List of lemur species
Encyclopedia
Lemur
Lemur
Lemurs are a clade of strepsirrhine primates endemic to the island of Madagascar. They are named after the lemures of Roman mythology due to the ghostly vocalizations, reflective eyes, and the nocturnal habits of some species...

s are strepsirrhine
Strepsirrhini
The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

 primate
Primate
A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...

s, all species of which are endemic to Madagascar
Madagascar
The Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...

. They include the smallest primate in the world, Madame Berthe's mouse lemur
Madame Berthe's Mouse Lemur
Madame Berthe's mouse lemur or Berthe's mouse lemur is the smallest of the mouse lemurs and the smallest primate in the world; the average body length is and seasonal weight is around...

, which weighs 30 grams (1.1 oz), and range up to the size of the indri
Indri
The indri , also called the babakoto, is one of the largest living lemurs. It is a diurnal tree-dweller related to the sifakas and, like all lemuroids, it is native to Madagascar.- Etymology :...

, which can weigh as much as 9.5 kilograms (20.9 lb). However, recently extinct species grew much larger. As of 2010, five families
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

, 15 genera
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

, and 101 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 and subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 of lemur
Lemur
Lemurs are a clade of strepsirrhine primates endemic to the island of Madagascar. They are named after the lemures of Roman mythology due to the ghostly vocalizations, reflective eyes, and the nocturnal habits of some species...

 are formally recognized. Of the 101 species and subspecies, the International Union for Conservation of Nature (IUCN) has classified
IUCN Red List
The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

 eight as Critically Endangered, 18 as Endangered, 15 as Vulnerable, four as Near Threatened, eight as Least Concern, 41 as Data Deficient, and seven have yet to be evaluated. From 2000 through 2008, 39 new species were described and nine other taxa resurrected.

The number of lemur species is likely to continue growing in the coming years, as field studies, cytogenetic
Cytogenetics
Cytogenetics is a branch of genetics that is concerned with the study of the structure and function of the cell, especially the chromosomes. It includes routine analysis of G-Banded chromosomes, other cytogenetic banding techniques, as well as molecular cytogenetics such as fluorescent in situ...

 and molecular genetic
Molecular genetics
Molecular genetics is the field of biology and genetics that studies the structure and function of genes at a molecular level. The field studies how the genes are transferred from generation to generation. Molecular genetics employs the methods of genetics and molecular biology...

 research continues. There is not complete agreement over the latest revisions to lemur taxonomy
Taxonomy
Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...

, with some experts preferring an estimated 50 lemur species. The debates are likely to continue, as some scholars label the explosive growth of species numbers as taxonomic inflation
Taxonomic inflation
Taxonomic inflation is a pejorative term for what is perceived to be an excessive increase in the number of recognised taxa in a given context, due not to the discovery of new taxa but rather to putatively arbitrary changes to how taxa are delineated....

. In many cases, classifications will ultimately depend upon which species concept is used. In the case of the lemurs of Madagascar, which have suffered extensively from deforestation
Deforestation
Deforestation is the removal of a forest or stand of trees where the land is thereafter converted to a nonforest use. Examples of deforestation include conversion of forestland to farms, ranches, or urban use....

 and habitat fragmentation
Habitat fragmentation
Habitat fragmentation as the name implies, describes the emergence of discontinuities in an organism's preferred environment , causing population fragmentation...

, nearly 25% of all species are either Endangered or Critically Endangered, most have yet to be extensively studied, and nearly all populations are in decline. For these reasons, taxonomists
Taxonomy
Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...

 and conservationist
Conservation movement
The conservation movement, also known as nature conservation, is a political, environmental and a social movement that seeks to protect natural resources including animal, fungus and plant species as well as their habitat for the future....

s favor splitting them into separate species to develop an effective strategy for the conservation of the full range of lemur diversity. Implicitly, this means that full species status will help grant genetically distinct populations added environmental protection.

At least 17 species and eight genera are believed to have become extinct
Extinction
In biology and ecology, extinction is the end of an organism or of a group of organisms , normally a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point...

 in the 2,000 years since humans first arrived in Madagascar. All known extinct species were large, ranging in weight from 10 to 200 kg (22 to 440.9 lb). The largest known subfossil lemur
Subfossil lemur
Subfossil lemurs are lemurs from Madagascar that are represented by recent remains dating from nearly 26,000 years ago to approximately 560 years ago. They include both living and extinct species, although the term more frequently refers to the extinct giant lemurs...

 was Archaeoindris fontoynonti, a giant sloth lemur
Palaeopropithecinae
The sloth lemurs is an extinct clade of lemurs that includes four genera. Despite their common names, members of Palaeopropithecidae were not closely related to sloths but were closely related to the other lemur species....

, which weighed more than a modern female gorilla
Gorilla
Gorillas are the largest extant species of primates. They are ground-dwelling, predominantly herbivorous apes that inhabit the forests of central Africa. Gorillas are divided into two species and either four or five subspecies...

. The extinction of the largest lemurs is often attributed to predation
Predation
In ecology, predation describes a biological interaction where a predator feeds on its prey . Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of its prey and the eventual absorption of the prey's tissue through consumption...

 by humans and possibly habitat destruction
Habitat destruction
Habitat destruction is the process in which natural habitat is rendered functionally unable to support the species present. In this process, the organisms that previously used the site are displaced or destroyed, reducing biodiversity. Habitat destruction by human activity mainly for the purpose of...

. Since all extinct lemurs were not only large (and thus ideal prey species), but also slow-moving (and thus more vulnerable to human predation), their presumably slow-reproducing and low-density populations were least likely to survive the introduction of humans. Gradual changes in climate have also been blamed, and may have played a minor role; however since the largest lemurs also survived the climatic changes from previous ice age
Ice age
An ice age or, more precisely, glacial age, is a generic geological period of long-term reduction in the temperature of the Earth's surface and atmosphere, resulting in the presence or expansion of continental ice sheets, polar ice sheets and alpine glaciers...

s and only disappeared following the arrival of humans, it is unlikely that climatic change was largely responsible.

There is strong evidence of extensive declines in extant populations since the introductions of humans, particularly among the larger and more specialized lemurs. As long as habitats continue to shrink, degrade and fragment, extinctions are likely to continue.

Taxonomic classification

Lemurs are classified under eight families, three of which are extinct:
  • Family: †Archaeolemuridae : monkey lemurs
    • Genus: †Archaeolemur
      Archaeolemur
      Archaeolemur is an extinct genus of lemurs that includes two species, Archaeolemur edwardsi and A. majori.The genus was widespread throughout Madagascar through much of the Holocene epoch, and its remains are often abundant at fossil sites across the island. The wide geographical and temporal range...

      (2 extinct species)
    • Genus: †Hadropithecus
      Hadropithecus
      Hadropithecus is a medium-sized, extinct genus of lemur, or strepsirrhine primate, from Madagascar that includes a single species, Hadropithecus stenognathus. Due to its rarity and lack of sufficient skeletal remains, it is one of the least understood of the extinct lemurs...

      (1 extinct species)
  • Family: Cheirogaleidae
    Cheirogaleidae
    Cheirogaleidae is the family of strepsirrhine primates that contains the various dwarf and mouse lemurs. Like all other lemurs, cheirogaleids live exclusively on the island of Madagascar.-Characteristics:...

    • Genus: Allocebus: hairy-eared dwarf lemur (1 extant species)
    • Genus: Cheirogaleus: dwarf lemurs (5 extant species)
    • Genus: Microcebus: mouse lemurs (18 extant species)
    • Genus: Mirza: giant mouse lemurs (2 extant species)
    • Genus: Phaner: fork-marked lemurs (4 extant species)
  • Family: Daubentoniidae: aye-aye
    • Genus: Daubentonia (1 extant species, 1 extinct species)
  • Family: Indriidae
    Indriidae
    The Indriidae are a family of strepsirrhine primates. They are medium to large sized lemurs with only four teeth in the toothcomb instead of the usual six...

    • Genus: Avahi: woolly lemurs (9 extant species)
    • Genus: Indri
      Indri
      The indri , also called the babakoto, is one of the largest living lemurs. It is a diurnal tree-dweller related to the sifakas and, like all lemuroids, it is native to Madagascar.- Etymology :...

