Piping Plover
Encyclopedia
The Piping Plover is a small sand-colored, sparrow-sized shorebird
Wader
Waders, called shorebirds in North America , are members of the order Charadriiformes, excluding the more marine web-footed seabird groups. The latter are the skuas , gulls , terns , skimmers , and auks...

 that nests and feeds along coastal sand and gravel beaches in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

. The adult has yellow-orange legs, a black band across the forehead from eye to eye, and a black ring around the neck. This chest band is usually thicker in males during the breeding season, and it's the only reliable way to tell the sexes apart. It is difficult to see when standing still as it blends well with open, sandy beach habitats. It typically runs in short starts and stops.

There are 2 subspecies of Piping Plovers: the eastern population is known as Charadrius melodus melodus and the mid-west population is known as Charadrius melodus circumcinctus. The bird's name is derived from its plaintive bell-like whistles which are often heard before the bird is visible.

Total population is currently estimated at about 6,410 individuals. A preliminary estimate showed 3,350 birds in 2003 on the Atlantic Coast alone, 52% of the total. The population has been increasing since 1991.

Their breeding habitat includes beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

es or sand flats on the Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 coast, the shores of the Great Lakes
Great Lakes
The Great Lakes are a collection of freshwater lakes located in northeastern North America, on the Canada – United States border. Consisting of Lakes Superior, Michigan, Huron, Erie, and Ontario, they form the largest group of freshwater lakes on Earth by total surface, coming in second by volume...

, and in the mid-west of Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

 and the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

. They nest on sand
Sand
Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.The composition of sand is highly variable, depending on the local rock sources and conditions, but the most common constituent of sand in inland continental settings and non-tropical coastal...

y or gravel beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

es or shoal
Shoal
Shoal, shoals or shoaling may mean:* Shoal, a sandbank or reef creating shallow water, especially where it forms a hazard to shipping* Shoal draught , of a boat with shallow draught which can pass over some shoals: see Draft...

s. These shorebirds forage for food on beaches, usually by sight, moving across the beaches in short bursts. Generally, Piping Plovers will forage for food around the high tide
Tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the moon and the sun and the rotation of the Earth....

 wrack zone
Wrack zone
The wrack zone is part of the shore just above the mean high tide line where kelp is deposited on the sand. This area is identified by the piles of kelp and other debris , and is often located on a slight "shelf" above the moist sand that slopes down toward the water....

 and along the waters edge. They mainly eat insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s, marine worm
Marine worm
Any worm that lives in a marine environment is considered a marine worm. Marine worms are found in several different phyla, including the Platyhelminthes, Nematoda, Annelida , Chaetognatha, Hemichordata, and Phoronida....

s, and crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s.

Description

The Piping Plover is a stout bird with a large rounded head, a short thick neck, and a stubby bill. It is a sand-colored, dull gray/khaki, sparrow
Sparrow
The sparrows are a family of small passerine birds, Passeridae. They are also known as true sparrows, or Old World sparrows, names also used for a genus of the family, Passer...

-sized shorebird. The adult has yellow-orange legs, a black band across the forehead from eye to eye, and a black ring around the neck during the breeding season. During nonbreeding season, the black bands become less pronounced. Its bill is orange with a black tip. It ranges from 15 – in length, with a wingspan of 35 – and a mass of 42 –.

Subspecies

Two subspecies are recognized, including nominate C. m. melodus of the Atlantic Coast
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 and C. m. circumcinctus of the Great Plains
Great Plains
The Great Plains are a broad expanse of flat land, much of it covered in prairie, steppe and grassland, which lies west of the Mississippi River and east of the Rocky Mountains in the United States and Canada. This area covers parts of the U.S...

. On average, circumcinctus is darker overall with more contrastingly dark cheeks and lores. Breeding male circumcinctus shows more extensive black on forehead and bill-base and more often shows complete breast-bands. Some overlap exists.

Vocalizations

Flight call is a soft, whistled peep given by standing and flying birds. Frequently heard alarm call is a soft pee-werp, which the second syllable lower pitched. Male's display song is a repeated, high-pitched pirp, pirp, pirp, pirp, pirp... or more drawn-out pooeep, pooeep, pooeep...,and once in a while ka-weep, ka-weep often repeated 40 or more times per flight. Display flight is often in the shape of a figure eight[4]

Habitat

The Piping Plover lives the majority of its life on open sand
Sand
Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.The composition of sand is highly variable, depending on the local rock sources and conditions, but the most common constituent of sand in inland continental settings and non-tropical coastal...

y beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

es or rocky shore
Rocky shore
A rocky shore is an intertidal area of seacoasts where solid rock predominates. Rocky shores are biologically rich environments, and make the ideal natural laboratory for studying intertidal ecology and other biological processes...

