Annonaceae
Encyclopedia
Annonaceae, also called the custard apple family
is a family of flowering plant
s consisting of tree
s, shrub
s or rarely liana
s.
With about 2300 to 2500 species and more than 130 genera,
it is the largest family in Magnoliales
. Only five genera, Annona
, Rollinia
, Uvaria
, Melodorum
and Asimina produce edible fruits.
Its type genus
is Annona
. The family is concentrated in the tropics
, with few species found in temperate
regions. About 900 species are Neotropic
al, 450 are Afrotropic
al, and the other species Indomalaya
n.
Compared to the species from the Neotropic
s, very little is known about many species from Indomalaya
. Only a few attempts have been made for the phylogeny-based reclassification
of the family, and those have been hampered by the Neotropic
bias in the available information, with the most of the work having been done on genera and tribes.
or evergreen
trees and shrubs, some lianas, with aromatic bark, leaves, and flowers.
Stems, stalks and leaves: Bark is fibrous and aromatic. Pith
septate (fine tangential bands divided by partitions) to diaphragmed (divided by thin partitions with openings in them).
Flowers: Flower stalks
are axillary to (on the opposite side of shoot from) leaf scars on old wood and sometimes from leaves on new shoots. The flowers are usually trimerous
; borne singly or in compound inflorescence
s; bisexual
and rarely unisexual
. The receptacle might become enlarged, elevated or flat. The outer whorls
are inserted below the ovaries, and have valvate (overlapping) or imbricate (nonoverlapping) segments. Usually two to four persistent sepal
s that are distinct or connate (fused) at the base. Six petals in two unequal whorls of three with larger outer whorls and fleshier inner whorls that might share the same nectar glands, or six to fifteen petals, with impressed veins on their inner face. Ten to twenty (or many more) stamens inserted below the ovary, spirally arranged and forming a ball or flat-topped mass with short and stout filaments and linear to oblong anthers which face outward and open longitudinally. Each flower can have from one to many pistils, distinct to connate, with stigmas distinct. Marginal placentation, each pistil bearing one locule
, with one to many ovules. Style short and thick, with terminal stigma.
Fruits and seeds: Fruits are distinct (berries) or coalesce (into syncarps). Seeds are one to many per pistil; have a fleshy and usually brightly colored cover, have ruminate endosperm (nutritive tissue surrounding the embryo) and are oily.
and inter-familial systematics have been well supported for Annonaceae by a combination of morphological
and molecular evidence. The APG II system
places Annonaceae as most closely related to the small Magnoliid family Eupomatiaceae
.
Combined plastid
DNA marker and morphological character analysis of Anaxagorea
and other Annonaceae and basal angiosperm genera suggest that Anaxagorea is an ancestral clade
, sister to the rest of the family. This may confirm the hypothesis that morphological traits shared with other Magnoliales species (such as 2-ranked phyllotaxis, monosulcate pollen, and laminate stamens) represent ancestral characters, while derived characters observed in other genera have evolved independently multiple times. These analyses also support a biogeographic history in which Annonaceae, and perhaps other basal angiosperm clades, originated in South America during the Cretaceous
, radiating due to migration events throughout Africa to Asia and Central to North America.
Guatteria
, with its approximately 265 species, is the largest genus of Annonaceae, and might be broken into three small genera based on molecular characters from multiple locations.
Tribe Saccopetaleae is another group of Annonaceae that is awaiting review as recent molecular studies suggest that this tribe is not monophyletic at all.
: custard apple
(A. reticulata
), cherimoya
(A. cherimola
), soursop
/guanábana/graviola (A. muricata
), sweetsop (A. squamosa
), ilama
(A. diversifolia), soncoya (A. purpurea
), atemoya
(a cross between A. cherimola and A. squamosa); and biriba (Rollinia
deliciosa, which may require reclassification under Annona.). The names of many of those fruits are sometimes used interchangeably. Recently, consumption of the neotropical annonaceous plant Annona muricata (soursop
, graviola, guanabana) has been strongly associated as a causal agent in "atypical Parkinsonsism." The causative agent, annonancin, is present in many of the annonaceae. It is thought to be responsible for up to 70% of Parkinsonian conditions in Guadeloupe. Exposure is typically through traditional food and "natural" medicines.
