is a partly enclosed coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea....
and river
River
A river is a natural watercourse, usually freshwater, flowing towards an ocean, a lake, a sea, or another river. In a few cases, a river simply flows into the ground or dries up completely before reaching another body of water. Small rivers may also be called by several other names, including...
s, although not to any great extent.
All goatfish have the ability to change their coloration depending on their current activity. One notable example, the diurnal goldsaddle goatfish
Goldsaddle goatfish
Also known as the Yellowsaddle goatfish.-Description:May be up to 50cm long, has long barbels; the body is usually yellow with a bluish sheen to the dorsal part but there is a golden yellow variant.-Habitat:...
(Parupeneus cyclostomus) will change from a lemon-yellow to a pale cream whilst feeding. Diurnal species also tend to be solitary, but will school as juveniles.
Goatfish are pelagic spawners; that is, they release many buoyant eggs into the water which become part of the plankton
Plankton
Plankton are any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. That is, plankton are defined by their ecological niche rather than phylogenetic or taxonomic classification...
. The eggs float freely with the currents until hatching. The postlarva floats in surface waters until it reaches around 5 or 6 centimetres in length, when it takes on the adult, bottom-feeding, lifestyle.
In ancient Rome until the end of the second century A.D. two species of goatfish (Mullus barbatus
Mullus barbatus
Mullus barbatus is a species of goatfish found in the Mediterranean Sea, east North Atlantic Ocean from Scandinavia to Senegal, and the Black Sea....
and Mullus surmuletus
Mullus surmuletus
Mullus surmuletus, the striped red mullet, is a species of goatfish found in the Mediterranean Sea, east North Atlantic Ocean, and the Black Sea....
) were highly sought-after and expensive, however not as a delicacy, but for aesthetic pleasure, since the fish assume a variety of colors and shades also during death (see also above). Therefore it was paramount to serve live fish and let it die before the eyes of the guests.
Species
- Genus MulloidichthysMulloidichthysMulloidichthys or Goatfishes is a tropical marine genus of perciform fish of the family Mullidae. Occurs mainly in coral and rocky reefs of the Atlantic, Indian, and Pacific Oceans. 6 species are recognized in this genus.-Species:...
- Mexican goatfish, Mulloidichthys dentatus (GillTheodore GillTheodore Nicholas Gill was an American ichthyologist, mammalogist, malacologist and librarian.Born and educated in New York City under private tutors, Gill early showed interest in natural history. He was associated with J...
, 1862). - Yellowstripe goatfish, Mulloidichthys flavolineatus (Lacépède, 1801).
- Yellow goatfishYellow goatfishThe yellow goatfish , also known as yellowsaddle or goldsaddle goatfish , is a white to pale pink fish with a yellow stripe from the eye to the caudal peduncle.-Description:...
, Mulloidichthys martinicus (CuvierGeorges CuvierGeorges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...
, 1829). - Mimic goatfish, Mulloidichthys mimicus Randall & Guézé, 1980.
- Orange goatfish, Mulloidichthys pfluegeri (SteindachnerFranz SteindachnerFranz Steindachner was an Austrian zoologist.- Work and career :Being interested in natural history, Steindachner took up the study of fossil fishes on the recommendation of his friend Eduard Suess...
, 1900). - Yellowfin goatfish, Mulloidichthys vanicolensis (ValenciennesAchille ValenciennesAchille Valenciennes was a French zoologist.Valenciennes was born in Paris, and studied under Georges Cuvier. Valenciennes' study of parasitic worms in humans made an important contribution to the study of parasitology...
, 1831).
- Mexican goatfish, Mulloidichthys dentatus (Gill
- Genus MullusMullusMullus known as Goatfishes or Mullets is subtropical marine genus of perciform fish of the family Mullidae. Occurs mainly in the Southwest Atlantic near the South American coast and in the Eastern Atlantic including the Mediterranean and the Black Sea. These fish are bottom dwelling over sands and...
- Argentine goatfish, Mullus argentinae Hubbs & Marini, 1933.
- Red goatfish, Mullus auratus JordanDavid Starr JordanDavid Starr Jordan, Ph.D., LL.D. was a leading eugenicist, ichthyologist, educator and peace activist. He was president of Indiana University and Stanford University.-Early life and education:...
