Candoia
Encyclopedia
Common names: bevel-nosed boas, keel-scaled boas.

Candoia is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of non-venomous boa
Boinae
The Boinae are a subfamily of boas found in Central and South America, Africa and Southeast Asia. Five genera comprising 28 species are currently recognized.-Geographic range:...

s found in mostly in New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

 and Melanesia
Melanesia
Melanesia is a subregion of Oceania extending from the western end of the Pacific Ocean to the Arafura Sea, and eastward to Fiji. The region comprises most of the islands immediately north and northeast of Australia...

. Four species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 are currently recognized.

Description

Members in this genus typically have a rounded and heavy body, with a flattened triangular shaped head, and an upturned nose. Colors and patterns vary greatly, but most are various shades of brown to black. They can vary in adult size from 60 cm to 1.8 m. Males are typically smaller than females and can be distinguished by their prominent cloacal spurs.

Geographic range

Found from Samoa
Samoa
Samoa , officially the Independent State of Samoa, formerly known as Western Samoa is a country encompassing the western part of the Samoan Islands in the South Pacific Ocean. It became independent from New Zealand in 1962. The two main islands of Samoa are Upolu and one of the biggest islands in...

 and Tokelau
Tokelau
Tokelau is a territory of New Zealand in the South Pacific Ocean that consists of three tropical coral atolls with a combined land area of 10 km2 and a population of approximately 1,400...

 west through Melanesia
Melanesia
Melanesia is a subregion of Oceania extending from the western end of the Pacific Ocean to the Arafura Sea, and eastward to Fiji. The region comprises most of the islands immediately north and northeast of Australia...

 to New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

 and the Maluku Islands
Maluku Islands
The Maluku Islands are an archipelago that is part of Indonesia, and part of the larger Maritime Southeast Asia region. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone...

.

Reproduction

Breeding occurs early in the year, typically after rains. Several males will approach and pursue a single female, though, there is no combat between the males. Females seem to only ovulate once every two or three years and give birth to litters averaging 10 or so neonates. The Solomon Island ground boa, C. carinata paulsoni is an exception as it is known to have particularly large litters, occasionally producing 30 or 40 small neonates.

Captivity

These snakes are frequently imported for the exotic pet trade, but are now being bred in captivity with some regularity by private individuals. Their small size and ease of care make them interesting captives, but wild caught specimens are known to not acclimate well. The stress of captivity manifests itself in the form of lack of interest in food. Their natural diet often presents a problem for hobbyists mainly familiar with using rodents as food.

Species

Species Taxon author Subsp.* Common name Geographic range
C. aspera
Candoia aspera
Candoia aspera, known commonly as the New Guinean boa, is a species of snake in the Boidae family. As its common name suggests, it is found in New Guinea....

(Günther
Albert C. L. G. Günther
Albert Karl Ludwig Gotthilf Günther FRS, also Albert Charles Lewis Gotthilf Günther , was a German-born British zoologist, ichthyologist, and herpetologist....

, 1877)
0 New Guinean boa New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

 (Irian Jaya
Western New Guinea
West Papua informally refers to the Indonesian western half of the island of New Guinea and other smaller islands to its west. The region is officially administered as two provinces: Papua and West Papua. The eastern half of New Guinea is Papua New Guinea.The population of approximately 3 million...

 and Papua New Guinea
Papua New Guinea
Papua New Guinea , officially the Independent State of Papua New Guinea, is a country in Oceania, occupying the eastern half of the island of New Guinea and numerous offshore islands...

 below 1300 m), including the nearby islands of Waigeu, Batanta
Batanta
Batanta is one of the four major islands in the Raja Ampat Islands in Papua province, Indonesia. Its area is 453 km² and its highest point is 1184 m.- Fauna :The following mammals are found on the island:...

, Misool
Misool
Misool is one of the four major islands in the Raja Ampat Islands in West Papua , Indonesia. Its area is 2034 km². The highest point is 535 m and the main town is Waigama, on the island's northern coast, and Lilinta....

 and Salawati
Salawati
Salawati is one of the four major islands in the Raja Ampat Islands in West Papua , Indonesia. Its area is 1623 km². The other main islands in the archipelago are Misool, Batanta and Waigeo....