Asimina triloba
(American pawpaw, prairie banana) has an Eastern U.S. distribution, and is currently under agricultural investigation as a commercial crop.
s. Pharmaceutical research has found antifungal, bacteriostatic, antimalarial, and especially cytostatic capability of some chemical constituents of the leaves and bark. A large number of chemical compounds, including flavonoid
s, alkaloid
s and acetogenins, have been extracted from the seeds and many other parts of these plants. Flavonoids and alkaloids have shown antibacterial properties, and have been used for treatment of medical conditions, such as skin disease, intestinal worms and inflammation
of the eye. Pharmaceutical products are currently in animal and human cell-line trials. Acetogenins are thought to have anti-HIV
and anti-cancer
effects. A wide variety of products have been developed and are available for cancer treatment.
is a family of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
s consisting of tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...
s, shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...
s or rarely liana
Liana
A liana is any of various long-stemmed, woody vines that are rooted in the soil at ground level and use trees, as well as other means of vertical support, to climb up to the canopy to get access to well-lit areas of the forest. Lianas are especially characteristic of tropical moist deciduous...
s.
With about 2300 to 2500 species and more than 130 genera,
it is the largest family in Magnoliales
Magnoliales
Magnoliales is an order of flowering plants.-Classification:The Magnoliales includes six families:* Annonaceae...
. Only five genera, Annona
Annona
Annona is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after Guatteria, containing approximately 110 species of mostly neotropical and afrotropical trees and shrubs....
, Rollinia
Rollinia
Rollinia is a genus in the family Annonaceae. While it is widely recognised as a distinct genus a recent monograph advocates its inclusion in Annona, which also contains custard apples and soursops....
, Uvaria
Uvaria
Uvaria is a genus of flowering plants in the soursop family, Annonaceae.-Selected species:* Uvaria calamistrata Hance* Uvaria chamae P.Beauv. – Finger-root* Uvaria grandiflora Uvaria is a genus of flowering plants in the soursop family, Annonaceae.-Selected species:* Uvaria calamistrata ...
, Melodorum
Melodorum
Melodorum is a genus of plant in family Annonaceae.-Species:Genus Melodorum contains 55 species, among which the following deserve mention:...
and Asimina produce edible fruits.
Its type genus
Type genus
In biological classification, a type genus is a representative genus, as with regard to a biological family. The term and concept is used much more often and much more formally in zoology than it is in botany, and the definition is dependent on the nomenclatural Code that applies:* In zoological...
is Annona
Annona
Annona is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after Guatteria, containing approximately 110 species of mostly neotropical and afrotropical trees and shrubs....
. The family is concentrated in the tropics
Tropics
The tropics is a region of the Earth surrounding the Equator. It is limited in latitude by the Tropic of Cancer in the northern hemisphere at approximately N and the Tropic of Capricorn in the southern hemisphere at S; these latitudes correspond to the axial tilt of the Earth...
, with few species found in temperate
Temperate
In geography, temperate or tepid latitudes of the globe lie between the tropics and the polar circles. The changes in these regions between summer and winter are generally relatively moderate, rather than extreme hot or cold...
regions. About 900 species are Neotropic
Neotropic
In biogeography, the Neotropic or Neotropical zone is one of the eight terrestrial ecozones. This ecozone includes South and Central America, the Mexican lowlands, the Caribbean islands, and southern Florida, because these regions share a large number of plant and animal groups.It is sometimes used...
al, 450 are Afrotropic
Afrotropic
The Afrotropic is one of the Earth's eight ecozones. It includes Africa south of the Sahara Desert, the southern and eastern fringes of the Arabian Peninsula, the island of Madagascar, southern Iran and extreme southwestern Pakistan, and the islands of the western Indian Ocean. It was formerly...
al, and the other species Indomalaya
Indomalaya
The Indomalaya ecozone is one of the eight ecozones that cover the planet's land surface. It extends across most of South and Southeast Asia and into the southern parts of East Asia....
n.
Compared to the species from the Neotropic
Neotropic
In biogeography, the Neotropic or Neotropical zone is one of the eight terrestrial ecozones. This ecozone includes South and Central America, the Mexican lowlands, the Caribbean islands, and southern Florida, because these regions share a large number of plant and animal groups.It is sometimes used...
s, very little is known about many species from Indomalaya
Indomalaya
The Indomalaya ecozone is one of the eight ecozones that cover the planet's land surface. It extends across most of South and Southeast Asia and into the southern parts of East Asia....
. Only a few attempts have been made for the phylogeny-based reclassification
Angiosperm Phylogeny Group
The Angiosperm Phylogeny Group, or APG, refers to an informal international group of systematic botanists who came together to try to establish a consensus on the taxonomy of flowering plants that would reflect new knowledge about plant relationships discovered through phylogenetic studies., three...
of the family, and those have been hampered by the Neotropic
Neotropic
In biogeography, the Neotropic or Neotropical zone is one of the eight terrestrial ecozones. This ecozone includes South and Central America, the Mexican lowlands, the Caribbean islands, and southern Florida, because these regions share a large number of plant and animal groups.It is sometimes used...
bias in the available information, with the most of the work having been done on genera and tribes.
Description
Mostly tropical, some mid-latitude, deciduousDeciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...
or evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...
trees and shrubs, some lianas, with aromatic bark, leaves, and flowers.
