Trochodendron
Encyclopedia
Trochodendron is a genus of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

s with one living species, Trochodendron aralioides
Trochodendron aralioides
Trochodendron aralioides, sometimes colloquially called "wheel tree", is a flowering plant and the sole living species in the genus Trochodendron, which also includes several extinct species. It was also often considered the sole species in the family Trochodendraceae, though botanists now include...

, and six extinct species known from the fossil record. It was often considered the sole genus in the family Trochodendraceae
Trochodendraceae
Trochodendraceae is a family of flowering plants with two living genera found in southeast Asia. The two living species share the feature of secondary xylem without vessels, which is quite rare in angiosperms...

, though botanists now also include the distinct genus Tetracentron
Tetracentron
Tetracentron is a genus of flowering plant, the sole living species being Tetracentron sinense. It was formerly considered the sole genus in the family Tetracentraceae, though modern botanists include it in the family Trochodendraceae together with the genus Trochodendron.It is native to southern...

in the family Trochodendraceae
Trochodendraceae
Trochodendraceae is a family of flowering plants with two living genera found in southeast Asia. The two living species share the feature of secondary xylem without vessels, which is quite rare in angiosperms...

.

Trochodendron is native to Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

, southern Korea
Korea
Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

, Taiwan
Taiwan
Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

 and the Ryukyu Islands
Ryukyu Islands
The , also known as the , is a chain of islands in the western Pacific, on the eastern limit of the East China Sea and to the southwest of the island of Kyushu in Japan. From about 1829 until the mid 20th century, they were alternately called Luchu, Loochoo, or Lewchew, akin to the Mandarin...

, with fossils know from North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 and Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

.

Trochodendron aralioides is an evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 or large shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 growing to 20 m tall. Trochodendron and Tetracentron lack vessel element
Vessel element
A vessel element is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are typically found in the angiosperms but absent from most gymnosperms such as the conifers....

s in their wood
Wood
Wood is a hard, fibrous tissue found in many trees. It has been used for hundreds of thousands of years for both fuel and as a construction material. It is an organic material, a natural composite of cellulose fibers embedded in a matrix of lignin which resists compression...

, a quite unusual feature in angiosperms
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

. This has long been considered a very primitive character, resulting in the classification of these two genera in a basal position in the angiosperms; however, genetic research by the Angiosperm Phylogeny Group
Angiosperm Phylogeny Group
The Angiosperm Phylogeny Group, or APG, refers to an informal international group of systematic botanists who came together to try to establish a consensus on the taxonomy of flowering plants that would reflect new knowledge about plant relationships discovered through phylogenetic studies., three...

 has shown it to be in a less basal position (early in the eudicots), suggesting the absence of vessel elements in these two genera is a secondarily evolved character, not a primitive one.

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