Antillocladius
Encyclopedia
Antillocladius is a genus of midges belonging to the family Chironomidae
Chironomidae
Chironomidae are a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae...

, subfamily Orthocladiinae
Orthocladiinae
Orthocladiinae is a subfamily of midges in the non-biting midge family . For lack of a better common name, they are simply referred to as orthoclads.-Genera:*Abiskomyia Edwards, 1937*Acamptocladius Brundin, 1956...

. The name Antillocladius means orthoclad (subfamily Orthocladiinae) that occurs in the Antilles
Antilles
The Antilles islands form the greater part of the West Indies in the Caribbean Sea. The Antilles are divided into two major groups: the "Greater Antilles" to the north and west, including the larger islands of Cuba, Jamaica, Hispaniola , and Puerto Rico; and the smaller "Lesser Antilles" on the...

, where it was primarily recorded.

Distribution

The genus Antillocladius is distributed in the Nearctic
Nearctic
The Nearctic is one of the eight terrestrial ecozones dividing the Earth's land surface.The Nearctic ecozone covers most of North America, including Greenland and the highlands of Mexico...

, Neotropical, Palaearctic (East Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

) and Indomalaya
Indomalaya
The Indomalaya ecozone is one of the eight ecozones that cover the planet's land surface. It extends across most of South and Southeast Asia and into the southern parts of East Asia....

n ecozone
Ecozone
An ecozone is the broadest biogeographic division of the Earth's land surface, based on distributional patterns of terrestrial organisms.Ecozones delineate large areas of the Earth's surface within which organisms have been evolving in relative isolation over long periods of time, separated from...

s. The genus was erected by Sæther (1981) based on A. antecalvus Sæther, 1981, from the British West Indies
British West Indies
The British West Indies was a term used to describe the islands in and around the Caribbean that were part of the British Empire The term was sometimes used to include British Honduras and British Guiana, even though these territories are not geographically part of the Caribbean...

: Saint Vincent
Saint Vincent (island)
Saint Vincent is a volcanic island in the Caribbean. It is the largest island of the chain called Saint Vincent and the Grenadines. It is located in the Caribbean Sea, between Saint Lucia and Grenada. It is composed of partially submerged volcanic mountains...

 and Saint Lucia
Saint Lucia
Saint Lucia is an island country in the eastern Caribbean Sea on the boundary with the Atlantic Ocean. Part of the Lesser Antilles, it is located north/northeast of the island of Saint Vincent, northwest of Barbados and south of Martinique. It covers a land area of 620 km2 and has an...

. Subsequently, Sæther (1982) described two new species from the southeastern United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 (Antillocladius arcuatus and Antillocladius pluspilalus) and emended the genus. Wang and Sæther (1993) described two additional new species from Palaearctic and Oriental China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

. Andersen and Contreras-Ramos (1999) described the first species from Continental South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

 (Ecuador
Ecuador
Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

). The genus was revised by Mendes, Andersen & Sæther (2004), with the addition of nine species, plus two Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

ese species transferred to the genus (Yamamoto, 2004). Recently the genus was revised by Mendes & Andersen (2008) with the addition of 7 new species from Brazil and a new combination of a species from India. The genus thus comprises:

Species

  • A. anandae Mendes, Andersen & Hagenlund, 2011 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. antecalvus Sæther, 1981 - recorded from Saint Vincent
    Saint Vincent (island)
    Saint Vincent is a volcanic island in the Caribbean. It is the largest island of the chain called Saint Vincent and the Grenadines. It is located in the Caribbean Sea, between Saint Lucia and Grenada. It is composed of partially submerged volcanic mountains...

    , Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

     and Venezuela
    Venezuela
    Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

  • A. arcuatus Sæther, 1982 - recorded from USA, Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

    , Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

     and Venezuela
    Venezuela
    Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

  • A. atalaia Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. axitiosus Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. biota Mendes, Andersen & Sæther, 2004 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. brazuca Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. calakmulensis Mendes, Andersen & Sæther, 2004 - recorded from Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • A. campususp Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. folius Mendes, Andersen & Sæther, 2004 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. gephyrus Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. herradurus Mendes, Andersen & Sæther, 2004 - recorded from Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • A. itatiaia Mendes, Andersen & Hagenlund, 2011 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. musci Mendes, Andersen & Sæther, 2004 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. plicatus Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. pluspilalus Sæther, 1982 - recorded from USA, Ecuador
    Ecuador
    Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

    , Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

     and Nicaragua
    Nicaragua
    Nicaragua is the largest country in the Central American American isthmus, bordered by Honduras to the north and Costa Rica to the south. The country is situated between 11 and 14 degrees north of the Equator in the Northern Hemisphere, which places it entirely within the tropics. The Pacific Ocean...

  • A. scalpellatus Wang & Sæther, 1993 - recorded from China
    China
    Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

     and Russia
    Russia
    Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

  • A. skartveiti Andersen & Contreras-Ramos, 1999 - recorded from Ecuador
    Ecuador
    Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

  • A. sooretama Mendes, Andersen & Sæther, 2004 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. subnubilus (Sinharay & Chaudhuri, 1979) - recorded from India
    India
    India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

  • A. tokarameneus (Sasa & Suzuki, 1995) - recorded from Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • A. ubatuba Mendes, Andersen & Sæther, 2004 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. ultimus Mendes & Andersen, 2008 - recorded from Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

  • A. venequatoriensis Mendes, Andersen & Sæther, 2004 - recorded from Venezuela
    Venezuela
    Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

     and Ecuador
    Ecuador
    Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

  • A. yakyijeus (Sasa & Suzuki, 2000) - recorded from Japan
    Japan
    Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  • A. zempoalensis Mendes, Andersen & Sæther, 2004 - recorded from Mexico
    Mexico
    The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

  • A. zhengi Wang & Sæther, 1993 - recorded from China
    China
    Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

     and Thailand
    Thailand
    Thailand , officially the Kingdom of Thailand , formerly known as Siam , is a country located at the centre of the Indochina peninsula and Southeast Asia. It is bordered to the north by Burma and Laos, to the east by Laos and Cambodia, to the south by the Gulf of Thailand and Malaysia, and to the...


Characteristics

The species are based on male genitalia characters and only two females are described, A. musci and A. antecalvus. The pupa
Pupa
A pupa is the life stage of some insects undergoing transformation. The pupal stage is found only in holometabolous insects, those that undergo a complete metamorphosis, going through four life stages; embryo, larva, pupa and imago...

e are known of only four species, A. antecalvus, A. folius, A. musci and A. pluspilalus. The larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e are known of only three species, A. folius, A. musci and A. pluspilalus.

The immatures of the genus are described from terrestrial and semi-terrestrial habitats, sharing the environment with many other genera in the subfamily Orthocladiinae.
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