Stems, stalks and leaves: Bark is fibrous and aromatic. Pith
Pith
Pith, or medulla, is a tissue in the stems of vascular plants. Pith is composed of soft, spongy parenchyma cells, which store and transport nutrients throughout the plant. In eudicots, pith is located in the center of the stem. In monocots, it extends also into flowering stems and roots...
septate (fine tangential bands divided by partitions) to diaphragmed (divided by thin partitions with openings in them).
- Branching distichous (arranged in two rows/on one plane) or spiral.
- Leaves are alternate, two-ranked, simple, pinnatePinnatePinnate is a term used to describe feather-like or multi-divided features arising from both sides of a common axis in plant or animal structures, and comes from the Latin word pinna meaning "feather", "wing", or "fin". A similar term is pectinate, which refers to a comb-like arrangement of parts...
ly veined, and have leaf stalksPetiole (botany)In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...
. Stipules absent.
Flowers: Flower stalks
Peduncle (botany)
In botany, a peduncle is a stem supporting an inflorescence, or after fecundation, an infructescence.The peduncle is a stem, usually green and without leaves, though sometimes colored or supporting small leaves...
are axillary to (on the opposite side of shoot from) leaf scars on old wood and sometimes from leaves on new shoots. The flowers are usually trimerous
Merosity
Merosity is the number of component parts in each whorl of a plant structure. It is most commonly used in the context of flowers, in which case it refers to the number of sepals in the calyx, the number of petals in the corolla, and the number of stamens in each whorl of the androecium...
; borne singly or in compound inflorescence
Inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Strictly, it is the part of the shoot of seed plants where flowers are formed and which is accordingly modified...
s; bisexual
Bisexual (disambiguation)
Bisexual may refer to:*Bisexuality, in human sexuality, describes a man or woman that is sexually or romantically attracted to persons of both sexes*Bisexual characteristics, having an ambiguous sexual identity Bisexual may refer to:*Bisexuality, in human sexuality, describes a man or woman that is...
and rarely unisexual
Gonochorism
In biology, gonochorism or unisexualism describes sexually reproducing species in which individuals have just one of at least two distinct sexes. The term is most often used with animals . The sex of an individual may change during its lifetime, this can for example be found in parrotfish...
. The receptacle might become enlarged, elevated or flat. The outer whorls
Perianth
The term perianth has two similar but separate meanings in botany:* In flowering plants, the perianth are the outer, sterile whorls of a flower...
are inserted below the ovaries, and have valvate (overlapping) or imbricate (nonoverlapping) segments. Usually two to four persistent sepal
Sepal
A sepal is a part of the flower of angiosperms . Collectively the sepals form the calyx, which is the outermost whorl of parts that form a flower. Usually green, sepals have the typical function of protecting the petals when the flower is in bud...
s that are distinct or connate (fused) at the base. Six petals in two unequal whorls of three with larger outer whorls and fleshier inner whorls that might share the same nectar glands, or six to fifteen petals, with impressed veins on their inner face. Ten to twenty (or many more) stamens inserted below the ovary, spirally arranged and forming a ball or flat-topped mass with short and stout filaments and linear to oblong anthers which face outward and open longitudinally. Each flower can have from one to many pistils, distinct to connate, with stigmas distinct. Marginal placentation, each pistil bearing one locule
Locule
A locule is a small cavity or compartment within an organ or part of an organism ....
, with one to many ovules. Style short and thick, with terminal stigma.
Fruits and seeds: Fruits are distinct (berries) or coalesce (into syncarps). Seeds are one to many per pistil; have a fleshy and usually brightly colored cover, have ruminate endosperm (nutritive tissue surrounding the embryo) and are oily.
Systematics
MonophylyMonophyly
In common cladistic usage, a monophyletic group is a taxon which forms a clade, meaning that it contains all the descendants of the possibly hypothetical closest common ancestor of the members of the group. The term is synonymous with the uncommon term holophyly...
and inter-familial systematics have been well supported for Annonaceae by a combination of morphological
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....
and molecular evidence. The APG II system
APG II system
The APG II system of plant classification is the second, now obsolete, version of a modern, mostly molecular-based, system of plant taxonomy that was published in April 2003 by the Angiosperm Phylogeny Group. It was a revision of the first APG system, published in 1998, and was superseded in 2009...
places Annonaceae as most closely related to the small Magnoliid family Eupomatiaceae
Eupomatiaceae
Eupomatiaceae is the botanical name for a family of flowering plants. Such a family has been recognised by most taxonomists.The APG II system, of 2003 , also recognizes this family, and assigns it to the order Magnoliales in the clade magnoliids...
.
