Daniel I. Arnon
Encyclopedia
Daniel Israel Arnon was a Polish-born
Poland
Poland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...

 American plant physiologist
Plant physiology
Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology , plant ecology , phytochemistry , cell biology, and molecular biology.Fundamental processes such as photosynthesis, respiration, plant nutrition,...

 whose research led to greater insights into the operation of photosynthesis
Photosynthesis
Photosynthesis is a chemical process that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight. Photosynthesis occurs in plants, algae, and many species of bacteria, but not in archaea. Photosynthetic organisms are called photoautotrophs, since they can...

 in plants. In 1973, he was awarded the National Medal of Science
National Medal of Science
The National Medal of Science is an honor bestowed by the President of the United States to individuals in science and engineering who have made important contributions to the advancement of knowledge in the fields of behavioral and social sciences, biology, chemistry, engineering, mathematics and...

 for "his fundamental research into the mechanism of green plant utilization of light to produce chemical energy and oxygen and for contributions to our understanding of plant nutrition."

Arnon was born on November 14, 1910, in Warsaw
Warsaw
Warsaw is the capital and largest city of Poland. It is located on the Vistula River, roughly from the Baltic Sea and from the Carpathian Mountains. Its population in 2010 was estimated at 1,716,855 residents with a greater metropolitan area of 2,631,902 residents, making Warsaw the 10th most...

. Summers spent on the family's farm helped foster Arnon's interest in agriculture. His father had lost the family's food wholesale business after World War I
World War I
World War I , which was predominantly called the World War or the Great War from its occurrence until 1939, and the First World War or World War I thereafter, was a major war centred in Europe that began on 28 July 1914 and lasted until 11 November 1918...

 and Arnon's readings of the works of Jack London
Jack London
John Griffith "Jack" London was an American author, journalist, and social activist. He was a pioneer in the then-burgeoning world of commercial magazine fiction and was one of the first fiction writers to obtain worldwide celebrity and a large fortune from his fiction alone...

 led him to save up his money to head to California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

. He enrolled in the University of California, Berkeley
University of California, Berkeley
The University of California, Berkeley , is a teaching and research university established in 1868 and located in Berkeley, California, USA...

 from Poland, and would spend his entire professional career at the school, until his retirement in 1978. He ultimately earned his Ph.D. in plant physiology at UC Berkeley under Dennis R. Hoagland and some of his earliest research focused on growing plants in nutrient-enriched water rather than in the soil. During World War II
World War II
World War II, or the Second World War , was a global conflict lasting from 1939 to 1945, involving most of the world's nations—including all of the great powers—eventually forming two opposing military alliances: the Allies and the Axis...

, Arnon served in the United States Army
United States Army
The United States Army is the main branch of the United States Armed Forces responsible for land-based military operations. It is the largest and oldest established branch of the U.S. military, and is one of seven U.S. uniformed services...

 in the Pacific Theater of Operations
Pacific Theater of Operations
The Pacific Theater of Operations was the World War II area of military activity in the Pacific Ocean and the countries bordering it, a geographic scope that reflected the operational and administrative command structures of the American forces during that period...

, where he used his prior experience with plant nutrition on Ponape Island, where there was no arable land
Arable land
In geography and agriculture, arable land is land that can be used for growing crops. It includes all land under temporary crops , temporary meadows for mowing or pasture, land under market and kitchen gardens and land temporarily fallow...

 available and he was able to grow food to feed the troops stationed there using gravel and nutrient-enriched water.

After returning from military service, Arnon performed research on chloroplast
Chloroplast
Chloroplasts are organelles found in plant cells and other eukaryotic organisms that conduct photosynthesis. Chloroplasts capture light energy to conserve free energy in the form of ATP and reduce NADP to NADPH through a complex set of processes called photosynthesis.Chloroplasts are green...

s and their role in the photosynthesis process. His work was able to demonstrate how energy from sunlight is used to form adenosine triphosphate
Adenosine triphosphate
Adenosine-5'-triphosphate is a multifunctional nucleoside triphosphate used in cells as a coenzyme. It is often called the "molecular unit of currency" of intracellular energy transfer. ATP transports chemical energy within cells for metabolism...

, the energy transport messenger within living cells, by adding a third phosphorus group to adenosine diphosphate
Adenosine diphosphate
Adenosine diphosphate, abbreviated ADP, is a nucleoside diphosphate. It is an ester of pyrophosphoric acid with the nucleoside adenosine. ADP consists of the pyrophosphate group, the pentose sugar ribose, and the nucleobase adenine....

. In 1954, Arnon reproduced the process in a laboratory, making him the first to successfully demonstrate the chemical function of photosynthesis, producing sugar and starch from inputs of carbon dioxide and water. He was elected a Fellow of the American Academy of Arts and Sciences
American Academy of Arts and Sciences
The American Academy of Arts and Sciences is an independent policy research center that conducts multidisciplinary studies of complex and emerging problems. The Academy’s elected members are leaders in the academic disciplines, the arts, business, and public affairs.James Bowdoin, John Adams, and...

 in 1962.

A resident of Kensington, California
Kensington, California
Kensington is an unincorporated community and census-designated place located in the East Bay, part of the San Francisco Bay Area, in Contra Costa County, California, United States. The population was 5,077 at the 2010 census.- Law and government :Kensington is an unincorporated area of Contra...

, Arnon died at age 84 on December 20, 1994, in Berkeley, California
Berkeley, California
Berkeley is a city on the east shore of the San Francisco Bay in Northern California, United States. Its neighbors to the south are the cities of Oakland and Emeryville. To the north is the city of Albany and the unincorporated community of Kensington...

, of complications resulting from cardiac arrest
Cardiac arrest
Cardiac arrest, is the cessation of normal circulation of the blood due to failure of the heart to contract effectively...

. He was survived by three daughters, two sons and eight grandchildren. His wife, the former Lucile Soule, died in 1986.
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