      : indri (1 extant species)
    • Genus: Propithecus: sifakas (9 extant species)
  • Family: Lemuridae
    Lemuridae
    Lemuridae is a family of prosimian primates native to Madagascar, and one of five families commonly known as lemurs. These animals were thought to be the evolutionary predecessors of monkeys and apes, but this is no longer considered correct...

    • Genus: Eulemur: true lemurs (12 extant species)
    • Genus: Hapalemur: bamboo lemurs (5 extant species, 3 extant subspecies)
    • Genus: Lemur
      Ring-tailed Lemur
      The ring-tailed lemur is a large strepsirrhine primate and the most recognized lemur due to its long, black and white ringed tail. It belongs to Lemuridae, one of five lemur families. It is the only member of the Lemur genus. Like all lemurs it is endemic to the island of Madagascar...

      : ring-tailed lemur (1 extant species)
    • Genus: †Pachylemur
      Pachylemur
      Pachylemur is an extinct genus of lemur most closely related to the ruffed lemurs . Its two representative species, Pachylemur insignis and Pachylemur jullyi, are only known from subfossil remains found at sites in central and southwestern Madagascar...

      (2 extinct species)
    • Genus: Prolemur: greater bamboo lemur (1 extant species)
    • Genus: Varecia: ruffed lemurs (2 extant species, 3 extant subspecies)
  • Family: Lepilemuridae: sportive lemurs
    • Genus: Lepilemur (26 extant species)
  • Family: †Megaladapidae: koala lemurs
    • Genus: †Megaladapis
      Megaladapis
      Koala lemurs, genus Megaladapis, belong to the family Megaladapidae, consisting of three extinct species of lemurs that once inhabited the island of Madagascar. The largest measured between in length.-Appearance and habits:...

      (3 extinct species)
  • Family: †Palaeopropithecidae: sloth lemurs
    • Genus: †Archaeoindris
      Archaeoindris
      Archaeoindris fontoynonti is an extinct species of Malagasy lemur that was the largest primate to evolve on Madagascar. It weighed about and measured around 1.5m in height, more than a silverback gorilla. Archaeoindris is one of eight known members of the Palaeopropithecinae subfamily...

      (1 extinct species)
    • Genus: †Babakotia
      Babakotia
      Babakotia is a medium-sized, extinct genus of lemur, or strepsirrhine primate, from Madagascar that contains a single species, Babakotia radofilai. Together with Palaeopropithecus, Archaeoindris, and Mesopropithecus, it forms the family Palaeopropithecidae, commonly known as the sloth lemurs...

      (1 extinct species)
    • Genus: †Mesopropithecus
      Mesopropithecus
      Mesopropithecus is an extinct genus of small to medium-sized lemur, or strepsirrhine primate, from Madagascar that includes three species, M. dolichobrachion, M. globiceps, and M. pithecoides. Together with Palaeopropithecus, Archaeoindris, and Babakotia, it is part of the sloth...

      (3 extinct species)
    • Genus: †Palaeopropithecus (3 extinct species)


The placement of lemurs within the order Strepsirrhini is currently under debate, although both sides agree upon the same phylogenetic tree
Phylogenetic tree
A phylogenetic tree or evolutionary tree is a branching diagram or "tree" showing the inferred evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical and/or genetic characteristics...

. The following table demonstrates the differences between the two taxonomies.

3 infraorders, 2 superfamilies 1 infraorder, 2 superfamilies
  • Order Primate
    Primate
    A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...

    s
    • Suborder Strepsirrhini
      Strepsirrhini
      The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

      : non-tarsier prosimians
      • Infraorder Chiromyiformes (aye-aye)
      • Infraorder Lemuriformes
        • Superfamily Cheirogaleoidea
          • Family Cheirogaleidae
            Cheirogaleidae
            Cheirogaleidae is the family of strepsirrhine primates that contains the various dwarf and mouse lemurs. Like all other lemurs, cheirogaleids live exclusively on the island of Madagascar.-Characteristics:...

        • Superfamily Lemuroidea
          • Family †Archaeolemuridae
            Monkey lemur
            The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

          • Family Indriidae
            Indriidae
            The Indriidae are a family of strepsirrhine primates. They are medium to large sized lemurs with only four teeth in the toothcomb instead of the usual six...

          • Family Lemuridae
            Lemuridae
            Lemuridae is a family of prosimian primates native to Madagascar, and one of five families commonly known as lemurs. These animals were thought to be the evolutionary predecessors of monkeys and apes, but this is no longer considered correct...

          • Family Lepilemuridae
            Sportive lemur
            The sportive lemurs are the medium sized primates that make up the Lepilemuridae family. The family consists of only one extant genus, Lepilemur, as well as the extinct genus Megaladapis. They are closely related to the other lemurs and exclusively live on the island of Madagascar...

          • Family †Megaladapidae
          • Family †Palaeopropithecidae
      • Infraorder Lorisiformes
        Lorisiformes
        Lorisiformes are a group of primates found throughout Africa and Asia. Members of this infraorder include the galagos and the lorises. As strepsirrhines, they are related to the lemurs.* Order Primates** Suborder Strepsirrhini: non-tarsier prosimians...

    • Suborder Haplorrhini
      Haplorrhini
      The haplorhines, the "dry-nosed" primates , are members of the Haplorhini clade: the prosimian tarsiers and the anthropoids...

      : tarsiers, monkeys and apes
  • Order Primate
    Primate
    A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...

    s
    • Suborder Strepsirrhini
      Strepsirrhini
      The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

      : non-tarsier prosimians
      • Infraorder Lemuriformes
        • Superfamily Lemuroidea
          • Family †Archaeolemuridae
            Monkey lemur
            The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

          • Family Cheirogaleidae
            Cheirogaleidae
            Cheirogaleidae is the family of strepsirrhine primates that contains the various dwarf and mouse lemurs. Like all other lemurs, cheirogaleids live exclusively on the island of Madagascar.-Characteristics:...

          • Family Daubentoniidae
            Aye-aye
            The aye-aye is a lemur, a strepsirrhine primate native to Madagascar that combines rodent-like teeth and a special thin middle finger to fill the same ecological niche as a woodpecker...

          • Family Indriidae
            Indriidae
            The Indriidae are a family of strepsirrhine primates. They are medium to large sized lemurs with only four teeth in the toothcomb instead of the usual six...

          • Family Lemuridae
            Lemuridae
            Lemuridae is a family of prosimian primates native to Madagascar, and one of five families commonly known as lemurs. These animals were thought to be the evolutionary predecessors of monkeys and apes, but this is no longer considered correct...

          • Family Lepilemuridae
            Sportive lemur
            The sportive lemurs are the medium sized primates that make up the Lepilemuridae family. The family consists of only one extant genus, Lepilemur, as well as the extinct genus Megaladapis. They are closely related to the other lemurs and exclusively live on the island of Madagascar...

          • Family †Megaladapidae
          • Family †Palaeopropithecidae
        • Superfamily Lorisoidea
    • Suborder Haplorrhini
      Haplorrhini
      The haplorhines, the "dry-nosed" primates , are members of the Haplorhini clade: the prosimian tarsiers and the anthropoids...

      : tarsiers, monkeys and apes

  • Key

    Scientific name Latin binomial name, or Latin name, of the species
    Common name Common name of the species, per Wilson, et al. Mammal Species of the World (2005) or Mittermeier, et al. "Lemur Diversity in Madagascar" (2008)
    Family Family within the Suborder Strepsirrhini
    Strepsirrhini
    The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

     to which the species belongs
    Classified Date in which the species was formally described and classified, as well as the binomial authority on the species
    Average size Average size of adult members of the species, in metric
    Metric system
    The metric system is an international decimalised system of measurement. France was first to adopt a metric system, in 1799, and a metric system is now the official system of measurement, used in almost every country in the world...

     and English weight units
    Estimated size Estimated Average size of adult members of the extinct species
    Conservation status Conservation status of the species, per IUCN as of 2008
    Extinction date Year of extinction or estimated date range for subfossil
    Subfossil
    Subfossil refers to remains whose fossilization process is not complete, either for lack of time or because the conditions in which they were buried were not optimal for fossilization....

     species, in BCE/CE
    Common Era
    Common Era ,abbreviated as CE, is an alternative designation for the calendar era originally introduced by Dionysius Exiguus in the 6th century, traditionally identified with Anno Domini .Dates before the year 1 CE are indicated by the usage of BCE, short for Before the Common Era Common Era...