{clade| style=font-size:75%;line-height:75% | label1=Magnoliidae Magnoliidae Magnoliids are a group of about 9,000 species of flowering plants, including magnolias, nutmeg, bay laurel, cinnamon, avocado, black pepper, and many others. They are characterized by trimerous flowers, pollen with one pore, and usually branching-veined leaves.-Classification:Traditionally,... |
1= }} |
Combined plastid
Plastid
Plastids are major organelles found in the cells of plants and algae. Plastids are the site of manufacture and storage of important chemical compounds used by the cell...
DNA marker and morphological character analysis of Anaxagorea
Anaxagorea
Anaxagorea is a genus of plant in family Annonaceae. It contains the following species Anaxagorea is a genus of plant in family Annonaceae. It contains the following species Anaxagorea is a genus of plant in family Annonaceae. It contains the following species (but this list may be incomplete, and...
and other Annonaceae and basal angiosperm genera suggest that Anaxagorea is an ancestral clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...
, sister to the rest of the family. This may confirm the hypothesis that morphological traits shared with other Magnoliales species (such as 2-ranked phyllotaxis, monosulcate pollen, and laminate stamens) represent ancestral characters, while derived characters observed in other genera have evolved independently multiple times. These analyses also support a biogeographic history in which Annonaceae, and perhaps other basal angiosperm clades, originated in South America during the Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...
, radiating due to migration events throughout Africa to Asia and Central to North America.
Guatteria
Guatteria
Guatteria is a genus of plant in family Annonaceae. It contains the following species :* Guatteria anomala, R.E. Fries* Guatteria atabapensis, Aristeg.* Guatteria augusti, Diels...
, with its approximately 265 species, is the largest genus of Annonaceae, and might be broken into three small genera based on molecular characters from multiple locations.
Tribe Saccopetaleae is another group of Annonaceae that is awaiting review as recent molecular studies suggest that this tribe is not monophyletic at all.
Food
The large, edible, pulpy fruits of some members typically called "anona" by Spanish and Portuguese speaking people of the family's Neotropical range, include species of AnnonaAnnona
Annona is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after Guatteria, containing approximately 110 species of mostly neotropical and afrotropical trees and shrubs....
: custard apple
Custard apple
Custard apple, a common name, can refer to:*Custard-apple, also called bull's heart, the fruit of the tree Annona reticulata.*The custard-apple tree itself, Annona reticulataThe term may also refer to:...
(A. reticulata
Annona reticulata
Annona reticulata is a small deciduous or semi-evergreen tree in the plant family Annonaceae.It is best known for its fruit, called custard-apple, a name it shares with fruits of other species from the same genus: A. cherimola and A. squamosa or sometimes it is called wild-sweetsop,...
), cherimoya
Cherimoya
The Cherimoya is the fruit of the species Annona cherimola, which is native to the Andes. Today they are grown throughout South and Central America.-Description:...
(A. cherimola
Annona cherimola
Annona cherimola , and originally called Chirimuya by the Inca people who lived where it was growing in the Andes of South America, is an edible fruit bearing species of the genus Annona from the family Annonaceae is now widely cultivated mostly for its sweet fruits that share the name...
), soursop
Soursop
The Soursop is tripti broadleaf, flowering, evergreen tree native to Central America, the Caribbean and northern South America, Colombia and Brazil, Mexico, Peru, Venezuela, Soursop is also native to sub-Saharan African countries that lie within the tropics. Today, it is also grown in some areas...
/guanábana/graviola (A. muricata
Annona muricata
Annona muricata is a member of the family of Custard apple trees called Annonaceae and a species of the genus Annona known mostly for its edible fruits Anona. Annona muricata produces fruits that are usually called Soursop due to its slightly acidic taste when ripe. A...
), sweetsop (A. squamosa
Annona squamosa
Annona squamosa a small well-branched tree or shrub that bears edible fruits called sugar-apple, species of the genus Annona and member of the family Annonaceae more willing to grow at lower altitudes than its relatives Annona reticulata and Annona cherimola making it the most widely cultivated of...
), ilama
Ilama (fruit)
The ilama is a tropical fruit tree found in Central America. The name is derived from the Spanish from the Nahuatl ilamatzapotl, of which the rough translation is "old woman's sapote". The name is also applied to a similar fruit, soncoya or cabeza de negro The ilama (also known as the tree of...
(A. diversifolia), soncoya (A. purpurea
Annona purpurea
Annona purpurea is an edible fruit and medicinal plant in the Annonaceae family. It is native to Mexico, Central America, and parts of South America. Its common names include soncoya, sincuya, and cabeza de negro. It is a tree reaching 6 to 10 meters in maximum height. It has hairy leaves and...
), atemoya
Atemoya
The atemoya, Annona × atemoya, is a hybrid of two fruits – the sugar apple and the cherimoya – which are both native to the American tropics. This fruit is popular in Taiwan, where it is known as the "pineapple sugar apple" , and is sometimes mistaken for a cross between the sugar...