& Gilbert, 1882. - The red mulletRed mulletThe red mullets or surmullets are two species of goatfish, Mullus barbatus and Mullus surmuletus, found in the Mediterranean Sea, east North Atlantic Ocean, and the Black Sea. Both "red mullet" and "surmullet" can also refer to the Mullidae in general.Though they can easily be distinguished—M...
s- Mullus barbatus barbatus LinnaeusCarolus LinnaeusCarl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...
, 1758. - Mullus barbatus ponticus Essipov, 1927.
- Striped red mullet, Mullus surmuletusMullus surmuletusMullus surmuletus, the striped red mullet, is a species of goatfish found in the Mediterranean Sea, east North Atlantic Ocean, and the Black Sea....
LinnaeusCarolus LinnaeusCarl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...
, 1758.
- Mullus barbatus barbatus Linnaeus
- Genus ParupeneusParupeneusParupeneus or Goatfishes is tropical marine genus of perciform fish of the family Mullidae. Seldom found in brackish waters, goatfish are most associated with the reefs of the Atlantic, Indian, and Pacific Oceans. 28 species are recognized in this genus.*) NE - Not Evaluated. T) Type species....
- Bicolor goatfish, Parupeneus barberinoidesParupeneus barberinoidesParupeneus barberinoides is a Goatfish from the Western Pacific. It occasionally makes its way into the aquarium trade. It grows to a size of 30cm in length....
(BleekerPieter BleekerPieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....
, 1852). - Dash-and-dot goatfishDash-and-dot goatfishThe dash-and-dot goatfish, Parupeneus barberinus, is a goatfish of the family Mullidae.-Description:The dash-and-dot goatfish has the twin chin barbels typical of goatfishes....
, Parupeneus barberinus (Lacépède, 1801). - Pointed goatfish, Parupeneus biaculeatus (RichardsonJohn Richardson (naturalist)Sir John Richardson was a Scottish naval surgeon, naturalist and arctic explorer.Richardson was born at Dumfries. He studied medicine at Edinburgh University, and became a surgeon in the navy in 1807. He traveled with John Franklin in search of the Northwest Passage on the Coppermine Expedition of...
, 1846). - Yellow-threaded goatfish, Parupeneus chrysonemus (JordanDavid Starr JordanDavid Starr Jordan, Ph.D., LL.D. was a leading eugenicist, ichthyologist, educator and peace activist. He was president of Indiana University and Stanford University.-Early life and education:...
& Evermann, 1903). - Yellow striped goatfish, Parupeneus chrysopleuron (TemminckCoenraad Jacob TemminckCoenraad Jacob Temminck was a Dutch aristocrat and zoologist.Temminck was the first director of the National Natural History Museum at Leiden from 1820 until his death. His Manuel d'ornithologie, ou Tableau systematique des oiseaux qui se trouvent en Europe was the standard work on European birds...
& SchlegelHermann SchlegelHermann Schlegel was a German ornithologist and herpetologist.-Early life and education:Schlegel was born at Altenburg, the son of a brassfounder. His father collected butterflies, which stimulated Schlegel's interest in natural history...
, 1843). - Whitesaddle goatfish, Parupeneus ciliatus (Lacépède, 1802).
- Parupeneus crassilabris (ValenciennesAchille ValenciennesAchille Valenciennes was a French zoologist.Valenciennes was born in Paris, and studied under Georges Cuvier. Valenciennes' study of parasitic worms in humans made an important contribution to the study of parasitology...
, 1831). - Goldsaddle goatfishGoldsaddle goatfishAlso known as the Yellowsaddle goatfish.-Description:May be up to 50cm long, has long barbels; the body is usually yellow with a bluish sheen to the dorsal part but there is a golden yellow variant.-Habitat:...
, Parupeneus cyclostomus (Lacépède, 1801). - Red Sea goatfish, Parupeneus forsskali (Fourmanoir & Guézé, 1976).
- Cinnabar goatfish, Parupeneus heptacanthus (Lacépède, 1802).
- Indian goatfish, Parupeneus indicus (ShawGeorge ShawGeorge Shaw was an English botanist and zoologist.Shaw was born at Bierton, Buckinghamshire and was educated at Magdalen Hall, Oxford, receiving his M.A. in 1772. He took up the profession of medical practitioner. In 1786 he became the assistant lecturer in botany at Oxford University...
, 1803). - Parupeneus insularis Randall & Myers, 2002.
- Parupeneus jansenii (BleekerPieter BleekerPieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....
, 1856). - Parupeneus louise Randall, 2004.