, the islands of Biak
Biak
Biak features a tropical rainforest climate with nearly identical temperatures throughout the course of the year. The average annual temperature in the city is 27 degrees celsius, which is also generally the average temperature of each day in Biak...

 and Japen
Yapen
Yapen is an island of western New Guinea, Indonesia. The Yapen Strait separates Yapen and the Biak Islands to the north. It is in Cenderawasih Bay. To the west is Mios Num Strait between it and Mios Num Island, and the east Kurudu Island. The southeast are the Amboi Islands and the southwest...

 in Geelvink Bay
Cenderawasih Bay
Cenderawasih Bay , also Teluk Sarera , formerly Geelvink Bay is a large bay in northern Province of Papua and West Papua, New Guinea, Indonesia, at . The Dutch name comes after a Dutch ship and family called Geelvinck...

, Seleo, Valise, Karkar
Karkar Island
Karkar Island is an oval-shaped volcanic island located in the Bismarck Sea, about 30 kilometres off the north coast of mainland Papua New Guinea in Madang Province. The island is about 25 km in length and 19 km in width. In the centre is an active volcano with two nested calderas...

, Umboi
Umboi Island
Umboi is a volcanic island between the mainland of Papua New Guinea and the island of New Britain. It is separated from New Britain by Dampier Strait and has an elevation of 1,548 m. Umboi is a Holocene complex volcano with no historic eruptions...

, Manus
Manus Island
Manus Island is part of Manus Province in northern Papua New Guinea and is the largest island of the Admiralty Islands. It is the fifth largest island in Papua New Guinea with an area of 2,100 km², measuring around 100 km × 30 km. According to the 2000 census, Manus Island had a...

, Los Negros
Los Negros Island
Los Negros Island is the third largest of the Admiralty Islands. It is significant because it contains the main airport of Manus Province on its eastern coastline, at Momote. It is connected to Lorengau, the capital of the province, on Manus Island via a highway and bridge...

, Lou, Baluan, the Rambatyo Islands in the Manus Group (Admiralty Islands
Admiralty Islands
The Admiralty Islands are a group of eighteen islands in the Bismarck Archipelago, to the north of New Guinea in the south Pacific Ocean. These are also sometimes called the Manus Islands, after the largest island. These rainforest-covered islands form part of Manus Province, the smallest and...

), New Britain
New Britain
New Britain, or Niu Briten, is the largest island in the Bismarck Archipelago of Papua New Guinea. It is separated from the island of New Guinea by the Dampier and Vitiaz Straits and from New Ireland by St. George's Channel...

, Duke of York
Duke of York Island, Papua New Guinea
Duke of York Island is the largest island of Duke of York Islands, Papua New Guinea, at . The island is named after Prince Edward, the brother of King George III of Great Britain....

, New Ireland
New Ireland (island)
New Ireland is a large island in Papua New Guinea, approximately 7,404 km² in area. It is the largest island of the New Ireland Province, lying northeast of the island of New Britain. Both islands are part of the Bismarck Archipelago, named after Otto von Bismarck, and they are separated by...

 and New Hanover
New Hanover Island
New Hanover Island, , also called Lavongai, is a large volcanic island in New Ireland Province, part of the Bismarck Archipelago of the New Guinea Islands region of Papua New Guinea, at...

 in the Bismarck Archipelago
Bismarck Archipelago
The Bismarck Archipelago is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea.-History:...

.
C. bibroni
Candoia bibroni
Candoia bibroni is a non-venomous boa species found in Melanesia and Polynesia. Two subspecies are currently recognized, including the nominate subspecies described here.-Description:...

(Duméril & Bibron
Gabriel Bibron
Gabriel Bibron was a French zoologist. He was born in Paris. Son of an employee of the Museum national d'histoire naturelle, he had a good foundation in natural history and was hired to collect vertebrates in Italy and Sicily. He classified a number of reptile species with André Marie Constant...