    Extant species

    All lemurs belong to the suborder Strepsirrhini
    Strepsirrhini
    The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

     within the order Primate
    Primate
    A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...

    s. The 101 extant species and subspecies are divided among 5 families and 15 genera. They range in weight from 30 g (1.1 oz) (Madame Berthe's mouse lemur
    Madame Berthe's Mouse Lemur
    Madame Berthe's mouse lemur or Berthe's mouse lemur is the smallest of the mouse lemurs and the smallest primate in the world; the average body length is and seasonal weight is around...

    ) to as much as 9.5 kg (20.9 lb) (indri
    Indri
    The indri , also called the babakoto, is one of the largest living lemurs. It is a diurnal tree-dweller related to the sifakas and, like all lemuroids, it is native to Madagascar.- Etymology :...

    ). Most are highly arboreal
    Arboreal locomotion
    Arboreal locomotion is the locomotion of animals in trees. In every habitat in which trees are present, animals have evolved to move in them. Some animals may only scale trees occasionally, while others are exclusively arboreal. These habitats pose numerous mechanical challenges to animals...

     and activity patterns range widely from nocturnal to diurnal to cathemeral
    Cathemeral
    A cathemeral organism is one that has sporadic and random intervals of activity during the day or night in which food is acquired, socializing with other organisms occurs, and any other activities necessary for livelihood are performed...

    . Having diversified over millions of years to fill every ecological niche
    Ecological niche
    In ecology, a niche is a term describing the relational position of a species or population in its ecosystem to each other; e.g. a dolphin could potentially be in another ecological niche from one that travels in a different pod if the members of these pods utilize significantly different food...

    , diets also vary widely, though fruit, leaves, and insects make up the majority of the diet for most species.

    Family: Cheirogaleidae

    Family Cheirogaleidae consists of the mouse lemur
    Mouse lemur
    The mouse lemurs are nocturnal lemurs of the genus Microcebus. Like all lemurs, mouse lemurs are native to Madagascar.Mouse lemurs have a combined head, body and tail length of less than , making them the smallest primates ; however, their weight fluctuates in response to daylight duration.Mouse...

    s (smallest of all primates), the dwarf lemur
    Dwarf lemur
    The dwarf lemurs are the lemurs of the genus Cheirogaleus. All of the species in this genus, like all other lemurs, are native to Madagascar.- Description :...

    s, and the fork-marked lemurs and their allies. There are 30 extant species.
    Scientific name Common name Classified Average size Conservation status References
    Allocebus trichotis Hairy-eared dwarf lemur
    Hairy-eared Dwarf Lemur
    The hairy-eared dwarf lemur , or hairy-eared mouse lemur, is a nocturnal lemur endemic to Madagascar. It is the only member of the genus Allocebus. This species is critically endangered and the population is estimated at 100-1000 individuals. They all live a single location in the northeastern part...

    1875, Günther
    Cheirogaleus crossleyi Furry-eared dwarf lemur
    Furry-eared Dwarf Lemur
    The furry-eared dwarf lemur , or Crossley's dwarf lemur, only found on the island of Madagascar, as with all other lemurs. It has a pelage coloration that is red-brown dorsally and gray ventrally. The eye-rings of this species are blackish and the ears are black inside and out. The dental formula...

    1870, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Cheirogaleus major Greater dwarf lemur
    Greater dwarf lemur
    The greater dwarf lemur , or the Geoffroy's dwarf lemur, is a lemur that is widely distributed over the primary and secondary forests near the eastern coast of Madagascar....

    1812, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Cheirogaleus medius Fat-tailed dwarf lemur
    Fat-tailed dwarf lemur
    The fat-tailed dwarf lemur , also known as the lesser dwarf lemur, western fat-tailed dwarf lemur, or spiny forest dwarf lemur, is one of the smallest primates. It is endemic to Madagascar....

    1812, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Cheirogaleus minusculus Lesser iron-gray dwarf lemur
    Lesser Iron-gray Dwarf Lemur
    The lesser iron-gray dwarf lemur , or large iron-gray dwarf lemur, is a small nocturnal lemur endemic to Madagascar.This species is only known from Ambositra....

    2000, Groves
    Colin Groves
    Colin Peter Groves is Professor of Biological Anthropology at the Australian National University in Canberra, Australia.Born in England on 24 June 1942, Colin Groves completed a Bachelor of Science at University College London in 1963, and a Doctor of Philosophy at the Royal Free Hospital School of...

    unknown
    Cheirogaleus sibreei Sibree's dwarf lemur
    Sibree's Dwarf Lemur
    Sibree's dwarf lemur is a small nocturnal lemur endemic to Madagascar.The name of this dwarf lemur commemorates the English missionary and naturalist James Sibree ....

    1896, Forsyth Major
    Charles Immanuel Forsyth Major
    Charles Immanuel Forsyth Major was a Swiss zoologist and vertebrate palaeontologist.Major was born in Glasgow and studied at Basel and Zurich Universities in Switzerland and later Göttingen in Germany...

    unknown
    Microcebus berthae Madame Berthe's mouse lemur
    Madame Berthe's Mouse Lemur
    Madame Berthe's mouse lemur or Berthe's mouse lemur is the smallest of the mouse lemurs and the smallest primate in the world; the average body length is and seasonal weight is around...

    2000, Rasoloarison et al.
    Microcebus bongolavensis Bongolava mouse lemur
    Bongolava mouse lemur
    The Bongolava mouse lemur is a species of mouse lemur endemic to Madagascar. It lives in western deciduous forest within a limited range, including Bongolava Forest and Ambodimahabibo Forest between the Sofia River and the Mahajamba River...

    2007, Olivieri et al.
    Microcebus danfossi Danfoss' mouse lemur
    Danfoss' Mouse Lemur
    Danfoss' mouse lemur , or Ambarijeby mouse lemur, is a species of mouse lemur endemic to Madagascar. It lives in western deciduous forest within a limited range between the Sofia River and the Maevarano River...

    2007, Olivieri et al.
    Microcebus griseorufus Reddish-gray mouse lemur
    Reddish-gray Mouse Lemur
    The reddish-gray mouse lemur also known as the gray-brown mouse lemur or rufous-gray mouse lemur‎, is found in Western Madagascar in the region around Beza Mahafaly Reserve, north to Lamboharana....

    1910, Kollman
    Microcebus jollyae Jolly's mouse lemur
    Jolly's Mouse Lemur
    Jolly's mouse lemur is a species of mouse lemur from Mananjary and Kianjavato in Madagascar.The species is named in honor of primatologist Alison Jolly....

    2006, Louis et al.
    Microcebus lehilahytsara Goodman's mouse lemur
    Goodman's Mouse Lemur
    Goodman's mouse lemur is a species of mouse lemur from Andasibe in eastern Madagascar. It is thought to have diverged from other mouse lemurs in the region about 2 million years ago....

    2005, Roos and Kappeler
    Microcebus macarthurii MacArthur's mouse lemur
    MacArthur's Mouse Lemur
    MacArthur's mouse lemur , or Anjiahely mouse lemur, is a species of mouse lemur known only from an Inter-River-System in the eastern Makira Natural Park in Madagascar....

    2008, Radespiel et al. unknown
    Microcebus mamiratra Claire's mouse lemur
    Claire's Mouse Lemur
    Claire's mouse lemur , or Nosy Be mouse lemur, is a newly described species of lemur from the genus of the mouse lemurs . It lives on the island Nosy Bé in the Antsiranana province of Madagascar, and on the mainland near the village of Manehoka, including Lokobe Reserve...

    2006, Andriantompohavana et al.
    Microcebus mittermeieri Mittermeier's mouse lemur
    Mittermeier's Mouse Lemur
    Mittermeier's mouse lemur is a species of mouse lemur known only from Anjanaharibe-Sud Special Reserve in Madagascar....

    2006, Louis et al.
    Microcebus murinus Gray mouse lemur
    Gray Mouse Lemur
    The gray mouse lemur , or lesser mouse lemur, is a small lemur, a type of strepsirrhine primate, found only on the island of Madagascar. Weighing , it is the largest of the mouse lemurs , a group which include the smallest primates in the world...