(a cross between A. cherimola and A. squamosa); and biriba (Rollinia
Rollinia
Rollinia is a genus in the family Annonaceae. While it is widely recognised as a distinct genus a recent monograph advocates its inclusion in Annona, which also contains custard apples and soursops....
deliciosa, which may require reclassification under Annona.). The names of many of those fruits are sometimes used interchangeably. Recently, consumption of the neotropical annonaceous plant Annona muricata (soursop
Soursop
The Soursop is tripti broadleaf, flowering, evergreen tree native to Central America, the Caribbean and northern South America, Colombia and Brazil, Mexico, Peru, Venezuela, Soursop is also native to sub-Saharan African countries that lie within the tropics. Today, it is also grown in some areas...
, graviola, guanabana) has been strongly associated as a causal agent in "atypical Parkinsonsism." The causative agent, annonancin, is present in many of the annonaceae. It is thought to be responsible for up to 70% of Parkinsonian conditions in Guadeloupe. Exposure is typically through traditional food and "natural" medicines.
Asimina triloba
Asimina triloba
Asimina triloba, the pawpaw, paw paw, paw-paw, or common pawpaw, is a species of Asimina in the same plant family as the custard-apple, cherimoya, sweetsop, ylang-ylang and soursop...
(American pawpaw, prairie banana) has an Eastern U.S. distribution, and is currently under agricultural investigation as a commercial crop.
Medical
The bark, leaves and roots of some species are used in folk medicineFolk medicine
-Description:Refers to healing practices and ideas of body physiology and health preservation known to a limited segment of the population in a culture, transmitted informally as general knowledge, and practiced or applied by anyone in the culture having prior experience.All cultures and societies...
s. Pharmaceutical research has found antifungal, bacteriostatic, antimalarial, and especially cytostatic capability of some chemical constituents of the leaves and bark. A large number of chemical compounds, including flavonoid
Flavonoid
Flavonoids , are a class of plant secondary metabolites....
s, alkaloid
Alkaloid
Alkaloids are a group of naturally occurring chemical compounds that contain mostly basic nitrogen atoms. This group also includes some related compounds with neutral and even weakly acidic properties. Also some synthetic compounds of similar structure are attributed to alkaloids...
s and acetogenins, have been extracted from the seeds and many other parts of these plants. Flavonoids and alkaloids have shown antibacterial properties, and have been used for treatment of medical conditions, such as skin disease, intestinal worms and inflammation
Inflammation
Inflammation is part of the complex biological response of vascular tissues to harmful stimuli, such as pathogens, damaged cells, or irritants. Inflammation is a protective attempt by the organism to remove the injurious stimuli and to initiate the healing process...
of the eye. Pharmaceutical products are currently in animal and human cell-line trials. Acetogenins are thought to have anti-HIV
HIV
Human immunodeficiency virus is a lentivirus that causes acquired immunodeficiency syndrome , a condition in humans in which progressive failure of the immune system allows life-threatening opportunistic infections and cancers to thrive...
and anti-cancer
Cancer
Cancer , known medically as a malignant neoplasm, is a large group of different diseases, all involving unregulated cell growth. In cancer, cells divide and grow uncontrollably, forming malignant tumors, and invade nearby parts of the body. The cancer may also spread to more distant parts of the...
effects. A wide variety of products have been developed and are available for cancer treatment.
Insecticides
Flavonoids and alkaloids contained in the leaves and bark of several species of the family have shown insecticidal properties.Other
- Some species of the family, such as Cananga odorata (ylang-ylangYlang-ylangCananga odorata, commonly called Ylang-ylang , cananga tree, ilang-ilang, kenanga , fragrant cananga, Macassar-oil plant or perfume tree),is a tree valued for its perfume...
) also have aromatic oil and are used for perfumePerfumePerfume is a mixture of fragrant essential oils and/or aroma compounds, fixatives, and solvents used to give the human body, animals, objects, and living spaces "a pleasant scent"...
s or spiceSpiceA spice is a dried seed, fruit, root, bark, or vegetative substance used in nutritionally insignificant quantities as a food additive for flavor, color, or as a preservative that kills harmful bacteria or prevents their growth. It may be used to flavour a dish or to hide other flavours...
s. - The strong bark is used for carrying burdens in the Amazon RainforestAmazon RainforestThe Amazon Rainforest , also known in English as Amazonia or the Amazon Jungle, is a moist broadleaf forest that covers most of the Amazon Basin of South America...
and for wooden implements, such as tool handles and pegs. The wood is valued as firewoodFirewoodFirewood is any wood-like material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form....
. - Yellow and brown dyes.
