European White Elm
Encyclopedia
Ulmus laevis Pall., the European White Elm, Fluttering Elm, Spreading Elm and, in the USA, Russian Elm, is a large deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 native to Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, from France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

 northeast to southern Finland
Finland
Finland , officially the Republic of Finland, is a Nordic country situated in the Fennoscandian region of Northern Europe. It is bordered by Sweden in the west, Norway in the north and Russia in the east, while Estonia lies to its south across the Gulf of Finland.Around 5.4 million people reside...

, east as far as the Urals, and southeast to Bulgaria
Bulgaria
Bulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...

 and the Crimea
Crimea
Crimea , or the Autonomous Republic of Crimea , is a sub-national unit, an autonomous republic, of Ukraine. It is located on the northern coast of the Black Sea, occupying a peninsula of the same name...

; there is also a disjunct population in the Caucasus
Caucasus
The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

. Moreover, a small number of trees found in Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

 is now considered a relict
Relict
A relict is a surviving remnant of a natural phenomenon.* In biology a relict is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas....

 population rather than an introduction by man, and possibly the origin of the European population.

Essentially endemic to alluvial forest, it is rarely encountered at elevations above 400 m. Most commonly found along rivers such as the Volga and Danube
Danube
The Danube is a river in the Central Europe and the Europe's second longest river after the Volga. It is classified as an international waterway....

, it is one of very few elms tolerant of prolonged waterlogged, anoxic
Hypoxia (environmental)
Hypoxia, or oxygen depletion, is a phenomenon that occurs in aquatic environments as dissolved oxygen becomes reduced in concentration to a point where it becomes detrimental to aquatic organisms living in the system...

 ground conditions. The White Elm is allogamous and is most closely related to the American Elm U. americana.

Description

The tree is similar in stature to the Wych Elm
Wych Elm
Ulmus glabra, the Wych elm or Scots elm, has the widest range of the European elm species, from Ireland eastwards to the Urals, and from the Arctic Circle south to the mountains of the Peloponnese in Greece; it is also found in Iran...

, if rather less symmetric, with a looser branch structure and less neatly rounded crown. It typically reaches a height and breadth of > 30 m, with a trunk < 2 m d.b.h
Diameter at breast height
Diameter at breast height, or DBH, is a standard method of expressing the diameter of the trunk or bole of a standing tree. DBH is one of the most common dendrometric measurements....

. The extensive shallow root system ultimately forms distinctive high buttresses around the base of the trunk. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

, alternate, simple ovate
Leaf shape
In botany, leaf shape is characterised with the following terms :* Acicular : Slender and pointed, needle-like* Acuminate : Tapering to a long point...

 with a markedly lop-sided base, < 10 cm long and < 7 cm broad, comparatively thin, often almost papery in texture and very translucent, smooth above with a downy underside. The apetalous wind-pollinated flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s appear before the leaves in early spring, produced in clusters of 15-30; they are 3–4 mm across on 20 mm long stems. In England, trees grown from seed flower in March, commencing at ages of between 7 and 12 years.
The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a winged samara
Samara (fruit)
A samara is a type of fruit in which a flattened wing of fibrous, papery tissue develops from the ovary wall. A samara is a simple dry fruit and indehiscent . It is a winged achene...

 < 15 mm long by 10 mm broad with a ciliate
Ciliate
The ciliates are a group of protozoans characterized by the presence of hair-like organelles called cilia, which are identical in structure to flagella but typically shorter and present in much larger numbers with a different undulating pattern than flagella...

 margin, the single round 5 mm seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

 maturing in late spring. U. laevis sheds its leaves earlier in the autumn than other species of European elm.

The tree is most reliably distinguished from other European elms by the long flower stems, and is most closely related to the American Elm
American Elm
Ulmus americana, generally known as the American Elm or, less commonly, as the White Elm or Water Elm, is a species native to eastern North America, occurring from Nova Scotia west to Alberta and Montana, and south to Florida and central Texas. The American elm is an extremely hardy tree that can...

 U. americana, from which it differs mainly in the irregular crown shape and frequent small sprout stems on the trunk and branches, features which also give the tree a winter silhouette; a useful diagnostic feature at this season.