    1777, Miller
    John Frederick Miller
    John Frederick Miller was an English illustrator, mainly of botanical subjects.Miller was the son of the artist Johann Sebastian Müller . Miller, along with his brother James, produced paintings from the sketches made by Sydney Parkinson on James Cook's first voyage...

    Microcebus myoxinus Pygmy mouse lemur 1852, Peters
    Wilhelm Peters
    Wilhelm Karl Hartwich Peters was a German naturalist and explorer.He was assistant to Johannes Peter Müller and later curator of the Berlin Zoological Museum. In September 1842 he travelled to Mozambique via Angola. He returned to Berlin with an enormous collection of natural history specimens...

    Microcebus ravelobensis Golden-brown mouse lemur 1998, Zimmerman et al.
    Microcebus rufus Brown mouse lemur 1834, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Microcebus sambiranensis Sambirano mouse lemur
    Sambirano Mouse Lemur
    The Sambirano mouse lemur is a small, recently discovered primate, and like the other mouse lemurs can only be found on the island of Madagascar. The dorsal side is both cinnamon and rufous-cinnamon, and is grey ventrally...

    2000, Rasoloarison et al.
    Microcebus simmonsi Simmons' mouse lemur
    Simmons' Mouse Lemur
    Simmons' mouse lemur is a species of mouse lemur known only from Betampona Special Reserve and Zahamena National Park in Madagascar....

    2006, Louis et al.
    Microcebus tavaratra Northern rufous mouse lemur
    Northern Rufous Mouse Lemur
    The northern rufous mouse lemur , northern brown mouse lemur, or Tavaratra mouse lemur is found in North-western Madagascar in Ankarana Reserve....

    2000, Rasoloarison et al.
    Microcebus arnholdi Arnhold's mouse lemur
    Arnhold's Mouse Lemur
    Arnhold's mouse lemur or Montagne d’Ambre mouse lemur is a species of mouse lemur endemic to Madagascar. Its holotype was first collected on 27 November 2005, and was first described in 2008. According to genetic tests, it is genetically distinct from its closest sister taxa, the Sambirano mouse...

    2008, E. Lewis, Jr. et al.
    Microcebus margotmarshae Margot Marsh's mouse lemur
    Margot Marsh's Mouse Lemur
    Margot Marsh's mouse lemur or the Antafondro mouse lemur is a species of mouse lemur endemic to Madagascar. Its holotype was first collected on 21 May 2006, proposed in 2006 by Andriantompohavana et al., and was formally described in 2008 by E...

    2006, Andriantompohavana et al.
    Mirza coquereli Coquerel's giant mouse lemur 1867, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Mirza zaza Northern giant mouse lemur
    Northern Giant Mouse Lemur
    The northern giant mouse lemur , or northern dwarf lemur, was discovered to be a species in 2005 by Kappeler et al. from the German primates centre and the University of Göttingen. Before, both populations of giant mouse lemurs were believed to belong to one species. The northern giant mouse lemurs...

    2005, Kappeler & Roos
    Phaner electromontis Amber Mountain fork-marked lemur 1991, Groves
    Colin Groves
    Colin Peter Groves is Professor of Biological Anthropology at the Australian National University in Canberra, Australia.Born in England on 24 June 1942, Colin Groves completed a Bachelor of Science at University College London in 1963, and a Doctor of Philosophy at the Royal Free Hospital School of...

     and Tattersall
    Ian Tattersall
    Ian Tattersall is a paleoanthropologist and a curator at the American Museum of Natural History. Tattersall received his PhD from Yale University in 1971. In addition to human evolution, he has worked extensively with lemurs. He is working with The Templeton Foundation.-Selected publications:* The...

    Phaner furcifer Masoala fork-marked lemur 1839, Blainville
    Henri Marie Ducrotay de Blainville
    Henri Marie Ducrotay de Blainville was a French zoologist and anatomist.Blainville was born at Arques, near Dieppe. In about 1796 he went to Paris to study painting, but he ultimately devoted himself to natural history, and attracted the attention of Georges Cuvier, for whom he occasionally...

    Phaner pallescens Pale fork-marked lemur 1991, Groves
    Colin Groves
    Colin Peter Groves is Professor of Biological Anthropology at the Australian National University in Canberra, Australia.Born in England on 24 June 1942, Colin Groves completed a Bachelor of Science at University College London in 1963, and a Doctor of Philosophy at the Royal Free Hospital School of...

     and Tattersall
    Ian Tattersall
    Ian Tattersall is a paleoanthropologist and a curator at the American Museum of Natural History. Tattersall received his PhD from Yale University in 1971. In addition to human evolution, he has worked extensively with lemurs. He is working with The Templeton Foundation.-Selected publications:* The...

    Phaner parienti Pariente's fork-marked lemur 1991, Groves
    Colin Groves
    Colin Peter Groves is Professor of Biological Anthropology at the Australian National University in Canberra, Australia.Born in England on 24 June 1942, Colin Groves completed a Bachelor of Science at University College London in 1963, and a Doctor of Philosophy at the Royal Free Hospital School of...

     and Tattersall
    Ian Tattersall
    Ian Tattersall is a paleoanthropologist and a curator at the American Museum of Natural History. Tattersall received his PhD from Yale University in 1971. In addition to human evolution, he has worked extensively with lemurs. He is working with The Templeton Foundation.-Selected publications:* The...



    Family: Lemuridae

    Family Lemuridae consists of the ring-tailed lemur
    Ring-tailed Lemur
    The ring-tailed lemur is a large strepsirrhine primate and the most recognized lemur due to its long, black and white ringed tail. It belongs to Lemuridae, one of five lemur families. It is the only member of the Lemur genus. Like all lemurs it is endemic to the island of Madagascar...

    , the brown lemur
    Brown lemur
    True lemurs, also known as brown lemurs, are the lemurs in genus Eulemur. They are medium sized primates that live exclusively on Madagascar....

    s, the bamboo lemur
    Bamboo lemur
    The bamboo or gentle lemurs are the lemurs in genus Hapalemur. They are medium sized primates that live exclusively on Madagascar...

    s and the ruffed lemur
    Ruffed lemur
    The ruffed lemurs of the genus Varecia are strepsirrhine primates and the largest extant lemurs within the family Lemuridae. Like all living lemurs, they are found only on the island of Madagascar...

    s. There are 21 extant species and six subspecies.
    Scientific name Common name Classified Average size Conservation status References
    Eulemur albifrons White-fronted brown lemur 1796, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Eulemur cinereiceps Gray-headed lemur
    Gray-headed Lemur
    The gray-headed lemur , or gray-headed brown lemur, is a medium-sized primate, a cathemeral species of lemur in the Lemuridae family. Until a taxonomic revision in 2008, it was known as the white-collared brown lemur or white-collared lemur . It lives in south-eastern Madagascar...

    1890, Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

     and Milne-Edwards
    Henri Milne-Edwards
    Henri Milne-Edwards was an eminent French zoologist.Henri Milne-Edwards was the 27th child of William Edwards, an English planter and militia colonel in Jamaica and Elisabeth Vaux, a French. He was born in Bruges, Belgium, where his parents had retired. At that time, Bruges was a part of the...

    Eulemur collaris Collared brown lemur
    Collared Brown Lemur
    The collared brown lemur , also known as the red-collared brown lemur or red-collared lemur, is a medium-sized strepsirrhine primate and one of twelve species of brown lemur in the Lemuridae family. It is only found in south-eastern Madagascar. Like most species of lemur, it is arboreal, moving...

    1812, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Eulemur coronatus Crowned lemur
    Crowned Lemur
    The crowned lemur is a lemur that is 31–36 cm long and weighs 2 kg. Its tail is about 42–51 cm long). The crowned lemur is endemic to the dry deciduous forests of the northern tip of Madagascar. It eats a diet of mostly flowers, fruits, and leaves...

    1842, Gray
    John Edward Gray
    John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

    Eulemur flavifrons Blue-eyed black lemur 1867, Gray
    John Edward Gray
    John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

    Eulemur fulvus Common brown lemur
    Common Brown Lemur
    The common brown lemur , or brown lemur, is a species of lemur in the Lemuridae family. It is found in Madagascar and Mayotte.-Range:...