s, often in high, dry sections away from water. They can be found on the Atlantic Coast
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 of the U.S. and Canada on the ocean or bay beaches and on the Great Lakes
Great Lakes
The Great Lakes are a collection of freshwater lakes located in northeastern North America, on the Canada – United States border. Consisting of Lakes Superior, Michigan, Huron, Erie, and Ontario, they form the largest group of freshwater lakes on Earth by total surface, coming in second by volume...

 shores. It builds its nests higher on the shore near beach grass and other objects. It is very rare to see a Piping Plover anywhere outside of sand or rocky beaches/shores while not migrating.

Migration and breeding

Piping Plovers migrate
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 north in the summer and winters to the south on the Gulf of Mexico
Gulf of Mexico
The Gulf of Mexico is a partially landlocked ocean basin largely surrounded by the North American continent and the island of Cuba. It is bounded on the northeast, north and northwest by the Gulf Coast of the United States, on the southwest and south by Mexico, and on the southeast by Cuba. In...

, the southern Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 coast of the United States and the Caribbean
Caribbean
The Caribbean is a crescent-shaped group of islands more than 2,000 miles long separating the Gulf of Mexico and the Caribbean Sea, to the west and south, from the Atlantic Ocean, to the east and north...

. They begin migrating north beginning in mid-March. Their breeding grounds extend from southern Newfoundland
Newfoundland and Labrador
Newfoundland and Labrador is the easternmost province of Canada. Situated in the country's Atlantic region, it incorporates the island of Newfoundland and mainland Labrador with a combined area of . As of April 2011, the province's estimated population is 508,400...

 south to the northern parts of South Carolina
South Carolina
South Carolina is a state in the Deep South of the United States that borders Georgia to the south, North Carolina to the north, and the Atlantic Ocean to the east. Originally part of the Province of Carolina, the Province of South Carolina was one of the 13 colonies that declared independence...

. They begin mating
Mating
In biology, mating is the pairing of opposite-sex or hermaphroditic organisms for copulation. In social animals, it also includes the raising of their offspring. Copulation is the union of the sex organs of two sexually reproducing animals for insemination and subsequent internal fertilization...

 and nesting on the beach in mid-April.

Males will begin claiming territories and pairing up in late March. When pairs are formed the male begins digging out several scrapes (nests
Bird nest
A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the...

) along the high shore near the beach grass line. The males also perform elaborate courtship ceremonies, including stone tossing and courtship flights featuring repeated dives. Scrapes, small depressions in the sand dug by kicking the sand, are often in the same area that Least Terns choose to colonize. Females will sit and evaluate the scrapes. The female will choose a good scrape and will decorate the nest with shells and debris to camouflage
Camouflage
Camouflage is a method of concealment that allows an otherwise visible animal, military vehicle, or other object to remain unnoticed, by blending with its environment. Examples include a leopard's spotted coat, the battledress of a modern soldier and a leaf-mimic butterfly...

 it. Once a scrape is seen as sufficient, the female will allow the male to copulate with her. The male begins a mating ritual of standing upright and "marching" towards the female, puffing himself up and quickly stomping his legs. If the female had seen the scrape as adequate she will allow the male to stand on her back and copulation occurs within a few minutes.

Most first time nest attempts in each breeding season are 4 egg nests. Nests appear as early as mid-to-late April. Females lay one egg every other day. Second, third and sometimes fourth nesting attempts may only have three or two eggs. Incubation of the nest is shared by both the male and the female. Incubation is generally 27 days and eggs usually all hatch on the same day.

Many conservationists use exclosure
Exclosure
An exclosure, in an area being used extensively for grazing, is a limited area from which unwanted animals, for example browsing animals, such as domestic cattle or wildlife, such as deer, are excluded by fencing or other means...

s, such as round turkey wire cages with screened top, to protect the nests from predators during incubation. These allow the adults to move in and out but stop predators from getting to the eggs. When the chicks hatch many areas will put up snow fencing to restrict driving and pets for the safety of the chicks. The threats to nests include predators such as crow
Crow
Crows form the genus Corvus in the family Corvidae. Ranging in size from the relatively small pigeon-size jackdaws to the Common Raven of the Holarctic region and Thick-billed Raven of the highlands of Ethiopia, the 40 or so members of this genus occur on all temperate continents and several...

s, cat
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

s, racoon
Racoon
Racoon is a Dutch rock band, formed in 1997. Their first big appearance was at the 1999 Noorderslagfestival. First album Till Monkeys Fly appeared in January 2000, produced by Michael Schoots . The first single, Feel Like Flying, became a hit and got a lot of airplay on the Dutch radiostation 3FM...

