Dubautia plantaginea
Encyclopedia
Dubautia plantaginea is a rare species of flowering plant in the aster family
Asteraceae
The Asteraceae or Compositae , is an exceedingly large and widespread family of vascular plants. The group has more than 22,750 currently accepted species, spread across 1620 genera and 12 subfamilies...

 known by the common name plantainleaf dubautia. It is endemic to Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

 where it is the only member of the silversword alliance
Silversword alliance
Silversword alliance refers to an adaptive radiation of over 50 species in the composite or sunflower family, Asteraceae. The group is endemic to Hawaii, and is derived from a single immigrant to the islands...

 that is found on all six of the largest islands
Hawaiian Islands
The Hawaiian Islands are an archipelago of eight major islands, several atolls, numerous smaller islets, and undersea seamounts in the North Pacific Ocean, extending some 1,500 miles from the island of Hawaii in the south to northernmost Kure Atoll...

 (Kaua'i, O'ahu, Moloka'i, Lana'i, Maui
Maui
The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

, Hawai'i). Two of the three subspecies are rare and endangered
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

. Like other Dubautia this plant is called na`ena`e.

This plant varies in morphology
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

, taking the form of a small shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 to a tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 up to 7 meters tall. There are three subspecies. The dwarf subspecies, ssp. humilis, is endemic to Maui
Maui
The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

, where there is only one population consisting of about 50 plants. This subspecies was federally listed as an endangered species in 1999. The ssp. magnifolia is a shrub or a tree which is endemic to Kaua'i, where there are no more than 2 populations remaining. It was listed as endangered in 2010.

These plants grow in moist and wet forest habitat with 75 to over 700 centimeters of precipitation annually.

Threats to this species and its habitat include landslide
Landslide
A landslide or landslip is a geological phenomenon which includes a wide range of ground movement, such as rockfalls, deep failure of slopes and shallow debris flows, which can occur in offshore, coastal and onshore environments...

s, rockslide
Rockslide
A rockslide is a type of landslide caused by rock failure in which part of the plane of failure passes through intact rock and where material collapses en masse and not in individual blocks.The mode of failure is different from that of a rock-fall....

s, erosion
Erosion
Erosion is when materials are removed from the surface and changed into something else. It only works by hydraulic actions and transport of solids in the natural environment, and leads to the deposition of these materials elsewhere...

, flood
Flood
A flood is an overflow of an expanse of water that submerges land. The EU Floods directive defines a flood as a temporary covering by water of land not normally covered by water...

ing, and invasive plant species
Invasive species
"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

 such as Hilo grass (Paspalum conjugatum). The ssp. magnifolia was decimated by Hurricane Iniki
Hurricane Iniki
Hurricane Iniki was the most powerful hurricane to strike the U.S. state of Hawaii in recorded history. Forming on September 5 during the strong El Niño of 1991–1994, Iniki was one of eleven Central Pacific tropical cyclones during the 1992 season. It attained tropical storm status on...

in 1992.
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