Arbutus
Encyclopedia
"Madroño" redirects here. In tropical America, this can refer to the mangosteen Garcinia madruno, an unrelated eudicot.


Arbutus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of at least 14 species of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

s in the family Ericaceae
Ericaceae
The Ericaceae, commonly known as the heath or heather family, is a group of mostly calcifuge flowering plants. The family is large, with roughly 4000 species spread across 126 genera, making it the 14th most speciose family of flowering plants...

, native to warm temperate regions of the Mediterranean, western Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, and North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

.

Description

Arbutus are small trees or shrubs with red flaking bark and edible red berries. Fruit development is delayed for about five months after pollination, so that flowers appear while the previous year's fruit are ripening.

Common names

North American members of the genus are called Madrones
Pacific Madrone
Arbutus menziesii, commonly known as the Pacific Madrone, is a species of Arbutus found on the west coast of North America, from British Columbia to California...

, from the Spanish
Spanish language
Spanish , also known as Castilian , is a Romance language in the Ibero-Romance group that evolved from several languages and dialects in central-northern Iberia around the 9th century and gradually spread with the expansion of the Kingdom of Castile into central and southern Iberia during the...

 name madroño (strawberry tree) although this terminology is not used in Canada. The Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

an species are also called Strawberry Trees from the superficial resemblance of the fruit to a strawberry
Strawberry
Fragaria is a genus of flowering plants in the rose family, Rosaceae, commonly known as strawberries for their edible fruits. Although it is commonly thought that strawberries get their name from straw being used as a mulch in cultivating the plants, the etymology of the word is uncertain. There...

; some species are sometimes referred to simply as the "Arbutus". In the United States, the name "Madrone" is used south of the Siskiyou Mountains
Siskiyou Mountains
The Siskiyou Mountains are a coastal mountain range in the northern Klamath Mountains in northwestern California and southwestern Oregon in the United States. They extend in an arc for approximately from east of Crescent City, California northeast along the north side of the Klamath River into...

 of southern Oregon/northern California and the name "Madrona" is used north of the Siskiyou Mountains according to the "Sunset Western Garden Book". In British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

, the trees are simply known by the name "Arbutus." All refer to the same tree, Arbutus menziesii
Pacific Madrone
Arbutus menziesii, commonly known as the Pacific Madrone, is a species of Arbutus found on the west coast of North America, from British Columbia to California...

, native to the Pacific Northwest and Northern California regions. It is Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

's only native broadleaved evergreen tree. Some species in the genera Epigaea
Epigaea
Epigaea is a genus comprising three species of flowering plants in the Ericaceae. The species are small creeping shrubs that are typically anywhere from 10 to 20 cm tall at full growth, forming large patches. The leaves are evergreen, alternate and simple, ranging amongst the three species...

, Arctostaphylos
Arctostaphylos
Arctostaphylos is a genus of plants comprised by the manzanitas and bearberries. They are shrubs or small trees.There are about 60 species of Arctostaphylos, ranging from ground-hugging arctic, coastal, and mountain species to small trees up to 6 m tall. Most are evergreen , with small oval...

and Gaultheria
Gaultheria
Gaultheria is a genus of about 170-180 species of shrubs in the family Ericaceae. The name memorializes Jean François Gauthier of Quebec, a mis-spelt honour bestowed by the Scandinavian Pehr Kalm in 1748. These plants are native to Asia, North and South America, and Australasia...

were formerly classified in Arbutus. As a result of its past classification, Epigaea repens
Epigaea
Epigaea is a genus comprising three species of flowering plants in the Ericaceae. The species are small creeping shrubs that are typically anywhere from 10 to 20 cm tall at full growth, forming large patches. The leaves are evergreen, alternate and simple, ranging amongst the three species...

(Mayflower) has an alternative common name of "trailing arbutus".

Species

A recent study which analyzed ribosomal DNA
Ribosomal DNA
Ribosomal DNA codes for ribosomal RNA. The ribosome is an intracellular macromolecule that produces proteins or polypeptide chains. The ribosome itself consists of a composite of proteins and RNA. As shown in the figure, rDNA consists of a tandem repeat of a unit segment, an operon, composed of...

 from Arbutus and related genera suggests that Arbutus is paraphyletic and the Mediterranean Basin
Mediterranean Basin
In biogeography, the Mediterranean Basin refers to the lands around the Mediterranean Sea that have a Mediterranean climate, with mild, rainy winters and hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation...

 species of Arbutus are more closely related to Arctostaphylos
Arctostaphylos
Arctostaphylos is a genus of plants comprised by the manzanitas and bearberries. They are shrubs or small trees.There are about 60 species of Arctostaphylos, ranging from ground-hugging arctic, coastal, and mountain species to small trees up to 6 m tall. Most are evergreen , with small oval...