    1812, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Eulemur macaco Black lemur
    Black Lemur
    The Black Lemur is a species of lemur from the family Lemuridae. Like all lemurs, it is endemic to Madagascar. Originally, the species was thought to have two subspecies, Eulemur macaco macaco and Eulemur macaco flavifrons, both of which were elevated to species status by Mittermeier et al. in...

    1766, Linnaeus
    Carolus Linnaeus
    Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

    Eulemur mongoz Mongoose lemur
    Mongoose Lemur
    The mongoose lemur is a lemur ranging from 12 to 18 inches long plus a tail of 16 to 25 inches. The mongoose lemur lives in dry deciduous forests on the island of Madagascar as well as in the humid forests on the islands of the Comoros...

    1766, Linnaeus
    Carolus Linnaeus
    Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

    Eulemur rubriventer Red-bellied lemur
    Red-bellied Lemur
    The Red-bellied Lemur is a medium sized prosimian with a luxuriant chestnut brown coat. This lemur is endemic to eastern Madagascan rainforests and is distinguished by patches of white skin below the eyes, giving rise to a "teardrop" effect, particularly conspicuous in the male.The species, first...

    1850, I. Geoffroy
    Eulemur rufifrons Red-fronted lemur
    Red-fronted Lemur
    The red-fronted lemur , also known as the red-fronted brown lemur or southern red-fronted brown lemur, is a species of lemur from Madagascar. Until 2001, it was considered a subspecies of the common brown lemur, E. fulvus. In 2001, E...

    1833, Bennett
    Eulemur rufus Red lemur
    Red Lemur
    The red lemur , also known as the rufous brown lemur or northern red-fronted lemur, is a species of lemur from Madagascar. Until 2001, the species E. rufus was considered a subspecies of the common brown lemur, E. fulvus, after which it was classified as its own species...

    1799, Audebert
    Eulemur sanfordi Sanford's brown lemur
    Sanford's Brown Lemur
    Sanford's brown lemur , or Sanford's lemur, is a species of prosimian primate in the Lemuridae family. The Sanford's brown lemur was previously considered a subspecies of the common brown lemur but as of 2005 many of the brown lemurs have been elevated to a full species status...

    1932, Archbold
    Hapalemur alaotrensis Lac Alaotra bamboo lemur 1975, Rumpler
    Hapalemur aureus Golden bamboo lemur
    Golden Bamboo Lemur
    The Golden Bamboo Lemur or Golden Lemur is a medium sized bamboo lemur endemic to southeastern Madagascar. It is listed as an endangered species due to habitat loss. The population is declining with only about 1000 individuals...

    1987, Meier et al.
    Hapalemur griseus gilberti Beanamalao bamboo lemur 2007, Rabarivola et al.
    Hapalemur griseus griseus Eastern lesser bamboo lemur 1795, Link
    Johann Heinrich Friedrich Link
    Johann Heinrich Friedrich Link was a German naturalist and botanist.Link was born at Hildesheim as a son of the minister August Heinrich Link , who taught him the love for nature through collection of 'natural objects'...

    Hapalemur griseus ranomafanensis Ranomafana bamboo lemur 2007, Rabarivola et al.
    Hapalemur meridionalis Southern lesser bamboo lemur
    Southern Lesser Bamboo Lemur
    The southern lesser bamboo lemur , also known as the southern bamboo lemur, rusty-gray bamboo lemur, and southern gentle lemur, is species of bamboo lemur endemic to southern Madagascar near Tôlanaro. It was originally proposed as a subspecies of the eastern lesser bamboo lemur and is of similar size...

    1987, Warter, et al.
    Hapalemur occidentalis Western lesser bamboo lemur
    Western Lesser Bamboo Lemur
    The Western Lesser Bamboo Lemur , also known as the Northern Bamboo Lemur and Western Gentle Lemur, is species of bamboo lemur endemic to Madagascar. The total length of this primate is 55–67 centimeters, more than half of which is tail, and average weight is just under 1 kilogram...

    1975, Rumpler
    Lemur catta Ring-tailed lemur
    Ring-tailed Lemur
    The ring-tailed lemur is a large strepsirrhine primate and the most recognized lemur due to its long, black and white ringed tail. It belongs to Lemuridae, one of five lemur families. It is the only member of the Lemur genus. Like all lemurs it is endemic to the island of Madagascar...

    1756, Linnaeus
    Carolus Linnaeus
    Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

    Prolemur simus Greater bamboo lemur
    Greater Bamboo Lemur
    The Greater Bamboo Lemur , also known as the Broad-nosed Bamboo Lemur and the Broad-nosed Gentle Lemur, is the largest bamboo lemur, at over five pounds or nearly 2.5 kilograms. It has greyish brown fur and white ear tufts, and has a head-body length of around one and a half feet, or forty to...

    1871, Gray
    John Edward Gray
    John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

    Varecia rubra Red ruffed lemur
    Red Ruffed Lemur
    The red ruffed lemur is one of two species in the genus Varecia, the ruffed lemurs; the other is the black-and-white ruffed lemur . Like all lemurs, it is native to Madagascar and occurs only in the rainforests of Masoala, in the northeast of the island...

    1812, É. Geoffroy
    Étienne Geoffroy Saint-Hilaire
    Étienne Geoffroy Saint-Hilaire was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories...

    Varecia variegata editorum Southern black-and-white ruffed lemur
    Black-and-white ruffed lemur
    The black-and-white ruffed lemur is the more endangered of the two species of ruffed lemurs, both of which are endemic to the island of Madagascar. Despite having a larger range than the red ruffed lemur, it has a much smaller population that is spread out, living in lower population densities...

    1953, Osman Hill
    Varecia variegata subcincta White-belted black-and-white ruffed lemur
    Black-and-white ruffed lemur
    The black-and-white ruffed lemur is the more endangered of the two species of ruffed lemurs, both of which are endemic to the island of Madagascar. Despite having a larger range than the red ruffed lemur, it has a much smaller population that is spread out, living in lower population densities...

    1833, A. Smith
    Varecia variegata variegata Black-and-white ruffed lemur
    Black-and-white ruffed lemur
    The black-and-white ruffed lemur is the more endangered of the two species of ruffed lemurs, both of which are endemic to the island of Madagascar. Despite having a larger range than the red ruffed lemur, it has a much smaller population that is spread out, living in lower population densities...

    1792, Kerr
    Robert Kerr (writer)
    Robert Kerr FRS was a scientific writer and translator from Scotland.Kerr was born in Roxburghshire as the son of a jeweller. He studied medicine at the University of Edinburgh and practised at the Edinburgh Foundling Hospital as a surgeon...



    Family: Lepilemuridae

    Family Lepilemuridae consists solely of the sportive lemur
    Sportive lemur
    The sportive lemurs are the medium sized primates that make up the Lepilemuridae family. The family consists of only one extant genus, Lepilemur, as well as the extinct genus Megaladapis. They are closely related to the other lemurs and exclusively live on the island of Madagascar...

    s. As of 2010, there are 26 extant species.
    Scientific name Common name Classified Average size Conservation status References
    Lepilemur aeeclis Antafia sportive lemur 2006, Andriaholinirina et al.
    Lepilemur ahmansonorum Ahmanson's sportive lemur
    Ahmanson's Sportive Lemur
    Ahmanson's sportive lemur , or Tsiombikibo sportive lemur, is a sportive lemur endemic to Madagascar. It is a relatively small sportive lemur with a total length of about 47 to 54 centimeters, of which 23-25 centimeters are tail. Wright's sportive lemur is found in western Madagascar, living in...

    2006, Louis, Jr.
    Lepilemur ankaranensis Ankarana sportive lemur
    Ankarana Sportive Lemur
    The Ankarana sportive lemur , or Ankarana weasel lemur, is a sportive lemur endemic to Madagascar. It is one of the smaller sportive lemurs with a total length of about 53 centimeters, including 25 centimeters of tail. Average body weight is approximately 750 grams...

    1975, Rumpler & Albignac
    Lepilemur betsileo Betsileo sportive lemur
    Betsileo Sportive Lemur
    The Betsileo sportive lemur is a sportive lemur endemic to Madagascar. It is a relatively large sportive lemur with a total length of about 58 to 27 centimeters, of which 32 to 33 centimeters are tail. The Betsileo Sportive Lemur is found in eastern Madagascar, living in primary and secondary...

