Prunus mahaleb
Encyclopedia
Prunus mahaleb is a species of cherry
Cherry
The cherry is the fruit of many plants of the genus Prunus, and is a fleshy stone fruit. The cherry fruits of commerce are usually obtained from a limited number of species, including especially cultivars of the wild cherry, Prunus avium....

 native to central and southern Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, western and central Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

, and northwest Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

, from Morocco
Morocco
Morocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...

 north to France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

, southern Belgium
Belgium
Belgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...

, and Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

, and east to northern Pakistan
Pakistan
Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

 and Kyrgyzstan
Kyrgyzstan
Kyrgyzstan , officially the Kyrgyz Republic is one of the world's six independent Turkic states . Located in Central Asia, landlocked and mountainous, Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the southwest and China to the east...

.

It is a deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 or large shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

, growing to 2–10 m (rarely up to 12 m) tall with a trunk up to 40 cm diameter. The bark
Bark
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines and shrubs. Bark refers to all the tissues outside of the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner...

 is grey-brown, with conspicuous lenticel
Lenticel
A lenticel is an airy aggregation of cells within the structural surfaces of the stems, roots, and other parts of vascular plants. It functions as a pore, providing a medium for the direct exchange of gasses between the internal tissues and atmosphere, thereby bypassing the periderm, which would...

s on young stems, and shallowly fissured on old trunks. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are 1.5-5 cm long, 1-4 cm. wide, alternate, clustered at the end of alternately arranged twigs, ovate to cordate, pointed, have serrate edges, longitudinal venation and are glabrous and green. The petiole
Petiole (botany)
In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...

 is 5-20 mm, and may or may not have two glands. The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are fragrant, pure white, small, 8-20 mm diameter, with an 8-15 mm pedicel
Pedicel (botany)
A pedicel is a stem that attaches single flowers to the main stem of the inflorescence. It is the branches or stalks that hold each flower in an inflorescence that contains more than one flower....

; they are arranged 3-10 together on a 3-4 cm long raceme
Raceme
A raceme is a type of inflorescence that is unbranched and indeterminate and bears pedicellate flowers — flowers having short floral stalks called pedicels — along the axis. In botany, axis means a shoot, in this case one bearing the flowers. In a raceme, the oldest flowers are borne...

. The flower pollination
Pollination
Pollination is the process by which pollen is transferred in plants, thereby enabling fertilisation and sexual reproduction. Pollen grains transport the male gametes to where the female gamete are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself...

 is mainly by bee
Bee
Bees are flying insects closely related to wasps and ants, and are known for their role in pollination and for producing honey and beeswax. Bees are a monophyletic lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila...

s. The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a small thin-fleshed cherry-like drupe
Drupe
In botany, a drupe is a fruit in which an outer fleshy part surrounds a shell of hardened endocarp with a seed inside. These fruits develop from a single carpel, and mostly from flowers with superior ovaries...

 8–10 mm in diameter, green at first, turning red then dark purple to black when mature, with a very bitter flavour; flowering is in mid spring with the fruit ripening in mid to late summer.

Ecology

Prunus mahaleb occurs in thickets and open woodland on dry slopes; in central Europe at altitudes up to 1,700 m, and in highlands at 1,200-2,000 m in southern Europe.

It demonstrates selective fruit abortion, producing a high proportion of excess flowers that result in low fruit set levels. This reduces the number of "poor quality" fruit and increases the viability of its seeds.

A scientific study discovered an ecological dependence between the plant and four species of frugivorous birds in southeastern Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

; Blackbirds and Blackcap
Blackcap
The Blackcap is a common and widespread sylviid warbler which breeds throughout temperate Europe, western Asia and northwestern Africa, and winters from northwestern Europe south to tropical Africa...

s proved to be the most important seed dispersers. When Prunus mahaleb is fruiting, these birds consume the fruit almost exclusively, and disperse the seeds to the locations favourable for the tree's growth. The way in which some birds consume the fruits and the habitats those birds use may act as a selective force in determining which genetic variations of the cherry flourish.

