Spea
Encyclopedia
Spea is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

n amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

 commonly referred to as the Western Spadefoot Toads. They differ greatly from true toad
Toad
A toad is any of a number of species of amphibians in the order Anura characterized by dry, leathery skin , short legs, and snoat-like parotoid glands...

s (those of the family Bufonidae) by having eye
Eye
Eyes are organs that detect light and convert it into electro-chemical impulses in neurons. The simplest photoreceptors in conscious vision connect light to movement...

s with vertical pupil
Pupil
The pupil is a hole located in the center of the iris of the eye that allows light to enter the retina. It appears black because most of the light entering the pupil is absorbed by the tissues inside the eye. In humans the pupil is round, but other species, such as some cats, have slit pupils. In...

s, no parotoid gland
Parotoid gland
The parotoid gland is an external skin gland on the back, neck, and shoulder of toads and some frogs and salamanders. It secretes a milky alkaloid substance to deter predators...

, and relatively smooth skin
Skin
-Dermis:The dermis is the layer of skin beneath the epidermis that consists of connective tissue and cushions the body from stress and strain. The dermis is tightly connected to the epidermis by a basement membrane. It also harbors many Mechanoreceptors that provide the sense of touch and heat...

. Their most distinctive feature is a spade-like projection on their hind feet, from which their common name is derived. This projection enables spadefoot toads to dig in loose soils with ease.

Species

Spea species were once classified with their Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

an cousins in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Pelobatidae
Pelobatidae
The European spadefoot toads are a family of frogs, the Pelobatidae, with only one extant genus Pelobates, containing four species. They are native to Europe, the Mediterranean, northwestern Africa and western Asia....

, but have since been reclassified to their own family, Scaphiopodidae with other North American species. There are four species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 in the genus Spea:
  • Plains Spadefoot Toad, Spea bombifrons
    Spea bombifrons
    The Plains Spadefoot Toad is a species of spadefoot toad which ranges from southwestern Canada, throughout the Great Plains of the western United States, and into northern Mexico...

    (Cope
    Edward Drinker Cope
    Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

    , 1863)
  • Western Spadefoot Toad, Spea hammondii
    Spea hammondii
    The Western Spadefoot Toad is a relatively smooth-skinned species of spadefoot toad. Its eyes are pale gold with vertical pupils. It has a green or grey dorsum often with skin tubercles tipped in orange, and it is a whitish color on the ventrum. It has a wedge-shaped black spade on each hind foot...

    (Baird
    Spencer Fullerton Baird
    Spencer Fullerton Baird was an American ornithologist, ichthyologist and herpetologist. Starting in 1850 he was assistant-secretary and later secretary of the Smithsonian Institution in Washington, D.C...

    , 1859)
  • Great Basin Spadefoot Toad, Spea intermontana (Cope
    Edward Drinker Cope
    Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

    , 1863)
  • Mexican Spadefoot Toad, Spea multiplicata
    Spea multiplicata
    The New Mexico Spadefoot Toad is a species of American spadefoot toad found in the southwestern United States and Mexico. Like other species of spadefoot toad, they get their name from a distinctive spade-like projections on their hind legs which enable them to dig in sandy soils...

    (Cope
    Edward Drinker Cope
    Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

    , 1863)

Related species

Genus Scaphiopus
Scaphiopus
Scaphiopus is a genus of North American amphibian commonly referred to as the Southern Spadefoot Toads. They differ greatly from true toads by having eyes with vertical pupils, no parotoid gland, and relatively smooth skin. Their most distinctive feature is a spade-like projection on their hind...

, Southern Spadefoot Toads
  • Couch's Spadefoot, Scaphiopus couchii
    Scaphiopus couchii
    Couch's Spadefoot Toad is a species of North American spadefoot toad. The epithet couchii is in honor of American naturalist Darius Nash Couch, who collected the first specimen while on a personal expedition to northern Mexico to collect plant, mineral and animal specimens for the Smithsonian...

    (Baird
    Spencer Fullerton Baird
    Spencer Fullerton Baird was an American ornithologist, ichthyologist and herpetologist. Starting in 1850 he was assistant-secretary and later secretary of the Smithsonian Institution in Washington, D.C...

    , 1854)
  • Eastern Spadefoot, Scaphiopus holbrookii
    Scaphiopus holbrookii
    The Eastern Spadefoot is a species of spadefoot found in North America. It has one spur on each of its back feet for burrowing. It spends almost all of its life deep underground; coming out only to breed, and sometimes eat. It remains in a type of hibernation almost all its life...

    (Harlan
    Richard Harlan
    Richard Harlan was an American naturalist, zoologist, physicist and paleontologist....

    , 1835)
  • Hurter's Spadefoot, Scaphiopus hurterii
    Scaphiopus hurterii
    Hurter's Spadefoot Toad is a species of spadefoot toad found in the south central United States and northern Mexico. It was once classified as a subspecies of the eastern spadefoot toad, but it has been granted its own species status...

    (Strecker, 1910)
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