Small Copper
Encyclopedia
The Small Copper, American Copper, or the Common Copper (Lycaena phlaeas) is a butterfly
Butterfly
A butterfly is a mainly day-flying insect of the order Lepidoptera, which includes the butterflies and moths. Like other holometabolous insects, the butterfly's life cycle consists of four parts: egg, larva, pupa and adult. Most species are diurnal. Butterflies have large, often brightly coloured...

 of the Lycaenids or gossamer-winged
Lycaenidae
The Lycaenidae are the second-largest family of butterflies, with about 6000 species worldwide, whose members are also called gossamer-winged butterflies...

 butterfly family.

Description

The upperside forewings are a bright orange with a dark outside edge border and with eight or nine black spots. The hindwings are dark with an orange border. Some females also have a row of blue spots inside the orange border and are known as form caeruleopunctata. The undersides are patterned in a similar way but are paler. The black spots on the forewings are outlined in yellow and the dark colouring is replaced by a pale brownish, gray. The hindwings are the same brown/grey colour with small black dots and a narrow orange border. The caterpillar
Caterpillar
Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...

s (larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e) are usually green, but some have a purple stripe down the middle of the back and along each side.

Range

It is widespread and common across Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

, and North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

, and also found in North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

 south through to Ethiopia
Ethiopia
Ethiopia , officially known as the Federal Democratic Republic of Ethiopia, is a country located in the Horn of Africa. It is the second-most populous nation in Africa, with over 82 million inhabitants, and the tenth-largest by area, occupying 1,100,000 km2...

.

It can be found almost anywhere in south/central England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

 and Wales
Wales
Wales is a country that is part of the United Kingdom and the island of Great Britain, bordered by England to its east and the Atlantic Ocean and Irish Sea to its west. It has a population of three million, and a total area of 20,779 km²...

 although never, it seems, in large numbers. Its distribution becomes more patchy in northern England, Scotland
Scotland
Scotland is a country that is part of the United Kingdom. Occupying the northern third of the island of Great Britain, it shares a border with England to the south and is bounded by the North Sea to the east, the Atlantic Ocean to the north and west, and the North Channel and Irish Sea to the...

 and Ireland
Ireland
Ireland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...

.

Habitat

It is found in a wide variety range of habitats from chalk downlands, heathland, woodland
Woodland
Ecologically, a woodland is a low-density forest forming open habitats with plenty of sunlight and limited shade. Woodlands may support an understory of shrubs and herbaceous plants including grasses. Woodland may form a transition to shrubland under drier conditions or during early stages of...

 clearings to churchyard
Churchyard
A churchyard is a patch of land adjoining or surrounding a church which is usually owned by the relevant church or local parish itself. In the Scots language or Northern English language this can also be known as a kirkyard or kirkyaird....

s and waste ground in cities.

Habits

In bright sun it is a very active little butterfly with the males setting up small territories which they will defend vigorously against rival males or indeed any unlucky passing insect. Even the shadow of a large bird passing overhead is enough to get him going. Females are pursued mercilessly and mating usually occurs low down in vegetation.

Life cycle

The eggs are laid singly and conspicuously on the upperside of foodplant leaves and the young caterpillar feeds on the underside of the leaf creating "windows" by leaving the upper epidermis of the leaf untouched. Pupation takes place in the leaf litter and the pupa
Pupa
A pupa is the life stage of some insects undergoing transformation. The pupal stage is found only in holometabolous insects, those that undergo a complete metamorphosis, going through four life stages; embryo, larva, pupa and imago...

 is thought to be tended by ants. There are between two and three broods a year, fewer further north. In exceptionally good years, a fourth brood sometimes occurs in the south and adults can still be seen flying into November. The species overwinters as a caterpillar.

Host-plants

Depending on the habitat, Common Sorrel (Rumex acetosa) and Sheep's Sorrel (Rumex acetosella) are the two main food-plants, although other Docks
Rumex
The docks and sorrels, genus Rumex L., are a genus of about 200 species of annual, biennial and perennial herbs in the buckwheat family Polygonaceae....

 (Rumex spp.) are occasionally used.

Systematics

Lycaena phlaeas belongs to the subgenus Thersamolycaena of the genus Lycaena. The species is in turn divided into several subspecies:
  • Lycaena phlaeas hibernicaRepublic of Ireland
    Republic of Ireland
    Ireland , described as the Republic of Ireland , is a sovereign state in Europe occupying approximately five-sixths of the island of the same name. Its capital is Dublin. Ireland, which had a population of 4.58 million in 2011, is a constitutional republic governed as a parliamentary democracy,...

    , Northern Ireland
    Northern Ireland
    Northern Ireland is one of the four countries of the United Kingdom. Situated in the north-east of the island of Ireland, it shares a border with the Republic of Ireland to the south and west...