- Some species are also grown as ornamental plantOrnamental plantOrnamental plants are plants that are grown for decorative purposes in gardens and landscape design projects, as house plants, for cut flowers and specimen display...
s, especially the Indian species Polyalthia longifoliaPolyalthia longifoliaPolyalthia longifolia is a lofty evergreen tree, native to India, commonly planted due to its effectiveness in alleviating noise pollution. It exhibits symmetrical pyramidal growth with willowy weeping pendulous branches and long narrow lanceolate leaves with undulate margins...
pendula.
Genera
- Afroguatteria
- AlphonseaAlphonseaAlphonsea is a genus of plant in family Annonaceae. It contains the following species :* Alphonsea curtisii, King* Alphonsea hainanensis, Merr. & Chun* Alphonsea kingii, Sinclair* Alphonsea lucida, King...
- Ambavia
- AnaxagoreaAnaxagoreaAnaxagorea is a genus of plant in family Annonaceae. It contains the following species Anaxagorea is a genus of plant in family Annonaceae. It contains the following species Anaxagorea is a genus of plant in family Annonaceae. It contains the following species (but this list may be incomplete, and...
- Ancana
- AnnickiaAnnickiaAnnickia is a genus of plant in family Annonaceae. It contains the following species :* Annickia kummeriae, Setten & Maas...
- AnnonaAnnonaAnnona is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after Guatteria, containing approximately 110 species of mostly neotropical and afrotropical trees and shrubs....
- Anomianthus
- AnonidiumAnonidiumAnonidium is a genus of plant in family Annonaceae. It contains the following species :* Anonidium usambarense R.E. Fries* Anonidium mannii Engl. & Diels - junglesop...
- ArtabotrysArtabotrysArtabotrys There are over 100 species of Old World tropics, with 31 species in Africa. It is part of the custard family. All are woody small trees or shrubs with recurved hooks with a tendency to climb. Leaves are simple and alternate, without hairs. Bisexual flowers are borne singly or in cluster...
- trail-grape - Asimina
- Asteranthe
- Balonga
- Bocagea
- Bocageopsis
- BoutiqueaBoutiqueaBoutiquea is a genus of plant in family Annonaceae. It contains the following species :* Boutiquea platypetala, Le Thomas...
- Cananga - Ylang-ylangYlang-ylangCananga odorata, commonly called Ylang-ylang , cananga tree, ilang-ilang, kenanga , fragrant cananga, Macassar-oil plant or perfume tree),is a tree valued for its perfume...
- Cardiopetalum
- Cleistochlamys
- Cleistopholis
- Craibella
- Cremastosperma
- CyathocalyxCyathocalyxCyathocalyx is a small genus with about 36 species distributed from southern India, Sri Lanka, through Malaysia, Indomalayan islands and reaches as far as Fiji in the South Pacific....
- Cyathostemma
- Cymbopetalum
- Dasoclema
- Dasymaschalon
- DeeringothamnusDeeringothamnusDeeringothamnus is a genus of flowering plants with two species, belonging to the Annonaceae family. They are native to Florida. Both are federally listed endangered species of the United States.-External links:** ***...
- false pawpaw - Dendrokingstonia
- Dennettia
- Desmopsis
- DesmosDesmosDesmos is a genus of trees and shrubs in the plant family Annonaceae.-Selected species:*Desmos cambodicus*Desmos chinensis*Desmos chryseus*Desmos cochinchinensis*Desmos dasymaschalus*Desmos dubius*Desmos dumosus...
- Diclinanona
- Dielsiothamnus
- Disepalum
- Duckeanthus
- DuguetiaDuguetiaDuguetia is a genus of trees and shrubs in the plant family Annonaceae with approximately 90 species in central and South America, and four species in west Africa.- Species :*Duguetia aberrans Maas *Duguetia adiscandra Jans.-Jac....
- Ellipeia
- Ellipeiopsis
- Enicosanthum
- Ephedranthus
- ExelliaExelliaExellia is a genus of extinct spadefish that lived in the Tethys Ocean during the early Paleogene. The adult form is shaped akin to a large spadefish or a short dolphinfish, with very large pelvic fins, and a long dorsal fin starting from in front of the eyes to near the base of the caudal peduncle...
- FissistigmaFissistigmaFissistigma is a genus of plant in family Annonaceae. It contains the following species :* Fissistigma cupreonitens, Merr. & Chun* Fissistigma tungfangense, Tsiang & P.T.Li...
- Fitzalania
- Friesodielsia
- Froesiodendron
- Fusaea
- Gilbertiella
- GoniothalamusGoniothalamusGoniothalamus is one of the largest palaeotropical genera of plant in family Annonaceae.-Species list:It contains the following species :Indian Region...
- Greenwayodendron
- GuamiaGuamiaGuamia is a monotypic genus of flowering plants in the soursop family, Annonaceae. The sole species it contains, Guamia mariannae, is endemic to the Marianas Islands.-External links:...