U. laevis suckers from roots, but not stools. Although the species is protandrous, levels of self-pollination are high

Pests and diseases

Like other European elm
Elm
Elms are deciduous and semi-deciduous trees comprising the genus Ulmus in the plant family Ulmaceae. The dozens of species are found in temperate and tropical-montane regions of North America and Eurasia, ranging southward into Indonesia. Elms are components of many kinds of natural forests...

s, natural populations of the European White Elm have little innate resistance to Dutch elm disease
Dutch elm disease
Dutch elm disease is a disease caused by a member of the sac fungi category, affecting elm trees which is spread by the elm bark beetle. Although believed to be originally native to Asia, the disease has been accidentally introduced into America and Europe, where it has devastated native...

, although research by Cemagref
Cemagref
Cemagref is a public research institute in France focusing on land management issues such as water resources and agricultural technology.- Organisation :...

 has isolated clones able to survive inoculation with the causal fungus, initially losing < 70% of their foliage, but regenerating strongly the following year. Moreover, the tree is not favoured by the vector bark beetles, which colonize it only when there are no other elm alternatives available, an uncommon situation in western Europe. Research in Spain has indicated that it is the presence of a triterpene
Triterpene
Triterpenes are terpenes consisting of six isoprene units and have the molecular formula C30H48.The pentacyclic triterpenes can be classified into lupane, oleanane or ursane groups.Animal- and plant-derived triterpenes exist, such as:*squalene...

, alnulin, that renders the tree bark unattractive to the beetles. Ergo: the tree's decline in western Europe is chiefly owing to woodland clearance in river valleys, not disease.

The species has a slight to moderate susceptibility to Elm Yellows
Elm Yellows
Elm Yellows is a plant disease of elm trees that is spread by leafhoppers or by root grafts. Elm Yellows, also known as Elm Phloem Necrosis, is very aggressive, with no known cure. Elm Yellows occurs in the Eastern United States and southern Ontario in Canada. It is caused by phytoplasmas which...

, but a very low susceptibility to the elm leaf beetle Xanthogaleruca luteola
Xanthogaleruca luteola
Xanthogaleruca luteola, commonly known as the Elm Leaf Beetle, is a serious pest of the elm. Indigenous to Europe, it was accidentally introduced to North America. Both the imagines and larvae feed on the emergent leaves of the elm...

.

Cultivation

Although ideally suited to wet ground conditions, the tree can still grow, albeit more slowly, on drier sites including chalk downland. However, one overriding factor in choosing a site is Exposure. White Elm is comparatively weak-wooded, much more so than Field Elm Ulmus minor, and thus an inappropriate choice for windy locations. In trials in southern England by Butterfly Conservation
Butterfly Conservation
Butterfly Conservation is an insect conservation organisation in the United Kingdom. It is one of the largest insect conservation organisations in the world.-History of the Organisation:...

, young trees of < 5 m height were badly damaged by gusts of 40 knots (75 km/h) in midsummer.

Ulmus laevis is occasionally planted as an ornamental tree. In the UK examples are few and far between but sometimes of great age. Mature specimens survive today amid diseased native elm near Torpoint
Torpoint
Torpoint is a civil parish and town on the Rame Peninsula in southeast Cornwall, England, United Kingdom. It is situated opposite the city of Plymouth across the Hamoaze which is the tidal estuary of the River Tamar....

 in Cornwall
Cornwall
Cornwall is a unitary authority and ceremonial county of England, within the United Kingdom. It is bordered to the north and west by the Celtic Sea, to the south by the English Channel, and to the east by the county of Devon, over the River Tamar. Cornwall has a population of , and covers an area of...

; others can be found in Edinburgh
Edinburgh
Edinburgh is the capital city of Scotland, the second largest city in Scotland, and the eighth most populous in the United Kingdom. The City of Edinburgh Council governs one of Scotland's 32 local government council areas. The council area includes urban Edinburgh and a rural area...

 (The Meadows area and Powderhall Road), London
London
London is the capital city of :England and the :United Kingdom, the largest metropolitan area in the United Kingdom, and the largest urban zone in the European Union by most measures. Located on the River Thames, London has been a major settlement for two millennia, its history going back to its...