Cultivation and uses

The plant is cultivated for a spice
Spice
A spice is a dried seed, fruit, root, bark, or vegetative substance used in nutritionally insignificant quantities as a food additive for flavor, color, or as a preservative that kills harmful bacteria or prevents their growth. It may be used to flavour a dish or to hide other flavours...

, which is fragrant and has the taste of bitter almond
Almond
The almond , is a species of tree native to the Middle East and South Asia. Almond is also the name of the edible and widely cultivated seed of this tree...

s. It is used in small quantities to sharpen sweet foods, such as the Turkish
Turkish cuisine
Turkish cuisine is largely the heritage of Ottoman cuisine, which can be described as a fusion and refinement of Central Asian, Middle Eastern and Balkan cuisines. Turkish cuisine has in turn influenced those and other neighbouring cuisines, including that of western Europe...

 sweet-bread çörek (chorak), the Greek
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

 sweet-bread tsoureki
Tsoureki
Tsoureki , çörek , panarët , choreg , or çörək is a sweet bread in the cuisines of the Balkans, Turkey, Azerbaijan, and Armenia. It is formed of braided strands of dough...

 or the Armenia
Armenia
Armenia , officially the Republic of Armenia , is a landlocked mountainous country in the Caucasus region of Eurasia...

n sweet-bread chorak. The chemical constituents are still uncertain, but the spice is prepared from the seeds, either by grinding and powdering the seed kernels, or in oil extracted from the seeds.

The wood
Wood
Wood is a hard, fibrous tissue found in many trees. It has been used for hundreds of thousands of years for both fuel and as a construction material. It is an organic material, a natural composite of cellulose fibers embedded in a matrix of lignin which resists compression...

 is hard, and is used in cabinet-making and for pipes.

The bark, wood, and seeds contain coumarin
Coumarin
Coumarin is a fragrant chemical compound in the benzopyrone chemical class, found in many plants, notably in high concentration in the tonka bean , vanilla grass , sweet woodruff , mullein , sweet grass , cassia cinnamon and sweet clover...

. They have anti-inflammatory, sedative and vasodilation effects.

Away from its native range, the species is grown as an ornamental tree for its strongly fragrant flowers, throughout temperate regions of the world; it has become naturalised
Naturalisation (biology)
In biology, naturalisation is any process by which a non-native organism spreads into the wild and its reproduction is sufficient to maintain its population. Such populations are said to be naturalised....

 in some areas, including Europe north of its native range (north to Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

 and Sweden
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

), and locally in Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

 and the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

.

A number of cultivar
Cultivar
A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

s have been selected, including 'Albomarginata', with variegated
Variegation
Variegation is the appearance of differently coloured zones in the leaves, and sometimes the stems, of plants. This may be due to a number of causes...

 foliage, 'Bommii', a dwarf with strongly pendulous branches, 'Globosa', a compact dwarf clone, 'Pendula', with drooping branching, and 'Xanthocarpa' with yellow fruit.

Etymology

The plant was described scientifically in the first edition of Linnaeus' Species Plantarum
Species Plantarum
Species Plantarum was first published in 1753, as a two-volume work by Carl Linnaeus. Its prime importance is perhaps that it is the primary starting point of plant nomenclature as it exists today. This means that the first names to be considered validly published in botany are those that appear...

(1753). The specific name was generally in use for the plant in a number of different languages, with slight differences in spelling. When it first began to be used is unknown, as it was to the contemporaries and post-contemporaries of Linnaeus. It does not appear in ancient literature. The Oxford English dictionary does offer an etymology: mahaleb comes from Arabic
Arabic language
Arabic is a name applied to the descendants of the Classical Arabic language of the 6th century AD, used most prominently in the Quran, the Islamic Holy Book...

محلب al-mahlab, probably meaning "a kind of perfume or sweet smell".
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