Trimorphodon
Encyclopedia
Trimorphodon is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of mildly venomous, rear-fanged, colubrid
Colubrid
A colubrid is a member of the snake family Colubridae. This broad classification of snakes includes about two-thirds of all snake species on earth. The earliest species of the snake family date back to the Oligocene epoch. With 304 genera and 1,938 species, Colubridae is the largest snake family...

 snake
Snake
Snakes are elongate, legless, carnivorous reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears. Like all squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales...

s. They are commonly known as lyre snakes, named after the distinctive V shaped pattern on their head that is said to resemble the shape of a lyre
Lyre
The lyre is a stringed musical instrument known for its use in Greek classical antiquity and later. The word comes from the Greek "λύρα" and the earliest reference to the word is the Mycenaean Greek ru-ra-ta-e, meaning "lyrists", written in Linear B syllabic script...

. In Mexico, they are commonly called "víbora de uña," or "nail viper." The word Trimorphodon is a combination of three Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...

 words, 'tri' - three, 'morph' - shape, and 'odon' - teeth, which refers to the three distinct kinds of teeth that lyre snakes have: recurved anterior teeth; shorter middle teeth, and large grooved fangs
Canine tooth
In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, fangs, or eye teeth, are relatively long, pointed teeth...

 at the rear of the jaw. There are two distinct species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 in the genus Trimorphodon, with seven subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

.

Range

Trimorphodon biscutatus ranges throughout the southwestern United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

, from Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

 to California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 as well as into northern Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

. Trimorphodon tau ranges through most of Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

, and down into Central America
Central America
Central America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...

, in Guatemala
Guatemala
Guatemala is a country in Central America bordered by Mexico to the north and west, the Pacific Ocean to the southwest, Belize to the northeast, the Caribbean to the east, and Honduras and El Salvador to the southeast...

, El Salvador
El Salvador
El Salvador or simply Salvador is the smallest and the most densely populated country in Central America. The country's capital city and largest city is San Salvador; Santa Ana and San Miguel are also important cultural and commercial centers in the country and in all of Central America...

, Honduras
Honduras
Honduras is a republic in Central America. It was previously known as Spanish Honduras to differentiate it from British Honduras, which became the modern-day state of Belize...

, Nicaragua
Nicaragua
Nicaragua is the largest country in the Central American American isthmus, bordered by Honduras to the north and Costa Rica to the south. The country is situated between 11 and 14 degrees north of the Equator in the Northern Hemisphere, which places it entirely within the tropics. The Pacific Ocean...

, and as far south as Costa Rica
Costa Rica
Costa Rica , officially the Republic of Costa Rica is a multilingual, multiethnic and multicultural country in Central America, bordered by Nicaragua to the north, Panama to the southeast, the Pacific Ocean to the west and the Caribbean Sea to the east....

. They are primarily found in rocky crevices and outcroppings.

Description

Lyre snakes usually grow to about 1m at adult size. They have a broad head with a narrow neck and large eye
Eye
Eyes are organs that detect light and convert it into electro-chemical impulses in neurons. The simplest photoreceptors in conscious vision connect light to movement...

s with vertical pupil
Pupil
The pupil is a hole located in the center of the iris of the eye that allows light to enter the retina. It appears black because most of the light entering the pupil is absorbed by the tissues inside the eye. In humans the pupil is round, but other species, such as some cats, have slit pupils. In...

s. They are tan
Tan (color)
Tan is a pale whiteish, tawny shade of white. The name is derived from tannum used in the tanning of leather.The first recorded use of tan as a shade name in English was in the year 1590....

 or brown
Brown
Brown is a color term, denoting a range of composite colors produced by a mixture of orange, red, rose, or yellow with black or gray. The term is from Old English brún, in origin for any dusky or dark shade of color....

 in color, with distinctive black
Black
Black is the color of objects that do not emit or reflect light in any part of the visible spectrum; they absorb all such frequencies of light...

 or dark brown bands that have lighter colored borders.