 (Riverside Walk, near Royal Botanic Gardens, Kew
Royal Botanic Gardens, Kew
The Royal Botanic Gardens, Kew, usually referred to as Kew Gardens, is 121 hectares of gardens and botanical glasshouses between Richmond and Kew in southwest London, England. "The Royal Botanic Gardens, Kew" and the brand name "Kew" are also used as umbrella terms for the institution that runs...

, and at Peckham
Peckham
Peckham is a district in south London, England, located in the London Borough of Southwark. It is situated south-east of Charing Cross. The area is identified in the London Plan as one of 35 major centres in Greater London...

 and Tooting
Tooting
Tooting is a district in south London, England, located in the London Borough of Wandsworth. It is situated south south-west of Charing Cross. The area is identified in the London Plan as one of 35 major centres in Greater London.-History:...

), Chelmsford
Chelmsford
Chelmsford is the county town of Essex, England and the principal settlement of the borough of Chelmsford. It is located in the London commuter belt, approximately northeast of Charing Cross, London, and approximately the same distance from the once provincial Roman capital at Colchester...

 (beside the Chelmer
River Chelmer
The River Chelmer is a river that flows entirely through the county of Essex, England.The source of the river is near Debden Green, a village near Thaxted. The source of the River Can is also nearby. The River Chelmer flows past Thaxted, south through the district of Uttlesford around the...

 at the Rivermead Campus) [2009], Brighton & Hove
Brighton & Hove
Brighton and Hove is a unitary authority area and city on the south coast of England. It is England's most populous seaside resort.In 1997 Brighton and Hove were joined to form the unitary authority of Brighton and Hove, which was granted city status by Queen Elizabeth II as part of the millennium...

, and near St. Albans.

The species was never widely introduced to the USA, but is represented at several arboreta. In the Far East, the tree has been planted in Xinjiang
Xinjiang
Xinjiang is an autonomous region of the People's Republic of China. It is the largest Chinese administrative division and spans over 1.6 million km2...

 province and elsewhere in Northern China; planting in Tongliao
Tongliao
Tongliao is a prefecture-level city in eastern Inner Mongolia, People's Republic of China. It borders Jilin province to the east, Liaoning to the south, Chifeng to the southwest, the Xilin Gol League to the west, and the Hinggan League to the north. The city was the administrative centre of the...

 City is known to have been particularly successful. White Elm is also known to have been introduced to Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

.

In recent years, the tree has enjoyed a small renaissance in England. A popular larval host plant of the White-letter Hairstreak
White-letter Hairstreak
The White-letter Hairstreak is a butterfly in the family Lycaenidae.-Appearance, behaviour and distribution:A dark little butterfly that spends the majority of its life in the tree tops, feeding on honeydew, making it best observed through binoculars. The uppersides are a dark brown with a small...

 Satyrium w-album butterfly across Europe, the elm is now being planted by Butterfly Conservation and other groups to restore local populations decimated by the effects of Dutch elm disease on native or archaeophytic elms. The Cheshire Wildlife Trust
Cheshire Wildlife Trust
The Cheshire Wildlife Trust is a wildlife trust covering the county of Cheshire and parts of the counties of Greater Manchester and Merseyside, England...

, for example, is planting 1,000 White Elms on its reserves in the former Vale Royal
Vale Royal
Vale Royal was, from 1974 to 2009, a local government district with borough status in Cheshire, England. It contained the towns of Northwich, Winsford and Frodsham.-Creation:...

 district of the county.

Uses

The timber of the White Elm is of poor quality and thus of little use to man, not even as firewood. Its density is significantly lower than that of other European elms.

Propagation

Usually easy to grow from seed sown to a depth of 6 mm in ordinary compost and kept well-watered. However, as seed viability can vary greatly from year to year, softwood cuttings taken in June may be a more reliable method. The cuttings strike very quickly, well within a fortnight, rapidly producing a dense matrix of roots.