s, fox
Fox
Fox is a common name for many species of omnivorous mammals belonging to the Canidae family. Foxes are small to medium-sized canids , characterized by possessing a long narrow snout, and a bushy tail .Members of about 37 species are referred to as foxes, of which only 12 species actually belong to...

 and various other animals to less extents. Ghost Crab
Ghost crab
Ghost crabs, also called sand crabs, are crabs of the genus Ocypode, common shore crabs in many countries. Characteristics of the genus include one claw being larger than the other, but this difference is not as marked as in male fiddler crabs....

s have been noted as being a possible predator of young chicks. These "exclosures" are what protect the nest from predators. Exclosures are not always used; sometimes they will draw more attention to the nest than they would without the exclosure. Other natural hazards to the eggs and chicks include storms, high winds, and abnormal high tides. Various human disturbances have caused the abandonment of nests and chicks as well. As a precaution it is best to stay away from any bird that appears distressed to prevent any unintended consequences.
After a chick hatches it is able to feed within hours. The adults' role is then to protect them from the elements by brooding them. They also alert them to any danger. Like many other species of plovers, adult Piping Plovers will often feign a "broken wing display
Distraction display
Distraction displays, also known as deflection display, diversionary display or paratrepsis, are anti-predator behaviours used to attract the attention of an enemy away from an object, typically the nest or young, that is being protected. They are particularly well known in birds but noted also in...

", drawing attention to themselves and away from the chicks when a predator may be threatening the chicks' safety. The "broken wing display" is also used during the nesting period to distract predators from the nest. A major defense mechanism in the chicks is their ability to blend in with the sand. It takes about 30 days before a chick achieves flight capability. They must be able to fly at least 50 yards before they can be considered as fledglings.

Migration south begins in August for some adults and fledglings, and by mid-September most Piping Plovers have headed south for winter.

Behavior

An inconspicuous bird of dry sandy beaches. Breeds in open sand, gravel, or shell-strewn beaches and alkali flats. Nest site is typically near small clumps of grass
Grass
Grasses, or more technically graminoids, are monocotyledonous, usually herbaceous plants with narrow leaves growing from the base. They include the "true grasses", of the Poaceae family, as well as the sedges and the rushes . The true grasses include cereals, bamboo and the grasses of lawns ...

, drift, or other windbreak. In winter prefers sand beaches and mudflat
Mudflat
Mudflats or mud flats, also known as tidal flats, are coastal wetlands that form when mud is deposited by tides or rivers. They are found in sheltered areas such as bays, bayous, lagoons, and estuaries. Mudflats may be viewed geologically as exposed layers of bay mud, resulting from deposition of...

s. Migrants seldom seen inland but occasionally show up at lake shores, river bars, or alkali flats. Forages visually in typical plover fashion, employing and run-stop-scan technique. Captures prey by leaning forward and picking at surface. Also employs a "foot-tremble" feeding method, causing prey to move and become more conspicuous. Feeds by day and night. Eats a wide variety of aquatic marine worm
Marine worm
Any worm that lives in a marine environment is considered a marine worm. Marine worms are found in several different phyla, including the Platyhelminthes, Nematoda, Annelida , Chaetognatha, Hemichordata, and Phoronida....

s, insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s, mollusks, and crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s. Seldom found in large numbers except at a few favored wintering or staging sites, where numbers sometimes reach 100 or more. More typically seen in pairs or in groups of 3 or 4. When approached, more often runs than flies. Very aggressive when nesting.

Status

The Piping Plover is globally threatened and endangered; it is uncommon and local within its range. The piping plover has been listed by the United States as "endangered" in the Great Lakes
Great Lakes
The Great Lakes are a collection of freshwater lakes located in northeastern North America, on the Canada – United States border. Consisting of Lakes Superior, Michigan, Huron, Erie, and Ontario, they form the largest group of freshwater lakes on Earth by total surface, coming in second by volume...

 region and "threatened" in the remainder of its breeding range. While it is federally threatened, the Piping Plover has been listed as state endangered in the states of Illinois, Indiana, Michigan, Minnesota, Nebraska, New York, New Jersey, Ohio, Pennsylvania, and Wisconsin.

In Eastern Canada, the Piping Plover is only found on coastal beaches. In 1985 it was declared an endangered species by the Committee on the Status of Endangered Wildlife in Canada. A large population in Ontario
Ontario
Ontario is a province of Canada, located in east-central Canada. It is Canada's most populous province and second largest in total area. It is home to the nation's most populous city, Toronto, and the nation's capital, Ottawa....

 has disappeared entirely.