- Longbarbel goatfish, Parupeneus macronemus (Lacépède, 1801).
- Pearly goatfish, Parupeneus margaritatus Randall & Guézé, 1984.
- Parupeneus moffitti Randall & Myers, 1993.
- Manybar goatfish, Parupeneus multifasciatusParupeneus multifasciatusParupeneus multifasciatus is a Goatfish from the Pacific Ocean. It occasionally makes its way into the aquarium trade. It grows to a size of 35cm in length....
(QuoyJean René Constant QuoyJean René Constant Quoy was a French zoologist.Along with Joseph Paul Gaimard he served as naturalist aboard La Coquille under Louis Isidore Duperrey during its circumnavigation of the globe , and the Astrolabe under the command of Jules Dumont d'Urville...
& GaimardJoseph Paul GaimardJoseph Paul Gaimard was a French naval surgeon and naturalist.Along with Jean René Constant Quoy he served as naturalist on the ships L'Uranie under Louis de Freycinet 1817-1820, and L'Astrolabe under Jules Dumont d'Urville 1826-1829...
, 1824). - Parupeneus orientalis (Fowler, 1933).
- Sidespot goatfish, Parupeneus pleurostigma (Bennett, 1831).
- Parupeneus porphyreus (Jenkins, 1903).
- Parupeneus posteli Fourmanoir & Guézé, 1967.
- Parupeneus procerigena Kim & Amaoka, 2001.
- Rosy goatfish, Parupeneus rubescens (Lacépède, 1801).
- Black-spot goatfishBlack-spot goatfishThe black-spot goatfish, Parupeneus signatus, is a goatfish of the family Mullidae, found in the western Pacific ocean, from north east New Zealand to New Guinea, in shallow waters...
, Parupeneus signatus (Günther, 1867). - Doublebar goatfish, Parupeneus trifasciatus (Lacépède, 1801).
- Bicolor goatfish, Parupeneus barberinoides
- Genus PseudupeneusPseudupeneusPseudupeneus known as Goatfishes is tropical-subtropical marine genus of perciform fish of the family Mullidae. Occurs mainly in the Atlantic and Eastern Pacific. These fish inhabit mainly the coastal waters of continental shelfs, but can be found in deep waters as well. 3 species are recognized.*)...
- Bigscale goatfish, Pseudupeneus grandisquamis (GillTheodore GillTheodore Nicholas Gill was an American ichthyologist, mammalogist, malacologist and librarian.Born and educated in New York City under private tutors, Gill early showed interest in natural history. He was associated with J...
, 1863). - Spotted goatfish, Pseudupeneus maculatus (BlochMarcus Elieser BlochMarcus Elieser Bloch was a German medical doctor and naturalist. He is generally considered one of the most important ichthyologists of the 18th century.- Life :...
, 1793). - West African goatfish, Pseudupeneus prayensis (CuvierGeorges CuvierGeorges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...
, 1829).
- Bigscale goatfish, Pseudupeneus grandisquamis (Gill
- Genus UpeneichthysUpeneichthysUpeneichthys or Goatfishes is Subtropical marine genus of perciform fish of the family Mullidae. Occurs mainly in Eastern Indian Ocean and Southwest Pacific, near Australians shores. 3 species are recognized in this genus.-Species:...
- Blue-lined goatfishBlue-lined goatfishThe blue-lined goatfish or red mullet, Upeneichthys lineatus, is a goatfish of the family Mullidae, found off eastern Australia and around New Zealand at depths of between 5 and 100 m...
, Upeneichthys lineatus (BlochMarcus Elieser BlochMarcus Elieser Bloch was a German medical doctor and naturalist. He is generally considered one of the most important ichthyologists of the 18th century.- Life :...
& SchneiderJohann Gottlob SchneiderJohann Gottlob Theaenus Schneider was a German classicist and naturalist.-Biography:Schneider was born at Collm in Saxony...
, 1801). - Upeneichthys stotti Platell, Potter & Clarke, 1998.
- Southern goatfish, Upeneichthys vlamingiiUpeneichthys vlamingiiUpeneichthys vlamingii or Bluespotted Goatfish, is a Bottom dwelling marine fish which inhabits the shallow costal waters along the southern Australian coastline. They grow to sizes of 30–40 cm....
(CuvierGeorges CuvierGeorges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...
, 1829).