Notable trees

The three largest known trees in Europe are at Gülitz
Gülitz-Reetz
Gülitz-Reetz is a municipality in the Prignitz district, in Brandenburg, Germany.-References:...

 in Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

 (3.1 m d.b.h.), at Komorów
Komorów, Swidnica County
Komorów is a village in the administrative district of Gmina Świdnica, within Świdnica County, Lower Silesian Voivodeship, in south-western Poland. Prior to 1945 it was in Germany....

 in Poland
Poland
Poland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...

 (2.96 m d.b.h.), and at Bergemolo in Piedmont
Piedmont
Piedmont is one of the 20 regions of Italy. It has an area of 25,402 square kilometres and a population of about 4.4 million. The capital of Piedmont is Turin. The main local language is Piedmontese. Occitan is also spoken by a minority in the Occitan Valleys situated in the Provinces of...

, Italy. The "Olmo di Bergemolo", over 200 years old, has a girth of 6.2 m (2008) and a height of 26 m. http://www.patriarchinatura.it/patriarchi_regione/piemonte.php A veteran between 300 and 500 years old, with a girth of 4.5 metres, survives in Rahnsdorf near Berlin. The UK Champion is at Ferry Farm, Harewood, Cornwall
Cornwall
Cornwall is a unitary authority and ceremonial county of England, within the United Kingdom. It is bordered to the north and west by the Celtic Sea, to the south by the English Channel, and to the east by the county of Devon, over the River Tamar. Cornwall has a population of , and covers an area of...

 (27 m high, 1.8 m d.b.h.). In the USA, a reputedly magnificent tree grows in Portland
Portland, Oregon
Portland is a city located in the Pacific Northwest, near the confluence of the Willamette and Columbia rivers in the U.S. state of Oregon. As of the 2010 Census, it had a population of 583,776, making it the 29th most populous city in the United States...

, Oregon
Oregon
Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

, but its dimensions and age are not known.

Subspecies and varieties

Several putative varieties have been identified. A variety celtidea from what is now the Ukraine
Ukraine
Ukraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...

 was reported by Rogowicz in the middle of the 19th century, but no examples are known to survive. Another variety parvifolia
Ulmus laevis var. parvifolia
Ulmus laevis var. parvifolia is a mutant variety of European White Elm found exclusively on the island of Ada Ciganlija, in the Sava at Belgrade, Serbia. -Description:...

 has been reported from Serbia
Serbia
Serbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...

.

Forms

In Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 several ornamental forms are recognized: f. aureovariegata, f. argentovariegata, f. rubra, and f. tiliifolia.

Cultivars

Compared with the other European species of elm, U. laevis has received scant horticultural attention, there being only five recorded cultivars, none of which are known to remain in cultivation, with the possible exception of 'Colorans' and 'Pendula':
Aureovariegata, Colorans, Ornata, Pendula, Punctata, Urticifolia
Ulmus laevis 'Urticifolia'
The European White Elm U. laevis cultivar Urticifolia was raised by Jacques as a chance seedling in 1830, and propagated by grafting. Listed in Revue Hort. 2: 128, 1832 as U. urticifolia. It was later mentioned by de Vries in Plant Breeding, 1906 .-Description:The tree was described as having...

.

Hybrid cultivars

U. laevis does not hybridize naturally, in common with the American Elm U. americana to which it is closely related. However, in experiments at the Arnold Arboretum, it was successfully crossed with U. thomasii and U. pumila; no such crosses have ever been released to commerce.

Accessions

North America
  • Arnold Arboretum. Acc. nos. 17910, 637-79, 6951, 753-80.
  • Brenton Arboretum
    Brenton Arboretum
    The Brenton Arboretum 121 acres is a new arboretum and botanical garden in early stages of development, located at 2629 Palo Circle, Dallas Center, Iowa, USA. The Arboretum aims to display all Iowa native trees and shrubs suitable to the site, as well as many other tree species which can grow in...

    , Dallas Center, Iowa
    Iowa
    Iowa is a state located in the Midwestern United States, an area often referred to as the "American Heartland". It derives its name from the Ioway people, one of the many American Indian tribes that occupied the state at the time of European exploration. Iowa was a part of the French colony of New...

    . No details available.
  • Brooklyn Botanic Garden
    Brooklyn Botanic Garden
    Brooklyn Botanic Garden is a botanical garden in the borough of Brooklyn in New York City. Located near the Prospect Heights, Crown Heights, and Park Slope neighborhoods, the garden includes a number of specialty "gardens within the Garden," plant collections, and the Steinhardt Conservatory,...