    2006, Louis, Jr.
    Lepilemur dorsalis Gray-backed sportive lemur
    Gray-backed Sportive Lemur
    The gray-backed sportive lemur , also known as gray's sportive lemur or back-striped sportive lemur, is a species of lemur in the Lepilemuridae family. It is endemic to Madagascar. It is threatened by habitat loss....

    1870, Gray
    John Edward Gray
    John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

    Lepilemur edwardsi Milne-Edwards' sportive lemur
    Milne-Edwards' Sportive Lemur
    The Milne-Edwards' sportive lemur , or Milne-Edwards' weasel lemur, is a species of lemur in the Lepilemuridae family. It is endemic to Madagascar. Its natural habitat is subtropical or tropical dry forests. It is threatened by habitat loss....

    1894, Forsyth Major
    Charles Immanuel Forsyth Major
    Charles Immanuel Forsyth Major was a Swiss zoologist and vertebrate palaeontologist.Major was born in Glasgow and studied at Basel and Zurich Universities in Switzerland and later Göttingen in Germany...

    Lepilemur fleuretae Fleurete's sportive lemur
    Fleurete's Sportive Lemur
    Fleurete's sportive lemur , or Andohahela sportive lemur, is a sportive lemur endemic to Madagascar. It is a medium sized sportive lemur with a total length of about 58 to 67 centimeters, of which about 30 centimeters are tail Fleurete's sportive lemur is found in southeastern Madagascar, living...

    2006, Louis, Jr.
    Lepilemur grewcockorum Grewcock's sportive lemur
    Grewcock's sportive lemur
    Grewcock's sportive lemur , or Anjiamangirana sportive lemur, is a sportive lemur endemic to Madagascar. It is a medium-sized sportive lemurs with a total length of about 55 to 63 centimeters, of which 26-30 centimeters are tail. Grewcock's sportive lemur is found in northwesetrn Madagascar,...

    2006, Louis, Jr.
    Lepilemur hubbardorum Hubbard's sportive lemur
    Hubbard's Sportive Lemur
    Hubbard's sportive lemur , or Zombitse sportive lemur, is a sportive lemur endemic to Madagascar. It has total length of about 51 to 59 centimeters, of which 23-25 centimeters are tail. Hubbard's sportive lemur is found in southwestern Madagascar, living in dry transitional forests.Originally...

    2006, Louis, Jr.
    Lepilemur jamesorum James' sportive lemur
    James' Sportive Lemur
    James' sportive lemur , or Manombo sportive lemur, is a sportive lemur endemic to Madagascar. It is one of the larger sportive lemurs with a total length of about 60 to 67 centimeters, of which 28 to 32 centimeters are tail...

    2006, Louis, Jr.
    Lepilemur leucopus White-footed sportive lemur
    White-footed Sportive Lemur
    The white-footed sportive lemur , or white-footed weasel lemur, is a species of lemur in the Lepilemuridae family. It is endemic to Madagascar. Its natural habitat is subtropical or tropical dry shrubland. It is threatened by habitat loss....

    1894, Forsyth Major
    Charles Immanuel Forsyth Major
    Charles Immanuel Forsyth Major was a Swiss zoologist and vertebrate palaeontologist.Major was born in Glasgow and studied at Basel and Zurich Universities in Switzerland and later Göttingen in Germany...

    Lepilemur microdon Small-toothed sportive lemur
    Small-toothed Sportive Lemur
    The small-toothed sportive lemur , or small-toothed weasel lemur, is a primate species in the family Lepilemuridae that—like all lemurs—is endemic to Madagascar. The species lives in dense rainforest in southeastern Madagascar, and can be found in Ranomafana and Andringitra National Parks...

    1894, Forsyth Major
    Charles Immanuel Forsyth Major
    Charles Immanuel Forsyth Major was a Swiss zoologist and vertebrate palaeontologist.Major was born in Glasgow and studied at Basel and Zurich Universities in Switzerland and later Göttingen in Germany...

    Lepilemur milanoii Daraina sportive lemur 2006, Louis, Jr.
    Lepilemur mittermeieri Mittermeier's sportive lemur
    Mittermeier's Sportive Lemur
    The Mittermeier's sportive lemur is a sportive lemur endemic to the Ampasindava Peninsula in Madagascar....

    2006, Rabarivola et al. unknown
    Lepilemur mustelinus Weasel sportive lemur 1851, I. Geoffroy
    Isidore Geoffroy Saint-Hilaire
    Isidore Geoffroy Saint-Hilaire was a French zoologist and an authority on deviation from normal structure. He coined the term ethology.He was born in Paris, the son of Étienne Geoffroy Saint-Hilaire...

    Lepilemur otto Otto's sportive lemur
    Otto's Sportive Lemur
    Otto's sportive lemur , or Ambodimahabibo sportive lemur, is a sportive lemur endemic to Madagascar. Its coloration is primarily grey-brown. It is known only from Ambodimahabibo, between the Mahajamba River in the west and the Sofia River in the north....

    2007, Craul et al. unknown
    Lepilemur petteri Petter's sportive lemur
    Petter's Sportive Lemur
    Petter's sportive lemur is a sportive lemur endemic to Madagascar. It is one of the smaller sportive lemurs with a total length of about 49 to 54 centimeters, of which 22-25 centimeters are tail. Petter's sportive lemur is found in southwestern Madagascar, living in dry spiny forests and some...

    2006, Louis, Jr.
    Lepilemur randrianasoloi Randrianasolo's sportive lemur 2006, Andriaholinirina et al.
    Lepilemur ruficaudatus Red-tailed sportive lemur
    Red-tailed Sportive Lemur
    The red-tailed sportive lemur , or red-tailed weasel lemur, is native to Madagascar like all lemurs. It is a nocturnal species feeding largely on leaves, though they also eat some fruit. Individuals weigh around 800 grams, and there is little sexual dimorphism...

    1867, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Lepilemur sahamalazensis Sahamalaza sportive lemur 2006, Andriaholinirina et al.
    Lepilemur scottorum Scott's sportive lemur
    Scott's Sportive Lemur
    Scott's sportive lemur , or Masoala sportive lemur, is a sportive lemur endemic to Madagascar. It has greyish brown fur and a black-tipped tail. It is named in honor of the Suzanne and Walter Scott Jr. Foundation....

    2008, Lei et al. unknown
    Lepilemur seali Seal's sportive lemur
    Seal's Sportive Lemur
    Seal's sportive lemur , or Anjanaharibe-Sud sportive lemur, is a sportive lemur endemic to Madagascar. It is a large sportive lemur with a total length of about 57 to 64 centimeters, of which 25 to 28 centimeters are tail. Seal's sportive lemur is found in northeastern Madagascar, living in...

    2006, Louis, Jr.
    Lepilemur septentrionalis Northern sportive lemur
    Northern Sportive Lemur
    The northern sportive lemur , also known as the Sahafary sportive lemur or northern weasel lemur, is a species of lemur in the Lepilemuridae family. It is endemic to Madagascar. It is threatened by habitat loss and has been named one of "The World's 25 Most Endangered Primates."-References:...

    1975, Rumpler & Albignac
    Lepilemur tymerlachsonorum Hawks' sportive lemur 2006, Louis, Jr.
    Lepilemur wrightae Wright's sportive lemur
    Wright's Sportive Lemur
    Wright's sportive lemur , or Kalambatritra sportive lemur, is a sportive lemur endemic to Madagascar. It is one of the larger sportive lemurs with a total length of about 52 to 64 centimeters, of which 24-27 centimeters are tail...

    2006, Louis, Jr.
    Lepilemur hollandorum Holland's sportive lemur
    Holland's Sportive Lemur
    Holland's sportive lemur , or Mananara-Nord sportive lemur, is a sportive lemur that is endemic to Madagascar. This lemur is found specifically in the Mananara-Nord Biosphere Reserve, but the limits of its habitat have yet to be determined. It lives in primary and secondary rainforests...

    2009, Ramaromilanto et al.


    Family: Indriidae

    Family Indriidae consists of the indri
    Indri
    The indri , also called the babakoto, is one of the largest living lemurs. It is a diurnal tree-dweller related to the sifakas and, like all lemuroids, it is native to Madagascar.- Etymology :...