Behavior

Trimorphodon species are primarily nocturnal, and feed on lizard
Lizard
Lizards are a widespread group of squamate reptiles, with nearly 3800 species, ranging across all continents except Antarctica as well as most oceanic island chains...

s, rodent
Rodent
Rodentia is an order of mammals also known as rodents, characterised by two continuously growing incisors in the upper and lower jaws which must be kept short by gnawing....

s and even bat
Bat
Bats are mammals of the order Chiroptera "hand" and pteron "wing") whose forelimbs form webbed wings, making them the only mammals naturally capable of true and sustained flight. By contrast, other mammals said to fly, such as flying squirrels, gliding possums, and colugos, glide rather than fly,...

s. They are a very secretive species, and excellent climbers. They live inside rock crevices and canyon walls where humans cannot easily find them, which makes research difficult.

Reproduction

They are oviparous, laying clutches of 10 or more egg
Egg (biology)
An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

s. Hybridization between species is not unknown in the areas where their range overlaps.

Species

  • Western Lyre Snake, Trimorphodon biscutatus
    Trimorphodon biscutatus
    The Western Lyre Snake is a mildly venomous colubrid snake native to the western United States, northern Mexico, and Central America.- Description :...

    • Trimorphodon biscutatus biscutatus (Duméril
      André Marie Constant Duméril
      André Marie Constant Duméril was a French zoologist. He was professor of anatomy at the Muséum National d'Histoire Naturelle from 1801 to 1812, when he became professor of herpetology and ichthyology...

      , Bibron
      Gabriel Bibron
      Gabriel Bibron was a French zoologist. He was born in Paris. Son of an employee of the Museum national d'histoire naturelle, he had a good foundation in natural history and was hired to collect vertebrates in Italy and Sicily. He classified a number of reptile species with André Marie Constant...

       & Duméril
      Auguste Duméril
      Auguste Henri André Duméril was a French zoologist. He was professor of Herpetology and Ichthyology at the Muséum National d'Histoire Naturelle in Paris from 1857. His father André Marie Constant Duméril was also a zoologist...

      , 1854)
    • Sonora
      Sonora
      Sonora officially Estado Libre y Soberano de Sonora is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided into 72 municipalities; the capital city is Hermosillo....

      n Lyre Snake, Trimorphodon biscutatus lambda (Cope
      Edward Drinker Cope
      Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

      , 1886)
    • Baja
      Baja California Peninsula
      The Baja California peninsula , is a peninsula in northwestern Mexico. Its land mass separates the Pacific Ocean from the Gulf of California. The Peninsula extends from Mexicali, Baja California in the north to Cabo San Lucas, Baja California Sur in the south.The total area of the Baja California...

       California
      California
      California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

       Lyre Snake, Trimorphodon biscutatus lyrophanes (Cope
      Edward Drinker Cope
      Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

      , 1860)
    • Trimorphodon biscutatus quadruplex (Smith
      Hobart M. Smith
      Hobart Muir Smith, born Frederick William Stouffer , is an American herpetologist. He was born in Stanwood, Iowa, United States. He has been credited with describing several new species of North American reptiles and amphibians...

      , 1941)
    • Texas
      Texas
      Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

       Lyre Snake, Trimorphodon biscutatus vilkinsonii
      Trimorphodon biscutatus vilkinsonii
      The Texas lyre snake is a subspecies of mildly venomous rear-fanged colubrid. The epithet vilkinsonii is in honor of amateur American naturalist Edward Wilkinson, who collected the first specimen near the city of Chihuahua...

      (Cope
      Edward Drinker Cope
      Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

      , 1886)
  • Mexican
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

     Lyre Snake, Trimorphodon tau
    • Trimorphodon tau latifascia (Peters
      Wilhelm Peters
      Wilhelm Karl Hartwich Peters was a German naturalist and explorer.He was assistant to Johannes Peter Müller and later curator of the Berlin Zoological Museum. In September 1842 he travelled to Mozambique via Angola. He returned to Berlin with an enormous collection of natural history specimens...

      , 1869)
    • Trimorphodon tau tau (Cope
      Edward Drinker Cope
      Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

      , 1870)
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