Cat behavior
Encyclopedia
Cat behavior generally refers to the behaviors and habits of domestic cats
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

, including body language
Body language
Body language is a form of non-verbal communication, which consists of body posture, gestures, facial expressions, and eye movements. Humans send and interpret such signals almost entirely subconsciously....

 and communication. Cat behavior may vary among breeds and individual cats. Many common behaviors include hunting
Hunting
Hunting is the practice of pursuing any living thing, usually wildlife, for food, recreation, or trade. In present-day use, the term refers to lawful hunting, as distinguished from poaching, which is the killing, trapping or capture of the hunted species contrary to applicable law...

 techniques and reactions to certain events as well as interactions with humans and other animals, such as dogs. Communication can vary greatly depending on a cat's temperament. In a family with multiple cats, social position can also affect behavior patterns with others. A cat's eating patterns can also vary depending on the owner's choice of food or eating times/quantities. In the case of a family having two or more cats, one cat may become dominant over the other cats.

Body language

Cats rely strongly on body language to communicate. Rubbing against an object, licking, and purring (sometimes) show affection. Purring can also mean that they are hurt or in pain.

Scent rubbing and spraying

This behavior
Behavior
Behavior or behaviour refers to the actions and mannerisms made by organisms, systems, or artificial entities in conjunction with its environment, which includes the other systems or organisms around as well as the physical environment...

 is used primarily to claim ownership of something, although unlike male cats, female cats do not usually spray. Some spayed females may spray to mimic males, but do not have enough scent to mark. Once male cats are neutered, the scent rubbing or spraying will, in most cases, decrease or stop. Some male cats continue to spray if not neutered early enough, though sometimes they continue to spray even if they were neutered early in life.

Kneading

Kneading is an activity common to all domestic cat
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

s whereby, when in a state of ease, they alternately push out and pull in their front paw
Paw
A paw is the soft foot of a mammal, generally a quadruped, that has claws or nails. A hard foot is called a hoof. Paws are used to pad feet for walking and increase friction.- Common characteristics :...

s, often alternating between right and left limbs. Some cats will actually appear to "nurse" or suck on clothing during kneading.

History

Multiple theories exist that explain why cats knead. Kneading may have an origin going back to cats' wild ancestors who had to tread down grass or foliage to make a temporary nest
Nest
A nest is a place of refuge to hold an animal's eggs or provide a place to live or raise offspring. They are usually made of some organic material such as twigs, grass, and leaves; or may simply be a depression in the ground, or a hole in a tree, rock or building...

 in which to rest. Alternatively, the behavior may be a remnant of a newborn's kneading of the mother's teat to stimulate milk excretion. Since most of the preferred "domestic traits" are neotenous or juvenile traits that persist in the adult, kneading may be a relict juvenile behavior that is not lost in modern adult domestic cats.

Kneading is often a precursor to sleep
Sleep
Sleep is a naturally recurring state characterized by reduced or absent consciousness, relatively suspended sensory activity, and inactivity of nearly all voluntary muscles. It is distinguished from quiet wakefulness by a decreased ability to react to stimuli, and is more easily reversible than...

ing.

Many cats purr
Purr
A purr is a sound made by all species of felids and is a part of cat communication. It varies between cats , and from species to species, but can be characterized as a tonal buzzing. Domestic cats purr in a frequency of 25 to 150 vibrations per second...

 while kneading. They also purr mostly when newborn, when feeding, or when trying to feed on their mother's teat. The common association between the two behaviors may corroborate the evidence in favor of the origin of kneading as a remnant instinct. Some experts consider kneading to "stimulate" the cat and make it feel good, in the same manner as a human stretching
Stretching
Stretching is a form of physical exercise in which a specific skeletal muscle is deliberately elongated, often by abduction from the torso, in order to improve the muscle's felt elasticity and reaffirm comfortable muscle tone. The result is a feeling of increased muscle control, flexibility and...

.