Historical and current conservation

In the 19th century and early 20th century the Piping Plover was utilized for its feathers, as were many other birds at the time, as decoration in women's hats. These decorations, called plumes, became a symbol of high society, especially those from larger rare birds. This led to its initial population decline. The Migratory Bird Treaty Act of 1918
Migratory Bird Treaty Act of 1918
The Migratory Bird Treaty Act of 1918 , codified at , is a United States federal law, at first enacted in 1916 in order to implement the convention for the protection of migratory birds between the United States and Great Britain...

 helped the population recover through the 1930s. The second decline in the Piping Plover's population and range has been attributed to increased development, shoreline stabilization efforts, habitat loss and human activity near nesting sites in the decades following World War II
World War II
World War II, or the Second World War , was a global conflict lasting from 1939 to 1945, involving most of the world's nations—including all of the great powers—eventually forming two opposing military alliances: the Allies and the Axis...

. The Great Lakes populations eventually shrank to only a couple dozen.

Critical nesting habitats are now being protected to help the population during its breeding season. Populations have significantly increased since the protection programs began, but the species remains in serious danger. Current conservation strategies include identification and preservation of known nesting sites, public education, limiting or preventing pedestrian and/or off-road vehicle traffic near nests and hatched chicks, limiting predation of free-ranging cats, dogs and other pets on breeding pairs, eggs and chicks, and removal of foxes, raccoons, skunks, and other predators. Roughly $3 million a year is being spent for the U.S. Atlantic Coast population alone.

In coastal areas such as Plymouth
Plymouth Beach, Massachusetts
Plymouth Beach is a small village located in Plymouth, Massachusetts, United States. It is located directly south of Plymouth Center, and is adjacent to Plimoth Plantation...

, Cape Cod
Cape Cod
Cape Cod, often referred to locally as simply the Cape, is a cape in the easternmost portion of the state of Massachusetts, in the Northeastern United States...

, Long Island
Long Island
Long Island is an island located in the southeast part of the U.S. state of New York, just east of Manhattan. Stretching northeast into the Atlantic Ocean, Long Island contains four counties, two of which are boroughs of New York City , and two of which are mainly suburban...

, North Manitou Island
North Manitou Island
North Manitou Island is located in Lake Michigan, approximately west-northwest of Leland, Michigan. It is nearly eight miles long and over four miles wide, with of shoreline. It has a land area of 57.876 km² and has no population...

 in Lake Michigan, and most recently, Cape Hatteras National Seashore
Cape Hatteras National Seashore
Cape Hatteras National Seashore preserves the portion of the Outer Banks of North Carolina from Bodie Island to Ocracoke Island, stretching over . Included within this section of barrier islands along N.C...

 on the Outer Banks
Outer Banks
The Outer Banks is a 200-mile long string of narrow barrier islands off the coast of North Carolina, beginning in the southeastern corner of Virginia Beach on the east coast of the United States....

 of North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

, beach access to pedestrians and Off Road Vehicles (ORVs) has been limited or even sometimes prohibited near nesting Piping Plovers as a result of management plans and lawsuits filed by environmental organizations such as the National Audubon Society
National Audubon Society
The National Audubon Society is an American non-profit environmental organization dedicated to conservation. Incorporated in 1905, Audubon is one of the oldest of such organizations in the world and uses science, education and grassroots advocacy to advance its conservation mission...

 and the Southern Environmental Law Center (SELC). As with any other threatened species, conflicts arise and compromises have to be made between those protecting the species and those who use the habitat.

Increased temporary restrictions to areas near Hatteras Village
Hatteras, North Carolina
Hatteras is an unincorporated community in Dare County, North Carolina, United States, on the Outer Banks island of Hatteras. North Carolina Highway 12 passes through the community linking it to Frisco and Ocracoke . It is south of the famous Cape Hatteras and the Cape Hatteras Lighthouse in Buxton...

, Ocracoke Island
Ocracoke, North Carolina
Ocracoke is a census-designated place and unincorporated town located at the southern end of Ocracoke Island, located entirely within Hyde County, North Carolina. The population was 769 as of the 2000 census...

, and other areas within the Cape Hatteras National Seashore
Cape Hatteras National Seashore
Cape Hatteras National Seashore preserves the portion of the Outer Banks of North Carolina from Bodie Island to Ocracoke Island, stretching over . Included within this section of barrier islands along N.C...

 during the period when Piping Plovers and other protected species are nesting have been viewed as excessive by some residents, visitors, businesses, and activist groups. The Plovers, however, have seen their highest nesting success since 1988 in the years since the temporary beach closures were put into effect.

External links

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