    , New York
    New York
    New York is a state in the Northeastern region of the United States. It is the nation's third most populous state. New York is bordered by New Jersey and Pennsylvania to the south, and by Connecticut, Massachusetts and Vermont to the east...

    . Acc. no. X02589.
  • Dominion Arboretum
    Dominion Arboretum
    The Dominion Arboretum is located at the Central Experimental Farm of Agriculture and Agri-Food Canada in Ottawa, Ontario, Canada.Originally begun in 1889 the Arboretum covers about 26 ha of rolling land between Prince of Wales Drive, Dow's Lake and the Rideau Canal. Carleton University is located...

    , Canada
    Canada
    Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

    . No details available
  • Longwood Gardens
    Longwood Gardens
    Longwood Gardens consists of over 1,077 acres of gardens, woodlands, and meadows in Kennett Square, Pennsylvania, United States in the Brandywine Creek Valley...

    . Acc. nos. 1964-0568, 1964-1119.
  • Morton Arboretum
    Morton Arboretum
    The Morton Arboretum, in Lisle, Illinois, covers 1,700 acres and is made up of gardens of various plant types and collections of trees from specific taxonomical and geographical areas. It includes native woodlands and a restored Illinois prairie. The Arboretum has over 4,100 different species of...

    , Illinois
    Illinois
    Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

    . Acc. nos. 1302-27, 446-48, 492-64, 27-98.

Europe
  • Arboretum de La Petite Loiterie
    Arboretum de la Petite Loiterie
    The Arboretum de la Petite Loiterie is a young arboretum located in Le Sentier, Monthodon, Indre-et-Loire, Centre, France. It is open a few days per month; an admission fee is charged....

    , Monthodon
    Monthodon
    Monthodon is a commune in the Indre-et-Loire department in central France....

    , France
    France
    The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

    . No details available
  • Arboretum Freiburg-Günterstal
    Arboretum Freiburg-Günterstal
    The Arboretum Freiburg-Günterstal is an arboretum maintained by the University of Freiburg. It is located in Günterstal in the Städtischen Forstamt Freiburg at Günterstalstraße 71, Freiburg, Baden-Württemberg, Germany, and open daily without charge.The arboretum dates back to 1896, or earlier,...

    , Germany
    Germany
    Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

    , no details available
  • Brighton & Hove
    Brighton & Hove
    Brighton and Hove is a unitary authority area and city on the south coast of England. It is England's most populous seaside resort.In 1997 Brighton and Hove were joined to form the unitary authority of Brighton and Hove, which was granted city status by Queen Elizabeth II as part of the millennium...

     City Council, UK, NCCPG Elm Collection. Ten trees at Hove Recreation Ground, Hove.
  • Copenhagen University, Botanic Garden. No details available.
  • ELTE Botanic Garden Budapest. Acc. nos. 1998-0718, 1998-0719.
  • Grange Farm Arboretum
    Grange Farm Arboretum
    The Grange Farm Arboretum is a small private arboretum comprising 3 hectares accommodating over 800 trees, mostly native and ornamental species or cultivars, notably oaks, ashes, walnuts and elms, growing on a calcareous loam....

    , Sutton St. James, Spalding
    Spalding, Lincolnshire
    Spalding is a market town with a population of 30,000 on the River Welland in the South Holland district of Lincolnshire, England. Little London is a hamlet directly south of Spalding on the B1172 road....

    , Lincolnshire
    Lincolnshire
    Lincolnshire is a county in the east of England. It borders Norfolk to the south east, Cambridgeshire to the south, Rutland to the south west, Leicestershire and Nottinghamshire to the west, South Yorkshire to the north west, and the East Riding of Yorkshire to the north. It also borders...

    , UK. Acc. no. 502.
  • Great Fontley Farm, Fareham
    Fareham
    The market town of Fareham lies in the south east of Hampshire, England, between the cities of Southampton and Portsmouth, roughly in the centre of the South Hampshire conurbation.It gives its name to the borough comprising the town and the surrounding area...