Action

The cat exerts firm downwards pressure
Pressure
Pressure is the force per unit area applied in a direction perpendicular to the surface of an object. Gauge pressure is the pressure relative to the local atmospheric or ambient pressure.- Definition :...

 with its paw, opening its toe
Toe
Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being digitigrade. Humans, and other animals that walk on the soles of their feet, are described as being plantigrade; unguligrade animals are those that walk on hooves at the tips of...

s to expose its claw
Claw
A claw is a curved, pointed appendage, found at the end of a toe or finger in most mammals, birds, and some reptiles. However, the word "claw" is also often used in reference to an invertebrate. Somewhat similar fine hooked structures are found in arthropods such as beetles and spiders, at the end...

s, then closes its claws as it lifts its paw. The process takes place with alternate paws at intervals of one to two seconds. They may do this while sitting on their owner's lap, which may prove painful if the cat is large or strong or has sharp claws (as the claws tend to dig into one's lap). Though cats will sit happily on a hard surface, they will only knead a soft or pliant surface, although some cats will reflexively "march" on hard surfaces instead of kneading them.

Courting

Cats, compared to many other mammals, have a unique courting style. The first step in courtship is the female coming into season, or "heat". Male cats will be able to smell a female cat in heat miles away and will therefore seek her out. Males will fight mercilessly for the right to be the first to mate with the female. After the dominant male has left, the less dominant males will then each mate with the female in turn. It is therefore possible that even if a male cat loses first breeding rights, he can still be the father. This is also the reason that a litter of kittens can have two or sometimes even three fathers.

Vocal calls

  • Purr
    Purr
    A purr is a sound made by all species of felids and is a part of cat communication. It varies between cats , and from species to species, but can be characterized as a tonal buzzing. Domestic cats purr in a frequency of 25 to 150 vibrations per second...

    ingPurring is often a sign of contentment. Some cats purr when they are in extreme pain, or in labour, simply to try to calm themselves down. Purring therefore can be a sign of pleasure or pain; usually it is the former. Scientists have not yet been able to discover how purring works, but it is suspected that it is caused by minute vibrations in the voice box.
  • Greeting
    Greeting
    Greeting is an act of communication in which human beings intentionally make their presence known to each other, to show attention to, and to suggest a type of relationship or social status between individuals or groups of people coming in contact with each other...

    A particular sort of vocalization, such as a low meow or chirp, possibly with simultaneous purring.
  • Distress
    Anxiety
    Anxiety is a psychological and physiological state characterized by somatic, emotional, cognitive, and behavioral components. The root meaning of the word anxiety is 'to vex or trouble'; in either presence or absence of psychological stress, anxiety can create feelings of fear, worry, uneasiness,...

    Mewing is a plea for help or attention often made by kittens. There are two basic types of this call, one more loud and frantic, the other more high-pitched. In older cats, it is more of a panicky repeated meow.
  • Attention
    Attention
    Attention is the cognitive process of paying attention to one aspect of the environment while ignoring others. Attention is one of the most intensely studied topics within psychology and cognitive neuroscience....

    Often simple meows and mews in both older cats and young kittens. A commanding meow is a command for attention, food, or to be let out.
  • Protest
    Protest
    A protest is an expression of objection, by words or by actions, to particular events, policies or situations. Protests can take many different forms, from individual statements to mass demonstrations...

    Whining meows.
  • Frustration
    Frustration
    This article concerns the field of psychology. The term frustration does, however, also concern physics. In this context, the term is treated in a different article, geometric frustration....

    A strong sigh or exhaled snort.
  • HappyA meow that starts low then goes up and comes back down.
  • Watching/interestCats will often "chatter" or "chirrup" on seeing something of interest. This is sometimes attributed to mimicking birdsong to attract prey or draw others' attention to it, but often birds are not present.

Panting

Unlike dogs, panting is an uncommon occurrence in cats. However, some cats can pant when stressed, such as a car ride. Most commonly, cats will pant in response to environmental changes, such as anxiety, fear, excitement, or heat. However, if panting is excessive or the cat appears in distress, it becomes important to identify the underlying cause, as panting may be a symptom of a more serious condition, such as a nasal blockage, heartworm
Heartworm
Heartworm is a parasitic roundworm that is spread from host to host through the bites of mosquitoes. The heartworm is a type of filaria, a small thread-like worm. The definitive host is the dog but it can also infect cats, wolves, coyotes, foxes and other animals, such as ferrets, sea lions and...

 disease, head trauma
Physical trauma
Trauma refers to "a body wound or shock produced by sudden physical injury, as from violence or accident." It can also be described as "a physical wound or injury, such as a fracture or blow." Major trauma can result in secondary complications such as circulatory shock, respiratory failure and death...