, Arctous, Comarostaphylis
Comarostaphylis
Comarostaphylis is a genus of shrubs in the heath family native to the Americas from California in the United States to Panama. These are hairy, glandular shrubs to small trees with shreddy bark, often quite similar to their close relatives, the manzanitas....

, Ornithostaphylos
Ornithostaphylos
Ornithostaphylos is a monotypic plant genus which contains the single species Ornithostaphylos oppositifolia, the Baja birdbush or Baja California birdbush. This is a shrub native to the chaparral plant community of the coastline of southern California and northern Baja California. This bush has...

and Xylococcus than to the western North American species of Arbutus, and that the split between the two groups of species occurred at the Paleogene
Paleogene
The Paleogene is a geologic period and system that began 65.5 ± 0.3 and ended 23.03 ± 0.05 million years ago and comprises the first part of the Cenozoic Era...

/Neogene
Neogene
The Neogene is a geologic period and system in the International Commission on Stratigraphy Geologic Timescale starting 23.03 ± 0.05 million years ago and ending 2.588 million years ago...

 boundary.

Old World

  • Arbutus andrachne
    Arbutus andrachne
    Arbutus andrachne, commonly called the Greek Strawberry Tree, is an evergreen shrub or small tree in the family Ericaceae, native to the Mediterranean region, the Middle East, and southwestern Asia.-Garden history:...

    (Greek Strawberry Tree). Southeastern Europe
    Europe
    Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

     and southwestern Asia
    Asia
    Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

    .
  • Arbutus canariensis
    Arbutus canariensis
    Arbutus canariensis is a species of shrub or tree in the Ericaceae family. It is endemic to the Canary Islands of Spain. It is threatened by habitat loss.-References:* Bañares, A. et al. 1998. . Downloaded on 20 August 2007....

    (Canary Madrone). Canary Islands
    Canary Islands
    The Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...

    .
  • Arbutus unedo (Strawberry Tree). Widespread in the Mediterranean region and also western France
    France
    The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

     and western Ireland
    Ireland
    Ireland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...

     as well as south to the Algarve in Portugal
    Portugal
    Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

    .
  • Note: the East Asian fruit Myrica rubra
    Myrica rubra
    Myrica rubra, also called yangmei , yamamomo , Chinese Bayberry, Japanese Bayberry, Red Bayberry,Yumberry, or Chinese strawberry tree is a subtropical tree grown for its sweet, crimson to dark purple-red, edible fruit.- Description :It is a small to medium-sized evergreen tree...

    is often mistranslated as Arbutus.
  • A. unedo and A. andrachne hybridise naturally where their ranges overlap; the hybrid has been named Arbutus × andrachnoides (syn. A. × hybrida, or A. andrachne × unedo), inheriting traits of both parent species, though fruits are not usually borne freely, and as a hybrid is unlikely to breed true from seed.

New World

  • Arbutus arizonica
    Arbutus arizonica
    Arbutus arizonica is a tree species of southwestern North America, mostly in Mexico but also the United States, from the Sierra Madre Occidental cordillera, in the Ericaceae family...

    (Arizona Madrone). New Mexico
    New Mexico
    New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

    , Arizona
    Arizona
    Arizona ; is a state located in the southwestern region of the United States. It is also part of the western United States and the mountain west. The capital and largest city is Phoenix...

     and western Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

     south to Jalisco
    Jalisco
    Jalisco officially Estado Libre y Soberano de Jalisco is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is located in Western Mexico and divided in 125 municipalities and its capital city is Guadalajara.It is one of the more important states...

    .
  • Arbutus glandulosa
    Arbutus glandulosa
    Arbutus glandulosa is a species of an Arbutus plant in the Ericaceae family. It is found in Guatemala and Mexico.-Conservation:Arbutus glandulosa is threatened by habitat loss, and is a Conservation dependent plant species.-References:...

    . Central and southern Mexico.
  • Arbutus menziesii (Pacific Madrone). West coast of North America
    North America
    North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

     from southern British Columbia
    British Columbia
    British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

     to central California
    California
    California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

    , on the west slopes of the Sierra Nevada and Pacific Coast Range mountains.
  • Arbutus peninsularis. Baja California Peninsula
    Baja California Peninsula
    The Baja California peninsula , is a peninsula in northwestern Mexico. Its land mass separates the Pacific Ocean from the Gulf of California. The Peninsula extends from Mexicali, Baja California in the north to Cabo San Lucas, Baja California Sur in the south.The total area of the Baja California...

     of Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

    .
  • Arbutus tessellata. Mexico
  • Arbutus xalapensis
    Arbutus xalapensis
    Arbutus xalapensis, commonly known as the Texas Madrone or Texas Madroño, is a species of flowering plant in the heather family that is native to the southwestern United States , south through most of Mexico and Central America to Nicaragua...

    (Texas Madrone; syn. A. texana). Texas
    Texas
    Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

    , New Mexico
    New Mexico
    New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

     and northeastern Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

    .