, 1844)
1 Fiji boa Melanesia
Melanesia
Melanesia is a subregion of Oceania extending from the western end of the Pacific Ocean to the Arafura Sea, and eastward to Fiji. The region comprises most of the islands immediately north and northeast of Australia...

 and Polynesia
Polynesia
Polynesia is a subregion of Oceania, made up of over 1,000 islands scattered over the central and southern Pacific Ocean. The indigenous people who inhabit the islands of Polynesia are termed Polynesians and they share many similar traits including language, culture and beliefs...

, including the eastern Solomon Islands
Solomon Islands
Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

 (Olu Malau, Reef, Rennell
Rennell Island
Rennell Island, locally known as Mungava, is the main island of two inhabited islands that make up the Rennell and Bellona Province in the Solomon Islands. Rennell Island has a land area of that is about long and wide. It is the second largest raised coral atoll in the world with the largest lake...

, San Critobal, Santa Ana, Santa Cruz
Santa Cruz Islands
The Santa Cruz Islands are a group of islands in the Pacific Ocean, part of Temotu Province of the Solomon Islands. They lie approximately 250 miles to the southeast of the Solomon Islands Chain...

, Bio, Bellona
Bellona Island
Bellona Island is an island of the Rennell and Bellona Province, Solomon Islands. Its length is about 10 km and its average width 2.5 km. Its area is about 17 km². It is almost totally surrounded by 30–70 m high cliffs, consisting primarily of raised coral...

, Three Sisters, Ugi and Vanikoro), the Banks Islands
Banks Islands
The Banks Islands are a group of islands in northern Vanuatu.Together with the Torres Islands to the northwest, they make up the northernmost province of Torba. The group lies about north of Maewo, and includes Gaua and Vanua Lava, two of the 13 largest islands in Vanuatu...

 (Vanua Lava Island), the New Hebrides
New Hebrides
New Hebrides was the colonial name for an island group in the South Pacific that now forms the nation of Vanuatu. The New Hebrides were colonized by both the British and French in the 18th century shortly after Captain James Cook visited the islands...

, all three of the Loyalty Islands
Loyalty Islands
The Loyalty Islands are an archipelago in the Pacific. They are part of the French territory of New Caledonia, whose mainland is away. They form the Loyalty Islands Province , one of the three provinces of New Caledonia...

, the Fiji Islands
Fiji
Fiji , officially the Republic of Fiji , is an island nation in Melanesia in the South Pacific Ocean about northeast of New Zealand's North Island...

 (Rotuma
Rotuma
Rotuma is a Fijian dependency, consisting of Rotuma Island and nearby islets. The island group is home to a small but unique indigenous ethnic group which constitutes a recognizable minority within the population of Fiji, known as "Rotumans"...

, the Yasawa Group
Yasawa Islands
The Yasawa Group is an archipelago of about 20 volcanic islands in the Western Division of Fiji, with an approximate total area of 135 square kilometers.- Geography :The Yasawa volcanic group consists of six main islands and numerous smaller islets...

 and the Lau Group
Lau Islands
The Lau Islands of Fiji are situated in the southern Pacific Ocean, just east of the Koro Sea. Of this chain of about one hundred islands and islets, about thirty are inhabited...

), Western Samoa
Samoa
Samoa , officially the Independent State of Samoa, formerly known as Western Samoa is a country encompassing the western part of the Samoan Islands in the South Pacific Ocean. It became independent from New Zealand in 1962. The two main islands of Samoa are Upolu and one of the biggest islands in...

 (Savii
Savai'i
Savaii is the largest and highest island in Samoa and the Samoa Islands chain. It is also the biggest landmass in Polynesia outside Hawaii and New Zealand. The island of Savai'i is also referred to by Samoans as Salafai, a classical Samoan term used in oratory and prose...

 and Upolu
Upolu
Upolu is an island in Samoa, formed by a massive basaltic shield volcano which rises from the seafloor of the western Pacific Ocean. The island is long, in area, and is the second largest in geographic area as well as the most populated of the Samoan Islands. Upolu is situated to the east of...

 islands) and American Samoa
American Samoa
American Samoa is an unincorporated territory of the United States located in the South Pacific Ocean, southeast of the sovereign state of Samoa...