- GuatteriaGuatteriaGuatteria is a genus of plant in family Annonaceae. It contains the following species :* Guatteria anomala, R.E. Fries* Guatteria atabapensis, Aristeg.* Guatteria augusti, Diels...
- haya minga, haya blanca - Guatteriella
- GuatteriopsisGuatteriopsisGuatteriopsis is a genus of plant in family Annonaceae. It contains the following species :* Guatteriopsis ramiflora, D. R. Simpson...
- Haplostichanthus
- Heteropetalum
- HexalobusHexalobusHexalobus is a genus of plant in family Annonaceae. It contains the following species :* Hexalobus mossambicensis, N. Robson* Hexalobus salicifolius, Engl. & Diels...
- Hornschuchia
- IsolonaIsolonaIsolona is a genus of plant in family Annonaceae .- Species of Isolona :It contains species :* Isolona campanulata, Engler & Diels,* Isolona cauliflora, Verdc....
- Letestudoxa
- LettowianthusLettowianthusLettowianthus is a genus of plant in family Annonaceae. It contains the following species :* Lettowianthus stellatus, Diels...
- MalmeaMalmeaMalmea is a genus of plant in family Annonaceae. It contains the following species :* Malmea cuspidata, Diels...
- Marsypopetalum
- Meiocarpidium
- MeiogyneMeiogyneMeiogyne is a genus of flowering plants with about 24 species belonging to the family Annonaceae. Its is native from south western India through Thailand, Malaysia, Java, Sumatra, Borneo, Hainan, The Philippines to Australia.- Description :...
- MelodorumMelodorumMelodorum is a genus of plant in family Annonaceae.-Species:Genus Melodorum contains 55 species, among which the following deserve mention:...
- MezzettiaMezzettiaMezzettia is a genus of plant in family Annonaceae. It contains the following species :* Mezzettia herveyana, Oliver...
- Mezzettiopsis
- MiliusaMiliusaMiliusa is a genus of plant in family Annonaceae. It contains the following species :* Miliusa amplexicaulis, Ridley* Miliusa nilagirica, Bedd.* Miliusa parviflora, Ridley...
- Mischogyne
- MitrellaMitrella (plant)Mitrella is a genus of plants in the Annonaceae family.-Species:Genus Mitrella contains the following species:* Mitrella aberrans* Mitrella beccarii* Mitrella dielsii* Mitrella kentii* Mitrella ledermannii...
- MitrephoraMitrephoraThe genus Mitrephora, of family Annonaceae, comprises around 40 species of small trees and shrubs found in tropical Asia and northern Australasia. The area of distribution extends from China in the North down to Queensland, Australia in the South. The genus is widely found in southern India and...
- MkiluaMkiluaMkilua is a genus of plant in family Annonaceae. It contains the following species :* Mkilua fragrans, Verdc....
- Monanthotaxis
- Monocarpia
- MonocyclanthusMonocyclanthusMonocyclanthus is a genus of plant in family Annonaceae. It contains the following species :* Monocyclanthus vignei, Keay...
- MonodoraMonodoraMonodora is a genus of plant in family Annonaceae. It contains approximately 35 species, distributed throughout tropical Africa.-Some species:* Monodora grandiflora Benth.* Monodora unwinii, Hutch. & Dalz....
- MosannonaMosannonaMosannona is a genus of plant in family Annonaceae. It contains the following species :* Mosannona pachiteae, Chatrou* Mosannona pacifica, Chatrou...
- NeostenantheraNeostenantheraNeostenanthera is a genus of plant in family Annonaceae. It contains the following species :* Neostenanthera hamata, Exell* Neostenanthera robsonii, Le Thomas...
- Neo-uvaria
- Onychopetalum
- Ophrypetalum
- Oreomitra
- OropheaOropheaOrophea is a genus of plant in family Annonaceae. It contains the following species :* Orophea hastata, King* Orophea palawanensis, Elm.* Orophea submaculata, Elm.* Orophea thomsoni, Beddome...
- OxandraOxandraOxandra is a genus of flowering plants in the soursop family, Annonaceae.-Selected species:* Oxandra lanceolata Baill.* Oxandra laurifolia A.Rich.* Oxandra leucodermis Warm.-External links:...
- blacklancewood, hayaHayaHaya may refer to:* Haya , a genus of plants in the family Caryophyllaceae* Haya , a genus of basal ornithopod dinosaur that lived during the Late Cretaceous from Mongolia... - Pachypodanthium
- Papualthia
- Petalolophus
- PhaeanthusPhaeanthusPhaeanthus is a genus of plant in family Annonaceae. It contains the following species :*Phaeanthus acuminata Merr. 1905*Phaeanthus crassipetalus Becc. *Phaeanthus cumingii Miq. 1858...