     (the largest extant lemur), the woolly lemur
    Woolly lemur
    The woolly lemurs, also known as avahis or woolly indris, are nine species of strepsirrhine primates. Like all other lemurs, they live only on the island of Madagascar....

    s and the sifaka
    Sifaka
    Sifakas are a genus of lemur from the family Indriidae within the order Primates. Their name of the family is an onomatopoeia of their characteristic "shi-fak" alarm call. Like all lemurs, they are found only on the island of Madagascar...

    s. There are 19 extant species.
    Scientific name Common name Classified Average size Conservation status References
    Avahi betsileo Betsileo woolly lemur
    Betsileo Woolly Lemur
    The Betsileo woolly lemur or Betsileo avahi is a species of woolly lemur native to southeastern Madagascar, in the District of Fandriana. It weighs about 1 kg. The pelage differs significantly from other southeastern woolly lemurs in that it is primarily light reddish brown on most of the...

    2007, Andriantompohavana et al. unknown
    Avahi cleesei Cleese's woolly lemur
    Cleese's Woolly Lemur
    The Bemaraha woolly lemur , also known as Cleese's woolly lemur, is a species of woolly lemur native to western Madagascar, named after John Cleese...

    2005, Thalmann and Geissmann
    Avahi laniger Eastern woolly lemur
    Eastern Woolly Lemur
    The eastern woolly lemur , also known as the eastern avahi or Gmelin's woolly lemur, is a species of woolly lemur native to eastern Madagascar, where it lives in humid forests. This nocturnal animal weighs 1-1.3 kg and reaches a length of 27–29 cm with a tail of 33–37 cm...

    1788, Gmelin
    Johann Friedrich Gmelin
    Johann Friedrich Gmelin was a German naturalist, botanist, entomologist, herpetologist and malacologist.- Education :Johann Friedrich Gmelin was born as the eldest son of Philipp Friedrich Gmelin in 1748 in Tübingen...

    Avahi meridionalis Southern woolly lemur
    Southern Woolly Lemur
    The southern woolly lemur , or southern avahi, has been recently recognized as a separate species of woolly lemur in 2006 by Zaramody et al. It is a nocturnal and pair-living species...

    2006, Zaramody et al. unknown
    Avahi mooreorum Moore's woolly lemur
    Moore's Woolly Lemur
    Moore's woolly lemur , or Masoala woolly lemur, is a woolly lemur endemic to Madagascar. It has brown-grey fur and a reddish tail. It is named in honor of the Gordon and Betty Moore Foundation of San Francisco....

    2008, Lei et al. unknown
    Avahi occidentalis Western woolly lemur
    Western Woolly Lemur
    The western woolly lemur or western avahi is a species of woolly lemur native to western Madagascar, where they live in dry deciduous forests. These nocturnal animals weigh 0.7-0.9 kg. It is a folivorous species....

    1898, von Lorenz-Liburnau
    Avahi peyrierasi Peyrieras' woolly lemur
    Peyrieras' Woolly Lemur
    The Peyrieras' woolly lemur or Peyrieras' avahi is a species of woolly lemur native to southeastern Madagascar. It weighs about 1 kg....

    2006, Zaramody et al. unknown
    Avahi ramanantsoavani Ramanantsoavana's woolly lemur
    Ramanantsoavana's Woolly Lemur
    The Ramanantsoavana's woolly lemur , also known as Manombo woolly lemur or Ramanantsoavana's avahi, is a species of woolly lemur native to southeastern Madagascar. It weighs about 1 kg. It was originally considered a subspecies of Southern woolly lemur , A. m...

    2006, Zaramody et al. unknown
    Avahi unicolor Sambirano woolly lemur
    Sambirano Woolly Lemur
    The Sambirano woolly lemur , also known as the Sambirano avahi or Unicolor woolly lemur, is a species of woolly lemur native to western Madagascar....

    2000, Thalmann and Geissmann
    Indri indri Indri
    Indri
    The indri , also called the babakoto, is one of the largest living lemurs. It is a diurnal tree-dweller related to the sifakas and, like all lemuroids, it is native to Madagascar.- Etymology :...

    1788, Gmelin
    Johann Friedrich Gmelin
    Johann Friedrich Gmelin was a German naturalist, botanist, entomologist, herpetologist and malacologist.- Education :Johann Friedrich Gmelin was born as the eldest son of Philipp Friedrich Gmelin in 1748 in Tübingen...

    Propithecus candidus Silky sifaka
    Silky Sifaka
    The silky sifaka , or silky simpona, is a large lemur characterized by long, silky white fur. It has a very restricted range in northeastern Madagascar, where it is known locally as the simpona...

    1871, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Propithecus coquereli Coquerel's sifaka
    Coquerel's Sifaka
    Coquerel's sifaka is a medium-sized lemur of the sifaka genus Propithecus. Like all lemurs, it is endemic to Madagascar.-Description:...

    1867, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Propithecus coronatus Crowned sifaka
    Crowned Sifaka
    The crowned sifaka is a sifaka endemic to Madagascar. It has a length of 87 to 102 centimeters, of which 47-57 centimeters are tail. The crowned sifaka lives in western Madagascar. It lives in dry deciduous forest....

    1871, Milne-Edwards
    Henri Milne-Edwards
    Henri Milne-Edwards was an eminent French zoologist.Henri Milne-Edwards was the 27th child of William Edwards, an English planter and militia colonel in Jamaica and Elisabeth Vaux, a French. He was born in Bruges, Belgium, where his parents had retired. At that time, Bruges was a part of the...

    Propithecus deckenii Von der Decken's sifaka 1870, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Propithecus diadema Diademed sifaka
    Diademed Sifaka
    The diademed sifaka , or diademed simpona, is an endangered species of sifaka, one of the lemurs endemic to certain rainforests in eastern Madagascar. This species is one of the world's largest living lemurs, with a total adult length of approximately 105 centimetres , half of which is tail...

    1832, Bennett
    Edward Turner Bennett
    Edward Turner Bennett was an English zoologist and writer. He was the elder brother of the botanist John Joseph Bennett. Bennett was born at Hackney and practiced as a surgeon, but his chief pursuit was always zoology...

    Propithecus edwardsi Milne-Edwards' sifaka
    Milne-Edwards' sifaka
    Milne-Edwards' sifaka , or Milne-Edwards' simpona, is a large arboreal, diurnal lemur endemic to the eastern coastal rainforest of Madagascar. Milne-Edwards' sifaka is characterized by a black body with a light-colored "saddle" on the lower part of its back. It is closely related to the diademed...

    1871, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...

    Propithecus perrieri Perrier's sifaka
    Perrier's Sifaka
    Perrier's sifaka is a sifaka endemic to Madagascar. It has a length of 85 to 92 centimeters, of which 42-46 centimeters are tail. Perrier's sifaka has a very limited range in northeastern Madagascar between the Irodo River to the north and the Lokia River to the south. It lives in dry deciduous...

    1931, Lavauden
    Propithecus tattersalli Golden-crowned sifaka
    Golden-crowned Sifaka
    The golden-crowned sifaka or Tattersall's sifaka is a medium-sized lemur characterized by mostly white fur, prominent furry ears and a golden-orange crown. It is one of the smallest sifakas , weighing around and measuring approximately from head to tail...

    1988, Simons
    Propithecus verreauxi Verreaux's sifaka
    Verreaux's Sifaka
    Verreaux's sifaka , or white sifaka, is a medium sized primate in one of the lemur families, Indriidae. It lives in Madagascar and can be found in a variety of habitats from rainforest to western Madagascar dry deciduous forests and dry and spiny forests...

    1867, A. Grandidier
    Alfred Grandidier
    Alfred Grandidier was a French naturalist and explorer.From a very wealthy family, he and his brother, Ernest Grandidier , undertook a voyage around the world...



    Family: Daubentoniidae

    Family Daubentoniidae contains only one surviving species, of the aye-aye
    Aye-aye
    The aye-aye is a lemur, a strepsirrhine primate native to Madagascar that combines rodent-like teeth and a special thin middle finger to fill the same ecological niche as a woodpecker...

    . Wide ranging genetic studies have shown that it separated from the ancestral lemurs long before any other branch of the modern Lemuriformes.
    Scientific name Common name Classified Average size Conservation status References
    Daubentonia madagascariensis Aye-aye
    Aye-aye
    The aye-aye is a lemur, a strepsirrhine primate native to Madagascar that combines rodent-like teeth and a special thin middle finger to fill the same ecological niche as a woodpecker...