  • Lycaena phlaeas eleus — England, Wales, Scotland
  • Lycaena phlaeas phlaeas — Europe, western Siberia
    Siberia
    Siberia is an extensive region constituting almost all of Northern Asia. Comprising the central and eastern portion of the Russian Federation, it was part of the Soviet Union from its beginning, as its predecessor states, the Tsardom of Russia and the Russian Empire, conquered it during the 16th...

    , Caucasus
    Caucasus
    The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

    , South Caucasus
    South Caucasus
    The South Caucasus is a geopolitical region located on the border of Eastern Europe and Southwest Asia also referred to as Transcaucasia, or The Trans-Caucasus...

     (type locality = Sweden
    Sweden
    Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

    )
  • Lycaena phlaeas polaris — northern Ural Mountains
    Ural Mountains
    The Ural Mountains , or simply the Urals, are a mountain range that runs approximately from north to south through western Russia, from the coast of the Arctic Ocean to the Ural River and northwestern Kazakhstan. Their eastern side is usually considered the natural boundary between Europe and Asia...

    , northern Siberia, Chukotka Peninsula (Russia
    Russia
    Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

    )
  • Lycaena phlaeas kamtschaticaKamchatka Peninsula
    Kamchatka Peninsula
    The Kamchatka Peninsula is a peninsula in the Russian Far East, with an area of . It lies between the Pacific Ocean to the east and the Sea of Okhotsk to the west...

     (Russia)
  • Lycaena phlaeas ganalica — Kamchatka Peninsula (Russia)
  • Lycaena phlaeas daimio — south Kurile Islands, Sakhalin
    Sakhalin
    Sakhalin or Saghalien, is a large island in the North Pacific, lying between 45°50' and 54°24' N.It is part of Russia, and is Russia's largest island, and is administered as part of Sakhalin Oblast...

     (Russian islands)
  • Lycaena phlaeas hypophlaeas — Altai Mountains, southern Siberia, Amur (Russia–NW China
    China
    Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

     border region), northern Ussuri (adjacent to Amur)
  • Lycaena phlaeas chinensis — southern Ussuri
  • Lycaena phlaeas oxianaKopet Dag mountains
    Kopet Dag
    The Kopet Dag, Kopet Dagh, or Koppeh Dagh , also known as the Turkmen-Khorasan Mountain Range is a mountain range on the frontier between Turkmenistan and Iran, extending about 650 km along the border, east of the Caspian Sea. The highest peak of the range in Turkmenistan is southwest of the...

    , Alai Mountains, Ghissar-Darvaz, Tian Shan mountains
    Tian Shan
    The Tian Shan , also spelled Tien Shan, is a large mountain system located in Central Asia. The highest peak in the Tian Shan is Victory Peak , ....

  • Lycaena phlaeas comedarum — eastern Pamir Mountains
    Pamir Mountains
    The Pamir Mountains are a mountain range in Central Asia formed by the junction or knot of the Himalayas, Tian Shan, Karakoram, Kunlun, and Hindu Kush ranges. They are among the world’s highest mountains and since Victorian times they have been known as the "Roof of the World" a probable...

  • Lycaena phlaeas stygiana — western Pamir Mountains
  • Lycaena phlaeas shima — Arabia
  • Lycanea phlaeas pseudophlaeasEthiopia
    Ethiopia
    Ethiopia , officially known as the Federal Democratic Republic of Ethiopia, is a country located in the Horn of Africa. It is the second-most populous nation in Africa, with over 82 million inhabitants, and the tenth-largest by area, occupying 1,100,000 km2...

    , Uganda
    Uganda
    Uganda , officially the Republic of Uganda, is a landlocked country in East Africa. Uganda is also known as the "Pearl of Africa". It is bordered on the east by Kenya, on the north by South Sudan, on the west by the Democratic Republic of the Congo, on the southwest by Rwanda, and on the south by...

  • Lycaena phlaeas ethiopica — Ruwenzori Mountains (SW Uganda)
  • Lycaena phlaeas abbottiKenya
    Kenya
    Kenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...

    , Tanzania
    Tanzania
    The United Republic of Tanzania is a country in East Africa bordered by Kenya and Uganda to the north, Rwanda, Burundi, and the Democratic Republic of the Congo to the west, and Zambia, Malawi, and Mozambique to the south. The country's eastern borders lie on the Indian Ocean.Tanzania is a state...

    , Malawi
    Malawi
    The Republic of Malawi is a landlocked country in southeast Africa that was formerly known as Nyasaland. It is bordered by Zambia to the northwest, Tanzania to the northeast, and Mozambique on the east, south and west. The country is separated from Tanzania and Mozambique by Lake Malawi. Its size...

  • Lycaena phlaeas flavens
  • Lycaena phlaeas coccinea
  • Lycaena phlaeas americana
  • Lycaena phlaeas arethusa
  • Lycaena phlaeas arctodon
  • Lycaena phlaeas feildeni

See also


External links

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