    , UK, Butterfly Conservation
    Butterfly Conservation
    Butterfly Conservation is an insect conservation organisation in the United Kingdom. It is one of the largest insect conservation organisations in the world.-History of the Organisation:...

    Elm Trials plantation, Home Field K2 and Platts N10, planted 2003, grown from cuttings of specimen at RBG Wakehurst Place. Acc. no. 1973-21048.
  • Hortus Botanicus Nationalis, Salaspils
    Salaspils
    Salaspils is a town in Latvia, the administrative centre of Salaspils municipality. The town is situated on the northern bank of the Daugava River 18 kilometers to the south-east of the city of Riga.-History:...

    , Latvia
    Latvia
    Latvia , officially the Republic of Latvia , is a country in the Baltic region of Northern Europe. It is bordered to the north by Estonia , to the south by Lithuania , to the east by the Russian Federation , to the southeast by Belarus and shares maritime borders to the west with Sweden...

    . Acc. nos. 18136, 18140.
  • Linnaean Gardens of Uppsala
    Linnaean Garden
    The Linnaean Garden or Linnaeus' Garden is the oldest of the botanical gardens belonging to Uppsala University in Sweden. It has been restored and is kept as an 18th century botanical garden, according to the specifications of Carolus Linnaeus.The garden was originally planned and planted by Olaus...

    , Sweden
    Sweden
    Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

    . Acc. no. 1930-1014.
  • Royal Botanic Garden Edinburgh
    Royal Botanic Garden Edinburgh
    The Royal Botanic Garden Edinburgh is a scientific centre for the study of plants, their diversity and conservation, as well as a popular tourist attraction. Originally founded in 1670 as a physic garden to grow medicinal plants, today it occupies four sites across Scotland — Edinburgh,...

    , UK. Acc. no. 20070643, from seed wild collected in Val d'Allier, France
    France
    The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

    .
  • Royal Botanic Gardens Kew, UK. Acc. nos. 1969-17302, 1973-11712.
  • Royal Botanic Gardens
    Royal Botanic Gardens
    Royal Botanic Gardens or Royal Botanical Gardens might refer to:* Royal Botanical Gardens in Canada* Royal Botanical Garden of Madrid in Spain* Royal Botanic Gardens, Trinidad in Port of Spain, Trinidad and Tobago...

    , Wakehurst Place, UK. Acc. no. 1973-21048.
  • Royal Horticultural Society
    Royal Horticultural Society
    The Royal Horticultural Society was founded in 1804 in London, England as the Horticultural Society of London, and gained its present name in a Royal Charter granted in 1861 by Prince Albert...

     Gardens, Wisley
    Wisley
    Wisley is a small village in Surrey, England. It lies between Cobham and Woking. It is the home of the Royal Horticultural Society's Wisley Garden. The River Wey runs through the village....

    , UK. No details available.
  • Sir Harold Hillier Gardens
    Sir Harold Hillier Gardens
    The Sir Harold Hillier Gardens is an arboretum comprising 72 hectares accommodating over 42,000 trees and shrubs in about 12,000 taxa, notably a collection of oaks, camellia, magnolia and rhododendron....

    , UK. Acc. no. 1981-2446.
  • Strona Arboretum Ulmus lamellosa
    Ulmus lamellosa
    Ulmus lamellosa, commonly called the Hebei Elm, is a small deciduous tree native to four Chinese provinces, Hebei, Henan, Nei Mongol, and Shanxi, to the west and south of Beijing .-Description:...

    , University of Life Sciences, Warsaw
    Warsaw
    Warsaw is the capital and largest city of Poland. It is located on the Vistula River, roughly from the Baltic Sea and from the Carpathian Mountains. Its population in 2010 was estimated at 1,716,855 residents with a greater metropolitan area of 2,631,902 residents, making Warsaw the 10th most...

    , Poland
    Poland
    Poland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...

    .
  • Tallinn
    Tallinn
    Tallinn is the capital and largest city of Estonia. It occupies an area of with a population of 414,940. It is situated on the northern coast of the country, on the banks of the Gulf of Finland, south of Helsinki, east of Stockholm and west of Saint Petersburg. Tallinn's Old Town is in the list...