    1788, Gmelin
    Johann Friedrich Gmelin
    Johann Friedrich Gmelin was a German naturalist, botanist, entomologist, herpetologist and malacologist.- Education :Johann Friedrich Gmelin was born as the eldest son of Philipp Friedrich Gmelin in 1748 in Tübingen...

    2.5 kg (5.5 lb)

    Extinct species

    All known extinct lemurs from Madagascar are known from recent, subfossil
    Subfossil
    Subfossil refers to remains whose fossilization process is not complete, either for lack of time or because the conditions in which they were buried were not optimal for fossilization....

     remains. Conditions for fossil
    Fossil
    Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

    ization were not ideal on the island, so little is known about ancestral lemur populations. All known extinct lemurs are thought to have died out after the arrival of humans.

    Scientific name Common name Family Estimated size Extinction date References
    Archaeoindris fontoynonti
    Archaeoindris
    Archaeoindris fontoynonti is an extinct species of Malagasy lemur that was the largest primate to evolve on Madagascar. It weighed about and measured around 1.5m in height, more than a silverback gorilla. Archaeoindris is one of eight known members of the Palaeopropithecinae subfamily...

    Sloth lemurs Palaeopropithecidae 1600 CE
    Archaeolemur edwardsi
    Archaeolemur
    Archaeolemur is an extinct genus of lemurs that includes two species, Archaeolemur edwardsi and A. majori.The genus was widespread throughout Madagascar through much of the Holocene epoch, and its remains are often abundant at fossil sites across the island. The wide geographical and temporal range...

    Monkey lemur
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    s (or baboon lemurs)
    Archaeolemuridae
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    1047–1280 CE
    Archaeolemur majori
    Archaeolemur
    Archaeolemur is an extinct genus of lemurs that includes two species, Archaeolemur edwardsi and A. majori.The genus was widespread throughout Madagascar through much of the Holocene epoch, and its remains are often abundant at fossil sites across the island. The wide geographical and temporal range...

    Monkey lemur
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    s (or baboon lemurs)
    Archaeolemuridae
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    1047–1280 CE
    Babakotia radofilai
    Babakotia
    Babakotia is a medium-sized, extinct genus of lemur, or strepsirrhine primate, from Madagascar that contains a single species, Babakotia radofilai. Together with Palaeopropithecus, Archaeoindris, and Mesopropithecus, it forms the family Palaeopropithecidae, commonly known as the sloth lemurs...

    Sloth lemurs Palaeopropithecidae 3050 BCE
    Daubentonia robusta
    Giant Aye-aye
    The giant aye-aye is an extinct relative of the aye-aye, the only other species in the genus Daubentonia. It lived in Madagascar, appears to have disappeared less than 1,000 years ago, is entirely unknown in life, and is only known from subfossil remains.As of 2004, giant aye-aye remains consisted...

    Giant aye-aye
    Giant Aye-aye
    The giant aye-aye is an extinct relative of the aye-aye, the only other species in the genus Daubentonia. It lived in Madagascar, appears to have disappeared less than 1,000 years ago, is entirely unknown in life, and is only known from subfossil remains.As of 2004, giant aye-aye remains consisted...

    Daubentoniidae 891–1027 CE
    Hadropithecus stenognathus
    Hadropithecus
    Hadropithecus is a medium-sized, extinct genus of lemur, or strepsirrhine primate, from Madagascar that includes a single species, Hadropithecus stenognathus. Due to its rarity and lack of sufficient skeletal remains, it is one of the least understood of the extinct lemurs...

    Monkey lemur
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    s (or baboon lemurs)
    Archaeolemuridae
    Monkey lemur
    The Monkey lemurs or Baboon lemurs are an extinct type of lemurs that includes one family, Archaeolemuridae, two genera and three species. Despite their common names, members of Archaeolemuridae were not as closely related to monkeys as they were to other lemurs....

    444–772 CE
    Megaladapis edwardsi Koala lemurs Megaladapidae 1280–1420 CE
    Megaladapis grandidieri Koala lemurs Megaladapidae 1280–1420 CE
    Megaladapis madagascariensis Koala lemurs Megaladapidae 1280–1420 CE
    Mesopropithecus dolichobrachion
    Mesopropithecus
    Mesopropithecus is an extinct genus of small to medium-sized lemur, or strepsirrhine primate, from Madagascar that includes three species, M. dolichobrachion, M. globiceps, and M. pithecoides. Together with Palaeopropithecus, Archaeoindris, and Babakotia, it is part of the sloth...

    Sloth lemurs Palaeopropithecidae 245–429 CE
    Mesopropithecus globiceps
    Mesopropithecus
    Mesopropithecus is an extinct genus of small to medium-sized lemur, or strepsirrhine primate, from Madagascar that includes three species, M. dolichobrachion, M. globiceps, and M. pithecoides. Together with Palaeopropithecus, Archaeoindris, and Babakotia, it is part of the sloth...

    Sloth lemurs Palaeopropithecidae 245–429 CE
    Mesopropithecus pithecoides
    Mesopropithecus
    Mesopropithecus is an extinct genus of small to medium-sized lemur, or strepsirrhine primate, from Madagascar that includes three species, M. dolichobrachion, M. globiceps, and M. pithecoides. Together with Palaeopropithecus, Archaeoindris, and Babakotia, it is part of the sloth...

    Sloth lemurs Palaeopropithecidae 245–429 CE
    Pachylemur insignis
    Pachylemur
    Pachylemur is an extinct genus of lemur most closely related to the ruffed lemurs . Its two representative species, Pachylemur insignis and Pachylemur jullyi, are only known from subfossil remains found at sites in central and southwestern Madagascar...

    Pachylemur
    Pachylemur
    Pachylemur is an extinct genus of lemur most closely related to the ruffed lemurs . Its two representative species, Pachylemur insignis and Pachylemur jullyi, are only known from subfossil remains found at sites in central and southwestern Madagascar...

    Lemuridae
    Lemuridae
    Lemuridae is a family of prosimian primates native to Madagascar, and one of five families commonly known as lemurs. These animals were thought to be the evolutionary predecessors of monkeys and apes, but this is no longer considered correct...

    1280–1420 CE
    Pachylemur jullyi
    Pachylemur
    Pachylemur is an extinct genus of lemur most closely related to the ruffed lemurs . Its two representative species, Pachylemur insignis and Pachylemur jullyi, are only known from subfossil remains found at sites in central and southwestern Madagascar...

    Pachylemur
    Pachylemur
    Pachylemur is an extinct genus of lemur most closely related to the ruffed lemurs . Its two representative species, Pachylemur insignis and Pachylemur jullyi, are only known from subfossil remains found at sites in central and southwestern Madagascar...

    Lemuridae
    Lemuridae
    Lemuridae is a family of prosimian primates native to Madagascar, and one of five families commonly known as lemurs. These animals were thought to be the evolutionary predecessors of monkeys and apes, but this is no longer considered correct...

    1280–1420 CE
    Palaeopropithecus ingens Sloth lemurs Palaeopropithecidae 1300–1620 CE
    Palaeopropithecus kelyus Sloth lemurs Palaeopropithecidae unknown
    Palaeopropithecus maximus Sloth lemurs Palaeopropithecidae 1300–1620 CE


    Footnotes

    In 2008, the red lemur, Eulemur rufus, was split into two species, Eulemur rufus (red lemur) and Eulemur rufifrons (red-fronted lemur). Also, Eulemur rufus was previously known as the red-fronted lemur, but was renamed the red lemur, while Eulemur rufifrons assumed its former name.Formerly referred to as Eulemur albocollaris or white-collared brown lemur, but was changed in 2008.This extinction date for Babakotia radofilai is based on a single radiocarbon date
    Radiocarbon dating
    Radiocarbon dating is a radiometric dating method that uses the naturally occurring radioisotope carbon-14 to estimate the age of carbon-bearing materials up to about 58,000 to 62,000 years. Raw, i.e. uncalibrated, radiocarbon ages are usually reported in radiocarbon years "Before Present" ,...

     from one specimen. For this reason, it is hard to tell when this species became extinct or how it is related to other lemur species.Average weights reported for this species are based on very small sample sizes or are general ranges for its genus and thus require further research.

    External links


    See also

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