 (Ta'u Island
Tau, American Samoa
Ta‘ū is the largest island in the Manu‘a Group and the easternmost volcanic island of the Samoan Islands. Ta‘ū is part of American Samoa. In the early 19th century, the island was sometimes called Opoun....

).
C. carinata
Candoia carinata
Candoia carinata, known commonly as the Pacific Ground Boa or the Pacific Keel-scaled Boa, is a species of snake in the Boidae family. It is found in Indonesia, New Guinea, and the Bismarck Archipelago....

T
(Schneider
Johann Gottlob Schneider
Johann Gottlob Theaenus Schneider was a German classicist and naturalist.-Biography:Schneider was born at Collm in Saxony...

, 1801)
1 Pacific boa Indonesia
Indonesia
Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

, including the northern peninsula of Sulawesi
Sulawesi
Sulawesi is one of the four larger Sunda Islands of Indonesia and is situated between Borneo and the Maluku Islands. In Indonesia, only Sumatra, Borneo, and Papua are larger in territory, and only Java and Sumatra have larger Indonesian populations.- Etymology :The Portuguese were the first to...

, Sangihe and Telaud Islands, the northern Maluku Islands
Maluku Islands
The Maluku Islands are an archipelago that is part of Indonesia, and part of the larger Maritime Southeast Asia region. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone...

 (Ceram
CERAM
CERAM is a materials science and testing organisation based in Stoke-on-Trent specialising in the field of ceramics. The CERAM Group owns testing facilities around the world.-History:The British Refractories Research Association was formed in 1920...

, Ambon
Ambon Island
Ambon Island is part of the Maluku Islands of Indonesia. The island has an area of , and is mountainous, well watered, and fertile. Ambon Island consists of 2 territories: The main city and seaport is Ambon , which is also the capital of Maluku province and Maluku Tengah Ambon Island is part of the...

, Haruku, Saparua
Saparua
Saparua is an island east of Ambon Island in the Indonesian province of Maluku. The main port is in the south at Kota Saparua.Saparua was the location of Indonesian national hero, Pattimura's rebellion against Dutch forces.Paperu is a village on the island....

, Banda
Banda Islands
The Banda Islands are a volcanic group of ten small volcanic islands in the Banda Sea, about south of Seram Island and about east of Java, and are part of the Indonesian province of Maluku. The main town and administrative centre is Bandanaira, located on the island of the same name. They rise...

, Goram and Tanimbar
Tanimbar Islands
The Tanimbar Islands, also called Timor Laut, are a group of about 65 islands in the Maluku province of Indonesia, including Fordata, Larat, Maru, Molu, Nuswotar, Selaru, Selu, Seira, Wotap, Wuliaru and Yamdena.-Geography:...

), Misool
Misool
Misool is one of the four major islands in the Raja Ampat Islands in West Papua , Indonesia. Its area is 2034 km². The highest point is 535 m and the main town is Waigama, on the island's northern coast, and Lilinta....

, Batanta
Batanta
Batanta is one of the four major islands in the Raja Ampat Islands in Papua province, Indonesia. Its area is 453 km² and its highest point is 1184 m.- Fauna :The following mammals are found on the island:...

, Salawatti, Anaguar (Saipan
Saipan
Saipan is the largest island of the United States Commonwealth of the Northern Mariana Islands , a chain of 15 tropical islands belonging to the Marianas archipelago in the western Pacific Ocean with a total area of . The 2000 census population was 62,392...

) and the Palau Group
Palau
Palau , officially the Republic of Palau , is an island nation in the Pacific Ocean, east of the Philippines and south of Tokyo. In 1978, after three decades as being part of the United Nations trusteeship, Palau chose independence instead of becoming part of the Federated States of Micronesia, a...

 (at least on Koror Island
Koror
Koror is the state comprising the main commercial centre of the Republic of Palau. It consists of several islands, the most prominent being Koror Island ....

