Colletes hederae
Encyclopedia
Colletes hederae, the Ivy Bee, is a species of plasterer bee belonging to the family Colletidae
Colletidae
Colletidae is a family of bees, and are often referred to collectively as plasterer bees or polyester bees, due to the method of smoothing the walls of their nest cells with secretions applied with their mouthparts; these secretions dry into a cellophane-like lining...

 subfamily Colletinae.

These mining bees are known from Austria
Austria
Austria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...

, Belgium
Belgium
Belgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...

, British Isles
British Isles
The British Isles are a group of islands off the northwest coast of continental Europe that include the islands of Great Britain and Ireland and over six thousand smaller isles. There are two sovereign states located on the islands: the United Kingdom of Great Britain and Northern Ireland and...

, Croatia
Croatia
Croatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...

, Cyprus
Cyprus
Cyprus , officially the Republic of Cyprus , is a Eurasian island country, member of the European Union, in the Eastern Mediterranean, east of Greece, south of Turkey, west of Syria and north of Egypt. It is the third largest island in the Mediterranean Sea.The earliest known human activity on the...

, France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

, Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

, Greece
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

, Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

, Luxembourg
Luxembourg
Luxembourg , officially the Grand Duchy of Luxembourg , is a landlocked country in western Europe, bordered by Belgium, France, and Germany. It has two principal regions: the Oesling in the North as part of the Ardennes massif, and the Gutland in the south...

, Netherlands
Netherlands
The Netherlands is a constituent country of the Kingdom of the Netherlands, located mainly in North-West Europe and with several islands in the Caribbean. Mainland Netherlands borders the North Sea to the north and west, Belgium to the south, and Germany to the east, and shares maritime borders...

, Serbia
Serbia
Serbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...

, Slovenia
Slovenia
Slovenia , officially the Republic of Slovenia , is a country in Central and Southeastern Europe touching the Alps and bordering the Mediterranean. Slovenia borders Italy to the west, Croatia to the south and east, Hungary to the northeast, and Austria to the north, and also has a small portion of...

, Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

 and Switzerland
Switzerland
Switzerland name of one of the Swiss cantons. ; ; ; or ), in its full name the Swiss Confederation , is a federal republic consisting of 26 cantons, with Bern as the seat of the federal authorities. The country is situated in Western Europe,Or Central Europe depending on the definition....

.
They have been only recently described (Schmidt & Westrich 1993) as a distinct species. Until then, they were confused with another species of Colletes
Colletes
The genus Colletes is a very large group of ground-nesting bees within the bee family Colletidae, with over 450 species worldwide, primarily in the Northern Hemisphere. These bees tend to be solitary, though sometimes nest close together in so-called aggregations...

the morphologically very similar Colletes halophilus.

The thorax of the adults is covered by orange-brown hair, while each abdominal segment has an apical orangey hair-band. The females are on average 13 millimetre (0.511811023622047 in) long, while the males are about 10 mm (0.393700787401575 in) long.

The adults emerge late in the year (the males from late August and the females a little later in early September) and remain on the wing until early November. The principal pollen forage plant is Ivy (Hedera helix
Hedera helix
Hedera helix is a species of ivy native to most of Europe and western Asia. It is labeled as an invasive species in a number of areas where it has been introduced.-Description:...

)(hence the specific epithet hederae) , but both sexes will also nectar at Ivy flowers too. When Ivy is scarce, other species of plants are also visited. The females supply the larval brood almost cells exclusively with nectar and pollen of ivy flowers. When Ivy flowering is delayed, females may also collect pollen at various members of the Daisy family (Asteraceae
Asteraceae
The Asteraceae or Compositae , is an exceedingly large and widespread family of vascular plants. The group has more than 22,750 currently accepted species, spread across 1620 genera and 12 subfamilies...

).

These are solitary bees, and do not live in colonies and do not overwinter as adults. They nest in clay-sandy soils, especially in loess hills and soft-rock cliffs. Like many other solitary bees, they can often be found nesting in dense aggregations, sometimes numbering many tens of thousands of nests. In parts of the west European range of the species, Colletes hederae are fequently parasitized by the larvae of the meloid beetle Stenoria analis, that feed on the supply of nectar and pollen prepared by females bees in their nests.

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