Hybrids

  • Arbutus 'Marina' (Marina Strawberry Tree), unknown parentage although Arbutus unedo variety rubra is undoubtedly one of the parents. It is also likely that either or both of A. canariensis and A. andrachne conferred not only structural size but leaf size and smoothness of bark. The parent species likely experienced different blooming times, as the hybrid blooms nearly continuously and sets copious fruit — a factor making this lovely tree inappropriate for planting over hardscapes as the uneaten fruits create a slippery mess.

Natural history

Arbutus species are used as food plants by some Lepidoptera
Lepidoptera
Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

 species including Emperor Moth, Pavonia pavonia.

Uses and symbolism

Several species are widely cultivated as ornamental plant
Ornamental plant
Ornamental plants are plants that are grown for decorative purposes in gardens and landscape design projects, as house plants, for cut flowers and specimen display...

s outside of their natural ranges, though cultivation is often difficult due to their intolerance of root disturbance. The hybrid Arbutus 'Marina' is much more adaptable and thrives under garden conditions.

The Arbutus unedo tree makes up part of the coat of arms
Coat of arms of Madrid
The Coat of arms of Madrid has its origin in the Middle Ages, although it acquired its present look in 1967. In 2004 a decision was taken to create a logo from the shield which is used at the moment like symbol of the city and its council...

 (El oso y el madroño, The Bear and the Strawberry Tree) of the city of Madrid
Madrid
Madrid is the capital and largest city of Spain. The population of the city is roughly 3.3 million and the entire population of the Madrid metropolitan area is calculated to be 6.271 million. It is the third largest city in the European Union, after London and Berlin, and its metropolitan...

, Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

. In the center of the city (Puerta del Sol) there is a statue of a bear eating the fruit of the Madroño tree. The image appears on city crests, taxi cabs, man-hole covers, and other city infrastructure. The fruit of the Madroño tree ferments on the tree if left to ripen, so some of the bears become drunk from eating the berries.

The Arbutus was important to the Straits Salish people of Vancouver Island, who used arbutus bark and leaves to create medicines for colds, stomach problems, and tuberculosis, and as the basis for contraceptives. The tree also figured into certain myths of the Straits Salish.

The fruit is edible but has minimal flavour and is not widely eaten. In Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

 the fruit is sometimes distilled
Distillation
Distillation is a method of separating mixtures based on differences in volatilities of components in a boiling liquid mixture. Distillation is a unit operation, or a physical separation process, and not a chemical reaction....

 (legally or not) into a potent brandy known as medronho
Medronho
The Aquardente de Medronhos is a strong spirit, a traditional fruit brandy from Portugal, obtained from the fruit of the Medronho tree, Arbutus unedo, also called strawberry tree...

. In Madrid
Madrid
Madrid is the capital and largest city of Spain. The population of the city is roughly 3.3 million and the entire population of the Madrid metropolitan area is calculated to be 6.271 million. It is the third largest city in the European Union, after London and Berlin, and its metropolitan...

 the fruit is distilled
Distillation
Distillation is a method of separating mixtures based on differences in volatilities of components in a boiling liquid mixture. Distillation is a unit operation, or a physical separation process, and not a chemical reaction....

 into Madroño, a sweet, fruity liqueur.

Arbutus is a great fuelwood tree since it burns hot and long. Many Pacific Northwest
Pacific Northwest
The Pacific Northwest is a region in northwestern North America, bounded by the Pacific Ocean to the west and, loosely, by the Rocky Mountains on the east. Definitions of the region vary and there is no commonly agreed upon boundary, even among Pacific Northwesterners. A common concept of the...

 states in the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 use the wood of A. menziesii primarily as a heat source, as the wood holds no value in the production of homes since it doesn't grow in straight timbers.

Myths

According to the Straits Salish, an anthropomorphic form of pitch
Pitch (resin)
Pitch is the name for any of a number of viscoelastic, solid polymers. Pitch can be made from petroleum products or plants. Petroleum-derived pitch is also called bitumen. Pitch produced from plants is also known as resin. Products made from plant resin are also known as rosin.Pitch was...

 would go fishing, but return to shore before it got too hot. One day he was too late getting back to shore and melted from the heat and several anthropomorphic trees rushed to get him - the first was Douglas Fir, who took most of the pitch, the Grand Fir
Grand Fir
Abies grandis is a fir native to the Pacific Northwest and Northern California of North America, occurring at altitudes of sea level to 1,800 m...

 received a small portion, and the Madrone received none - which is why they say it still has no pitch.

Also, according to the Great Flood legends of several tribes in the northwest, the madrone helped people survive by providing an anchor on top of a mountain. Because of this the Saanich
Saanich
The Saanich or W̱SÁNEĆ are indigenous nations from the north coast of the Olympic Peninsula in Washington, the Gulf and San Juan Islands, southern Vancouver Island and the southern edge of the Lower Mainland in British Columbia.*BOḰEĆEN – Pauquachin...

people do not burn madrone out of thanks for saving them.

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