- Blue-lined goatfish
- Genus UpeneusUpeneusUpeneus or Goatfishes is Subtropical marine genus of perciform fish of the family Mullidae. Occurs mainly in Eastern Indian Ocean and Southwest Pacific. 28 species are recognized in this genus.-Species:...
- Band-tail goatfish, Upeneus arge JordanDavid Starr JordanDavid Starr Jordan, Ph.D., LL.D. was a leading eugenicist, ichthyologist, educator and peace activist. He was president of Indiana University and Stanford University.-Early life and education:...
& Evermann, 1903. - Asymmetrical goatfish, Upeneus asymmetricus Lachner, 1954.
- Upeneus australiae Kim & Nakaya, 2002.
- Upeneus crosnieri Fourmanoir & Guézé, 1967.
- Upeneus davidaromi Golani, 2001.
- Gilded goatfish, Upeneus doriae (Günther, 1869).
- Upeneus filifer (OgilbyWilliam OgilbyWilliam Ogilby was an Irish barrister and naturalist.Ogilby was honorary secretary of the Zoological Society of London from 1839 to 1846....
, 1910). - Upeneus francisi Randall & Guézé, 1992.
- Upeneus guttatus (Day, 1868).
- Bensasi goatfish, Upeneus japonicus (HouttuynMartinus HouttuynMaarten Houttuyn or Houttuijn , Latinised as Martinus Houttuyn, was a Dutch naturalist.Houttuyn was born in Hoorn, studied medicine in Leiden and moved to Amsterdam in 1753. He published many books on natural history. His areas of interest encompassed Pteridophytes, Bryophytes and Spermatophytes...
, 1782). - Dark-barred goatfish, Upeneus luzonius JordanDavid Starr JordanDavid Starr Jordan, Ph.D., LL.D. was a leading eugenicist, ichthyologist, educator and peace activist. He was president of Indiana University and Stanford University.-Early life and education:...
& Seale, 1907. - Upeneus mascareinsis Fourmanoir & Guézé, 1967.
- Goldband goatfish, Upeneus moluccensis (BleekerPieter BleekerPieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....
, 1855). - Upeneus mouthami Randall & Kulbicki, 2006.
- Dwarf goatfish, Upeneus parvus PoeyFelipe Poey-Biography:Poey was born in Havana, the son of French and Spanish parents. He spent several years of his life in Pau then studied law in Madrid. He became a lawyer in Spain but was forced to leave due to his liberal ideas, returning to Cuba in 1823. He began to concentrate on the study of the...
, 1852. - Por's goatfish, Upeneus pori Ben-Tuvia & Golani, 1989.
- Upeneus quadrilineatus Cheng & Wang, 1963.
- Upeneus subvittatus (TemminckCoenraad Jacob TemminckCoenraad Jacob Temminck was a Dutch aristocrat and zoologist.Temminck was the first director of the National Natural History Museum at Leiden from 1820 until his death. His Manuel d'ornithologie, ou Tableau systematique des oiseaux qui se trouvent en Europe was the standard work on European birds...
& SchlegelHermann SchlegelHermann Schlegel was a German ornithologist and herpetologist.-Early life and education:Schlegel was born at Altenburg, the son of a brassfounder. His father collected butterflies, which stimulated Schlegel's interest in natural history...
, 1843). - Sulphur goatfish, Upeneus sulphureus CuvierGeorges CuvierGeorges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...
, 1829. - Ochre-banded goatfish, Upeneus sundaicus (BleekerPieter BleekerPieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....
, 1855). - Finstripe goatfish, Upeneus taeniopterus CuvierGeorges CuvierGeorges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...
, 1829. - Freckled goatfish, Upeneus tragula RichardsonJohn Richardson (naturalist)Sir John Richardson was a Scottish naval surgeon, naturalist and arctic explorer.Richardson was born at Dumfries. He studied medicine at Edinburgh University, and became a surgeon in the navy in 1807. He traveled with John Franklin in search of the Northwest Passage on the Coppermine Expedition of...
, 1846. - Yellowstriped goatfish, Upeneus vittatus (ForsskålPeter ForsskålPeter Forsskål, sometimes spelled Pehr Forsskål, Peter Forskaol, Petrus Forskål or Pehr Forsskåhl, was a Swedish explorer, orientalist, naturalist and an apostle of Carl Linnaeus.-Early life:...
, 1775). - Upeneus xanthogrammus Gilbert, 1892.
- Band-tail goatfish, Upeneus arge Jordan