). New Guinea: the islands of Geelvink Bay and those along the north coast, including Liki, Djamna, Karkar and the Umboi islands. Manus and nearby islands. The Bismarck Archipelago, including the islands of Mussau
Mussau Island
Mussau Island is the largest island of St. Matthias Islands, Papua New Guinea, at . It is one of the northernmost islands of Papua New Guinea....

 and Tench. The islands of Milne Bay Province
Milne Bay Province
Milne Bay is a province of Papua New Guinea. Its capital is Alotau. The province covers 14,000 km² of land and 252,990 km² of sea, within the province there are more than 600 islands, about 160 of which are inhabited...

, at least on Samarai
Samarai
Samarai is an island and former administrative capital in Milne Bay Province, Papua New Guinea. Located off the south-eastern tip of New Guinea in the China Strait Samarai has an area of just ....

, Kiriwina
Kiriwina
Kiriwina is the largest of the Trobriand Islands, with an area of 290.5 km². It is part of the Milne Bay Province of Papua New Guinea. Most of the 12,000 people who live in the Trobriands live on Kiriwina. The Kilivila language, also known as Kiriwina, is spoken on the island...

, Kitava, Goodenough
Goodenough Island
Goodenough Island in the Solomon Sea is the westernmost of the three large islands of the D'Entrecasteaux Islands in Milne Bay Province of Papua New Guinea. It lies to the east of mainland New Guinea and south west of the Trobriand Islands.It should not be confused with Goodenough's Island...

, Fergusson
Fergusson Island
Fergusson Island is the largest island of the D'Entrecasteaux Islands, in Papua New Guinea. It has an area of just over 500 square miles , and mostly consists of mountainous regions, covered by rain forests...

, Dobu
Dobu Island
Dobu Island is an island, part of D'Entrecasteaux Islands in Papua New Guinea. It is located south of Fergusson Island and north of Normanby Island.The people of Dobu were the subject of a seminal anthropological study by Reo Fortune...

, Normanby, Slade, Misima
Misima Island
Misima is a volcanic island in the northwest of Louisiade Archipelago within Milne Bay Province of Papua New Guinea. The island measures 40 km by 10 km and has an area of 202,5 km². It is located some 20 km north of the northwest extreme of the barrier reef of Vanatinai at Isu Raua Raua Island,...

, Woodlark
Woodlark Island
Woodlark Island, known to its inhabitants simply as Woodlark or Muyua, is an island in Milne Bay Province, Papua New Guinea. It is called Murua by the inhabitants of some other islands in the province...

, Sudest and Rossel
Rossel Island (Yela)
Rossel Island is the easternmost island of the Louisiade Archipelago, which itself is part of the Milne Bay Province of Papua New Guinea. The volcanic island measures 34 km east-west, and is up to 11 km wide. With an area of 262.5 km², it is the second largest island of the archipelago, after...

. The islands of Buka
Buka, Papua New Guinea
Buka is situated on Buka Island, Bougainville Province, Papua New Guinea. It has been the capital of the province during the Bougainville civil war...

, Bougainville
Bougainville Island
Bougainville Island is the main island of the Autonomous Region of Bougainville of Papua New Guinea. This region is also known as Bougainville Province or the North Solomons. The population of the province is 175,160 , which includes the adjacent island of Buka and assorted outlying islands...

 and those nearby. The Solomon Islands, including Santa Cruz, Rennell and Bellona.
C. superciliosa
Candoia superciliosa
Candoia superciliosa, known commonly as the Palau bevel-nosed boa, is a species of snake in the Boidae family. As its common name suggests, it is found in Palau....

(Günther, 1863) 0 Palau bevel-nosed boa Belau
Palau
Palau , officially the Republic of Palau , is an island nation in the Pacific Ocean, east of the Philippines and south of Tokyo. In 1978, after three decades as being part of the United Nations trusteeship, Palau chose independence instead of becoming part of the Federated States of Micronesia, a...

.

*) Not including the nominate subspecies.

T
) Type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

.

External links

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