The Guanay Cormorant or Guanay Shag (Phalacrocorax bougainvillii or Leucocarbo bougainvillii) is a member of the cormorant

Cormorant

The bird family Phalacrocoracidae is represented by some 40 species of cormorants and shags. Several different classifications of the family have been proposed recently, and the number of genera is disputed.- Names :...

 family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 found in on the Pacific

Pacific Ocean

The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

 coast of Peru

Peru

Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

 and northern Chile

Chile

Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

. (The Argentinian

Argentina

Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

 population on the Patagonia

Patagonia

Patagonia is a region located in Argentina and Chile, integrating the southernmost section of the Andes mountains to the southwest towards the Pacific ocean and from the east of the cordillera to the valleys it follows south through Colorado River towards Carmen de Patagones in the Atlantic Ocean...

n Atlantic coast appears to be extinct.) After breeding it spreads south to southern parts of Chile and north to Ecuador

Ecuador

Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

, and has also been recorded as far north as Panama

Panama

Panama , officially the Republic of Panama , is the southernmost country of Central America. Situated on the isthmus connecting North and South America, it is bordered by Costa Rica to the northwest, Colombia to the southeast, the Caribbean Sea to the north and the Pacific Ocean to the south. The...

 and Colombia

Colombia

Colombia, officially the Republic of Colombia , is a unitary constitutional republic comprising thirty-two departments. The country is located in northwestern South America, bordered to the east by Venezuela and Brazil; to the south by Ecuador and Peru; to the north by the Caribbean Sea; to the...

 – probably a result of mass dispersal due to food shortage in El Niño years. Its major habitat

Habitat (ecology)

A habitat is an ecological or environmental area that is inhabited by a particular species of animal, plant or other type of organism...

s include shallow seawater and rocky shores.

The Guanay Cormorant is similar in coloration to the Rock Cormorant, Phalacrocorax magellanicus, but larger, measuring 78 cm from the tip of the bill

Beak

The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 to the end of the tail. Its bill is greyish with some red at the base. The face is red with a green eye-ring. It has roseate feet. The head

Head

In anatomy, the head of an animal is the rostral part that usually comprises the brain, eyes, ears, nose and mouth . Some very simple animals may not have a head, but many bilaterally symmetric forms do....

, neck

Neck

The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...

 and back are black as are the outer parts of the thigh

Thigh

In humans the thigh is the area between the pelvis and the knee. Anatomically, it is part of the lower limb.The single bone in the thigh is called the femur...

s. The throat

Throat

In vertebrate anatomy, the throat is the anterior part of the neck, in front of the vertebral column. It consists of the pharynx and larynx...

 patch, breast

Breast

The breast is the upper ventral region of the torso of a primate, in left and right sides, which in a female contains the mammary gland that secretes milk used to feed infants.Both men and women develop breasts from the same embryological tissues...

 and belly

Abdomen

In vertebrates such as mammals the abdomen constitutes the part of the body between the thorax and pelvis. The region enclosed by the abdomen is termed the abdominal cavity...

 are white. In breeding plumage

Plumage

Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

 it has a few white feathers on the sides of the head and neck.

Breeding occurs year round with a peak in November and December. The nest

Bird nest

A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the...

 is built of guano

Guano

Guano is the excrement of seabirds, cave dwelling bats, and seals. Guano manure is an effective fertilizer due to its high levels of phosphorus and nitrogen and also its lack of odor. It was an important source of nitrates for gunpowder...

 on flat surfaces on offshore islands or remote headlands. There are up to 3 nests per square meter in high-density colonies

Bird colony

A bird colony is a large congregation of individuals of one or more species of bird that nest or roost in close proximity at a particular location. Many kinds of birds are known to congregate in groups of varying size; a congregation of nesting birds is called a breeding colony...

. The Guanay Cormorant lays two or three eggs

Egg (biology)

An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

 of approximately 63x40 mm in size.

It feeds mainly on the Peruvian anchoveta

Peruvian anchoveta

The Peruvian anchoveta is a fish of the anchovy family, Engraulidae.Anchoveta are pelagic fish in the southeastern Pacific Ocean, and are regularly caught on the coasts of Peru, and Chile. They live for up to 4 years, reaching 20 cm, with recruitment occurring after only about 6 months when...

, Engraulis ringens, and the Peruvian silverside, Odontesthes regia, which thrive in the cold Humboldt Current

Humboldt Current

The Humboldt Current , also known as the Peru Current, is a cold, low-salinity ocean current that flows north-westward along the west coast of South America from the southern tip of Chile to northern Peru. It is an eastern boundary current flowing in the direction of the equator, and can extend...

. The Guanay Cormorant is the main producer of guano

Guano

Guano is the excrement of seabirds, cave dwelling bats, and seals. Guano manure is an effective fertilizer due to its high levels of phosphorus and nitrogen and also its lack of odor. It was an important source of nitrates for gunpowder...

.

Habitat

Habitat (ecology)

A habitat is an ecological or environmental area that is inhabited by a particular species of animal, plant or other type of organism...

 loss and degradation and over-fishing have resulted in a steady decline of the population of about 30% from an estimated figure of 3 million birds in 1984. This species is listed as NT (near threatened) by IUCN.

The scientific name commemorates the French explorer Louis Antoine de Bougainville

Louis Antoine de Bougainville

Louis-Antoine, Comte de Bougainville was a French admiral and explorer. A contemporary of James Cook, he took part in the French and Indian War and the unsuccessful French attempt to defend Canada from Britain...

.