     Botanic Garden, Estonia
    Estonia
    Estonia , officially the Republic of Estonia , is a state in the Baltic region of Northern Europe. It is bordered to the north by the Gulf of Finland, to the west by the Baltic Sea, to the south by Latvia , and to the east by Lake Peipsi and the Russian Federation . Across the Baltic Sea lies...

    . No accession details available.
  • Thenford House arboretum, Oxfordshire, UK. No details available.
  • 'The Leys', University Parks, Oxford, UK. Acc. no. 02678.
  • Westonbirt Arboretum
    Westonbirt Arboretum
    Westonbirt, The National Arboretum is managed by the Forestry Commission. Westonbirt Arboretum is located near the historic market town of Tetbury in Gloucestershire, England, and is perhaps the most important and widely known arboretum in the United Kingdom.Planted in the heyday of Victorian plant...

    , Tetbury
    Tetbury
    Tetbury is a town and civil parish within the Cotswold district of Gloucestershire, England. It lies on the site of an ancient hill fort, on which an Anglo-Saxon monastery was founded, probably by Ine of Wessex, in 681. The population of the parish was 5,250 in the 2001 census.In the Middle Ages,...

    , Glos.
    Gloucestershire
    Gloucestershire is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn, and the entire Forest of Dean....

    , UK, planted 1997. Acc. no. 1995/322

Nurseries

North America
None known
Europe
  • Arboretum Waasland, Nieuwkerken-Waas, Belgium
    Belgium
    Belgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...

    .
  • Boomkwekerij Oirschot, Oirschot
    Oirschot
    Oirschot is a municipality and a town in the southern Netherlands. It is situated 12 km from the city of Eindhoven and 20 km from the city of Tilburg in the province Noord-Brabant...

    , Netherlands
    Netherlands
    The Netherlands is a constituent country of the Kingdom of the Netherlands, located mainly in North-West Europe and with several islands in the Caribbean. Mainland Netherlands borders the North Sea to the north and west, Belgium to the south, and Germany to the east, and shares maritime borders...

  • Crown Nursery, Ufford, Suffolk
    Suffolk
    Suffolk is a non-metropolitan county of historic origin in East Anglia, England. It has borders with Norfolk to the north, Cambridgeshire to the west and Essex to the south. The North Sea lies to the east...

    , UK
  • Dulford Nurseries, Cullompton
    Cullompton
    Cullompton is a civil parish and town in Devon, England, locally known as Cully. It is miles north-north-east of Exeter and lies on the River Culm. In 2010 it had a population of 8,639 and is growing rapidly....

    , Devon
    Devon
    Devon is a large county in southwestern England. The county is sometimes referred to as Devonshire, although the term is rarely used inside the county itself as the county has never been officially "shired", it often indicates a traditional or historical context.The county shares borders with...

    , UK
  • Landcare Tree Nursery, Old Sodbury
    Old Sodbury
    Old Sodbury is a small village in the valley of the River Frome just below and to the west of the Cotswold escarpment and to the east of Chipping Sodbury and Yate in South Gloucestershire, England. The village extends from Chipping Sodbury in the West to the Cotswold Edge in the East and is on the...

    , South Gloucestershire
    South Gloucestershire
    South Gloucestershire is a unitary district in the ceremonial county of Gloucestershire, in South West England.-History:The district was created in 1996, when the county of Avon was abolished, by the merger of former area of the districts of Kingswood and Northavon...

    , UK
  • Lorenz von Ehren, Hamburg
    Hamburg
    -History:The first historic name for the city was, according to Claudius Ptolemy's reports, Treva.But the city takes its modern name, Hamburg, from the first permanent building on the site, a castle whose construction was ordered by the Emperor Charlemagne in AD 808...

    , Germany
    Germany
    Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

  • UmbraFlor, Spello
    Spello
    Spello is an ancient town and comune of Italy, in the province of Perugia in east central Umbria, on the lower southern flank of Mt. Subasio. It is 6 km NNW of Foligno and 10 km SSE of Assisi.The old walled town lies on a regularly NW-SE sloping ridge that eventually meets the plain...

    , Italy
    Italy
    Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...


External links

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