- Phoenicanthus
- PiptostigmaPiptostigmaPiptostigma is a genus of plant in family Annonaceae. It contains the following species :* Piptostigma calophyllum, Mildler. & Diels* Piptostigma fugax, A.Chev. ex Hutch. & Dlaz....
- Platymitra
- PolyalthiaPolyalthiaPolyalthia is a genus of plant in family Annonaceae. It contains the following species :* Polyalthia angustissima Ridl.* Polyalthia chrysotricha Ridley* Polyalthia elmeri Merr....
- PolyceratocarpusPolyceratocarpusPolyceratocarpus is a genus of plant in family Annonaceae. It contains the following species :* Polyceratocarpus scheffleri, Engl. & Diels...
- PopowiaPopowiaPopowia is a genus of plant in family Annonaceae. It contains the following species :* Popowia beddomeana, Hook.f. & Thomson* Popowia fusca, King* Popowia pauciflora, Maingay ex Hook.f. & Th....
- Porcelia
- Pseudartabotrys
- Pseudephedranthus
- PseudoxandraPseudoxandraPseudoxandra is a genus of plant in family Annonaceae. It contains the following species :* Pseudoxandra williamsii, R. E. Fr....
- PseuduvariaPseuduvariaPseuduvaria is a genus of plant in family Annonaceae. It contains the following species :* Pseuduvaria cerina, J.Sincl.* Pseuduvaria galeata, J. Sincl.* Pseuduvaria nervosa, J. Sincl....
- Pyramidanthe
- Raimondia
- Reedrollinsia
- RichellaRichellaRichella is a genus of plant in family Annonaceae. It contains the following species :* Richella hainanensis, Tsiang & P.T.Li...
- RolliniaRolliniaRollinia is a genus in the family Annonaceae. While it is widely recognised as a distinct genus a recent monograph advocates its inclusion in Annona, which also contains custard apples and soursops....
- wild sugar-apple - Ruizodendron
- SageraeaSageraeaSageraea is a genus of plant in family Annonaceae. It contains the following species :* Sageraea grandiflora, Dunn* Sageraea laurifolia, Blatter* Sageraea thwaitesii, Hook.f. & Thomson...
- Sanrafaelia
- SapranthusSapranthusSapranthus is a genus of flowering woody plants in the family Annonaceae.Their flowers are pollinated by flies, and smell accordingly like decaying organic matter.-Species:* Sapranthus borealis* Sapranthus campechianus...
- Schefferomitra
- Sphaerocoryne
- Stelechocarpus - kepel
- StenanonaStenanonaStenanona is a genus of plant in family Annonaceae. It contains the following species :* Stenanona panamensis, Standl....
- TetrameranthusTetrameranthusTetrameranthus is a genus of plant in family Annonaceae. It contains the following species :* Tetrameranthus globuliferus, Westra...
- ToussaintiaToussaintiaToussaintia is a genus of plant in family Annonaceae. It contains the following species :* Toussaintia orientalis, Verdc....
- Tridimeris
- TrigynaeaTrigynaeaTrigynaea is a genus of plant in family Annonaceae. It contains the following species :* Trigynaea triplinervis, D.M. Johnson & N.A. Murray...
- TrivalvariaTrivalvariaTrivalvaria is a genus of plant in family Annonaceae. It contains the following species :* Trivalvaria nervosa, James Sincl....
- UnonopsisUnonopsisUnonopsis is a genus of plant in family Annonaceae. It contains the following species :* Unonopsis magnifolia, R.E. Fr.* Unonopsis velutina, P. Maas...
- UvariaUvariaUvaria is a genus of flowering plants in the soursop family, Annonaceae.-Selected species:* Uvaria calamistrata Hance* Uvaria chamae P.Beauv. – Finger-root* Uvaria grandiflora Uvaria is a genus of flowering plants in the soursop family, Annonaceae.-Selected species:* Uvaria calamistrata ...
- UvariastrumUvariastrumUvariastrum is a genus of plant in family Annonaceae. It contains the following species :* Uvariastrum zenkeri, Engl. & Diels...
- UvariodendronUvariodendronUvariodendron is a genus of plant in family Annonaceae. It contains the following species :* Uvariodendron anisatum, Verdc.* Uvariodendron connivens, R.E.Fries...
- UvariopsisUvariopsisUvariopsis is a genus of plant in family Annonaceae. It contains the following species :* Uvariopsis bisexualis, Verdc.* Uvariopsis submontana, Kenfack, Gosline & Gereau...
- Woodiellantha
- XylopiaXylopiaXylopia is a genus of plant in family Annonaceae. This genus is solely composed of trees, distribution of which is pantropical . It contains the following species :* Xylopia africana, Oliv....