German wasp
Encyclopedia
The German wasp, or European wasp, Vespula germanica, is a wasp
Wasp
The term wasp is typically defined as any insect of the order Hymenoptera and suborder Apocrita that is neither a bee nor an ant. Almost every pest insect species has at least one wasp species that preys upon it or parasitizes it, making wasps critically important in natural control of their...

 found in much of the Northern Hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of its equator—the word hemisphere literally means “half sphere”. It is also that half of the celestial sphere north of the celestial equator...

, native to Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, northern Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

, and temperate Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

. It has been introduced and is well-established in many other places, including North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

, South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

 (Argentina
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

 and Chile
Chile
Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

), Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

 and New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

. German wasps are part of the family Vespidae and are sometimes mistakenly referred to as paper wasp
Paper wasp
Paper wasps are -long wasps that gather fibers from dead wood and plant stems, which they mix with saliva, and use to construct water-resistant nests made of gray or brown papery material...

s because they build a grey paper nest—although, strictly speaking, paper wasps are part of the subfamily Polistinae
Polistinae
The Polistinae are eusocial wasps closely related to the more familiar yellowjackets, but placed in their own subfamily, containing four tribes; with some 1100 species total, it is the second most diverse subfamily within Vespidae, and while most species are tropical or subtropical, they include...

. In North America, they are also known as yellowjacket
Yellowjacket
Yellowjacket is the common name in North America for predatory wasps of the genera Vespula and Dolichovespula. Members of these genera are known simply as "wasps" in other English-speaking countries...

s.

Identification

The German wasp is about 13mm (0.5 inch) long, and has typical wasp colours of black and yellow. It is very similar to the common wasp (Vespula vulgaris), but seen head on, its face has three tiny black dots. German wasps also have black dots on their abdomen, while the common wasp's analogous markings are fused with the black rings above them, forming a different pattern.

Nests

The nest is made from chewed plant fibres, mixed with saliva
Saliva
Saliva , referred to in various contexts as spit, spittle, drivel, drool, or slobber, is the watery substance produced in the mouths of humans and most other animals. Saliva is a component of oral fluid. In mammals, saliva is produced in and secreted from the three pairs of major salivary glands,...

. They are generally found close to or in the ground, rather than higher up on bushes and trees like hornet
Hornet
Hornets are the largest eusocial wasps; some species can reach up to in length. The true hornets make up the genus Vespa and are distinguished from other vespines by the width of the vertex , which is proportionally larger in Vespa and by the anteriorly rounded gasters .- Life cycle :In...

s. It has open cells and a petiole
Petiole (insect)
In entomology, the term petiole is most commonly used to refer to the constricted first metasomal segment of members of the Hymenopteran suborder Apocrita; it may be used to refer to other insects with similar body shapes, where the metasomal base is constricted...

 attaching the nest to the substrate. The wasps produce a chemical which repels ant
Ant
Ants are social insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. More than...

s, and secrete it around the base of this petiole in order to avoid ant predation
Predation
In ecology, predation describes a biological interaction where a predator feeds on its prey . Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of its prey and the eventual absorption of the prey's tissue through consumption...

.

A solitary female queen
Eusociality
Eusociality is a term used for the highest level of social organization in a hierarchical classification....

 starts the nest, building 20–30 cells before initial egg-laying. This phase begins in spring, depending on climatic conditions. She fashions a petiole and produces a single cell at the end of it. Six further cells are then added around this to produce the characteristic hexagonal shape of the nest cells.

Once the larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e have hatched as workers
Eusociality
Eusociality is a term used for the highest level of social organization in a hierarchical classification....

, they take up most of the colony’s foraging, brood care and nest maintenance. A finished nest may be 20–30 cm across and contain 3,000 individuals.

Each wasp colony
Colony (biology)
In biology, a colony reference to several individual organisms of the same species living closely together, usually for mutual benefit, such as stronger defense or the ability to attack bigger prey. Some insects live only in colonies...

 includes one queen and a number of sterile workers. Colonies usually last only one year, all but the queen dying at the onset of winter. However, in mild climates such as New Zealand, around 10% of the colonies survive the winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queen overwinters in a crack or other sheltered location.

This common and widespread wasp collects insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s including caterpillar
Caterpillar
Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...

s to feed to its larvae, and is therefore generally beneficial. The adults feed on nectar and sweet fruit, and are also attracted to human food and food waste, particularly sodas and meats.

The nests are subject to predation by the Honey Buzzard
Honey Buzzard
The European Honey Buzzard , is a bird of prey in the family Accipitridae which also includes many other diurnal raptors such as kites, eagles and harriers....

, which excavates them to obtain the larva. The hoverfly
Hoverfly
Hoverflies, sometimes called flower flies or syrphid flies, make up the insect family Syrphidae. As their common name suggests, they are often seen hovering or nectaring at flowers; the adults of many species feed mainly on nectar and pollen, while the larvae eat a wide range of foods...

 Volucella pellucens
Volucella pellucens
Volucella pellucens is a hover-fly. It occurs in much of Europe, and across Asia to Japan.It is about 15–16 mm in length with a broad body. It is mainly black, but the front part of the abdomen has a broad yellow band, giving it the appearance of a bee or wasp...

and some of its relatives lay their eggs in the wasp's nest, and the larva feeds on the wasp's young.
This species is considered a pest in most areas outside its native range, though its long residency in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 is such that it is not treated with any level of urgency there, in contrast to areas such as South America, where the introduction is more recent, and the impacts far more dramatic, prompting a greater degree of concern over control measures (e.g.).

Along with the closely related common wasp
Common wasp
The common wasp, Vespula vulgaris, known in the US as the yellowjacket, is found in much of the Northern Hemisphere and has been introduced to Australia and New Zealand. It is a eusocial vespid which builds its grey paper nest in or on a structure capable of supporting it...

 and two species of Polistes
Polistes
Wasps of the cosmopolitan genus Polistes are the most familiar of the polistine wasps, and are the most common type of paper wasp. It is also the single largest genus within the family Vespidae, with over 300 recognized species and subspecies...

, the German wasp is likewise considered to be a pest in New Zealand. It was probably introduced in the late 19th century, but did not appear in large numbers until around 1940. It is common in the beech
Nothofagus
Nothofagus, also known as the southern beeches, is a genus of 35 species of trees and shrubs native to the temperate oceanic to tropical Southern Hemisphere in southern South America and Australasia...

 forest since it is one of the two wasps that feeds on the honeydew
Honeydew (secretion)
Honeydew is a sugar-rich sticky liquid, secreted by aphids and some scale insects as they feed on plant sap. When their mouthpart penetrates the phloem, the sugary, high-pressure liquid is forced out of the gut's terminal opening. Honeydew is particularly common as a secretion in the Hemipteran...

 exuded by the native beech scale insect which lives in the bark of the trees. It has a serious effect on the forest ecology
Ecology
Ecology is the scientific study of the relations that living organisms have with respect to each other and their natural environment. Variables of interest to ecologists include the composition, distribution, amount , number, and changing states of organisms within and among ecosystems...

since there is less honeydew available for the native birds. In domestic situations nests have been known to become very large, sometimes taking up entire attic spaces in houses. This is put down to the comparatively mild winters experienced in New Zealand, as opposed to the wasp's usual European habitat.

An unusual attempt at wasp control is related from Abercairney in Scotland, where until the 1950s children were encouraged to compete in the Wasp Cup, awarded to the competitor who handed in the most queen wasps. The wasps were stuck to card and a payment of 1d was made for each; totals of forty were not uncommon.

External links

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