, or drug poison
Poison
In the context of biology, poisons are substances that can cause disturbances to organisms, usually by chemical reaction or other activity on the molecular scale, when a sufficient quantity is absorbed by an organism....

ing. Other problems, such as fatigue, weight loss, poor appetite, excessive drinking, vomiting, or diarrhea, may also be present. If the panting appears to be in response to heat or fear, the owner should remove the stimulation and continue to observe his or her pet. If panting continues, the owner should consult a veterinarian. In many cases, feline panting, especially if accompanied by other symptoms, such as cough
Cough
A cough is a sudden and often repetitively occurring reflex which helps to clear the large breathing passages from secretions, irritants, foreign particles and microbes...

ing or shallow breathing (dyspnea
Dyspnea
Dyspnea , shortness of breath , or air hunger, is the subjective symptom of breathlessness.It is a normal symptom of heavy exertion but becomes pathological if it occurs in unexpected situations...

), is considered to be abnormal, and treated as a medical emergency.

Righting reflex

The righting reflex is the ability for cats to land on their feet with little or no injury. They can do this more easily than other animals due to their flexible spine and floating collar bone. Cats also use vision and/or their vestibular apparatus to help tell which way to turn. They then can stretch themselves out and relax their muscles. Cats do not always land unharmed. They can break bones or die from excessive falls.

Food eating patterns

Cats are obligate carnivores, and can survive without vegetation. Felines in the wild will usually hunt smaller mammals regularly throughout the day to keep themselves nourished. Domestic cats, however, are used to a relaxed lifestyle and, therefore, will eat even smaller amounts, but more regularly. Many cats will find and chew small quantities of long grass but this is not for its nutritional value, it is a purely mechanical function. The eating of grass triggers a regurgitation reflex to help expel indigestible matter, like hairballs and the bones of prey.

Socialization

Kittens are naturally afraid of people at first, but if handled and well-cared for in the first 16 weeks, they will develop trust in the humans who care for them. To decrease the odds of a cat being unsocial or hostile towards humans, kittens should be socialized at an early age.

It is a challenge to socialize an adult feral cat
Feral cat
A feral cat is a descendant of a domesticated cat that has returned to the wild. It is distinguished from a stray cat, which is a pet cat that has been lost or abandoned, while feral cats are born in the wild; the offspring of a stray cat can be considered feral if born in the wild.In many parts of...

. Socialized adult feral cats tend to trust only those people they have learned over time can be trusted, and can be very fearful around strangers.

Cats can be extremely friendly companions. The strength of the cat–human bond is mainly correlated with how much consideration is given to the cat's feelings by his human companion. The formula for a successful relationship thus has much in common with human to human relationships.

Some people regard cats as sneaky, shy, or aloof animals. Cats have an inherent distrust for predator species such as humans, and often seek to minimize any contact with people they do not perceive as trustworthy. Feline shyness and aggression around people with cat social skills is often a result of lack of socialization, abuse or neglect. Cats relate to humans differently than more social animals, enjoying some time on their own each day as well as time with humans.

Cats have a strong "escape" instinct. Attempts to corner, capture or herd a cat can thus provoke powerful fear-based escape behavior. Socialization greatly reduces the number of humans that a cat will respond to in this way. Socialization
Socialization
Socialization is a term used by sociologists, social psychologists, anthropologists, political scientists and educationalists to refer to the process of inheriting and disseminating norms, customs and ideologies...

 is a process of learning that many humans can be trusted.

There is a widespread belief that relationships between dog
Dog
The domestic dog is a domesticated form of the gray wolf, a member of the Canidae family of the order Carnivora. The term is used for both feral and pet varieties. The dog may have been the first animal to be domesticated, and has been the most widely kept working, hunting, and companion animal in...

s and cats are problematic. However, both species can develop amicable relationships by reading each others' body language correctly. The animals can better read each others' language when they first encounter each other at a young age, due to the fact that they are learning to communicate simultaneously. The order of adoption may also cause significant differences in their relationship. Sometimes, the dog may be simply looking to play with the cat while the cat may take the approach a lot more seriously and lash out with its sharp claws, causing painful injury. Such an incident may cause an irreversible animosity between the cat and dog.
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