Black-tailed Tityra
Encyclopedia
The Black-tailed Tityra (Tityra cayana) is a medium-sized passerine
Passerine
A passerine is a bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds or, less accurately, as songbirds, the passerines form one of the most diverse terrestrial vertebrate orders: with over 5,000 identified species, it has roughly...

 bird of tropical South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

. The tityras have been placed in the cotinga
Cotinga
The cotingas are a large family of passerine bird species found in Central America and tropical South America. Cotingas are birds of forests or forest edges, which mostly eat fruit or insects and fruit. Comparatively little is known about this diverse group, although all have broad bills with...

 or the tyrant flycatcher
Tyrant flycatcher
The tyrant flycatchers are a family of passerine birds which occur throughout North and South America. They are considered the largest family of birds on Earth, with more than 400 species. They are the most diverse avian family in every country in the Americas, except for the United States and...

 families
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 (Cotingidae and Tyrannidae) by various authors. But the weight of evidence strongly suggest they and their closest relatives are better separated as Tityridae
Tityridae
Tityridae is family of suboscine passerine birds found in forest and woodland in the Neotropics. The approximately 30 species in this family were formerly spread over the families Tyrannidae, Pipridae and Cotingidae . As yet, no widely accepted common name exists for the family, although Tityras...

; the AOU
American Ornithologists' Union
The American Ornithologists' Union is an ornithological organization in the USA. Unlike the National Audubon Society, its members are primarily professional ornithologists rather than amateur birders...

 for example advocates this separation.

Description

This is a short-tailed robust bird with a long hook-tipped bill; like other tityras it has a peculiar vestigial ninth primary feather. The adult Black-tailed Tityra is about 20–22 cm (7.9–8.7 in) long and weighs about 60–70 g (2.1–2.5 oz). The male is dull white above and white below. The rectrices, the primary and secondary remiges and a cap extending to below the eyes are black; the tertiary remiges are silvery grey. Females have dark brown rather than black on wings and tail and some brown pattern on head, back and underside. In both sexes there is a patch of rosy-red bare skin around the eye, extending to the bill which is red-based with a black tip. The iris and feet are dark. , the juvenile plumage was unknown.

Voice

This species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 has a buzzing short call rendered as ed, rek, urd or wenk. A double beeza-buzza and triple weenk, weenk, weenk are also given. It more frequently calls in flight than when perched.

Taxonomy

Two subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 are recognized; they are sometimes treated as distinct species but widely intergrade where their ranges meet:
  • Black-tailed Tityra proper, Tityra cayana cayana (Linnaeus, 1766) – Northern South America
    South America
    South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

     east of the Andes
    Andes
    The Andes is the world's longest continental mountain range. It is a continual range of highlands along the western coast of South America. This range is about long, about to wide , and of an average height of about .Along its length, the Andes is split into several ranges, which are separated...

    , from Trinidad
    Trinidad
    Trinidad is the larger and more populous of the two major islands and numerous landforms which make up the island nation of Trinidad and Tobago. It is the southernmost island in the Caribbean and lies just off the northeastern coast of Venezuela. With an area of it is also the fifth largest in...

     across the Amazonas
    Amazon Basin
    The Amazon Basin is the part of South America drained by the Amazon River and its tributaries that drains an area of about , or roughly 40 percent of South America. The basin is located in the countries of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela...

     basin to Piauí
    Piauí
    Piauí is one of the states of Brazil, located in the northeastern part of the country.Piauí has the shortest coastline of any of the non-landlocked Brazilian states at 66 km , and the capital, Teresina, is the only state capital in the north east to be located inland...

     and Mato Grosso
    Mato Grosso
    Mato Grosso is one of the states of Brazil, the third largest in area, located in the western part of the country.Neighboring states are Rondônia, Amazonas, Pará, Tocantins, Goiás and Mato Grosso do Sul. It also borders Bolivia to the southwest...

     (Brazil
    Brazil
    Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

    ) and adjacent Bolivia
    Bolivia
    Bolivia officially known as Plurinational State of Bolivia , is a landlocked country in central South America. It is the poorest country in South America...

    .
Red bill base wide. Male greyish on back. Female with distinct brownish cap and little streaking on breast and back; throat white.
  • Brazilian Tityra, Tityra cayana braziliensis (Swainson, 1837) – Piauí, Mato Grosso and adjacent Bolivia southeastwards to Misiones
    Misiones Province
    Misiones is one of the 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamiсa region. It is surrounded by Paraguay to the northwest, Brazil to the north, east and south, and Corrientes Province of Argentina to the southwest.- History :The province was...

     and possibly Corrientes Province
    Corrientes Province
    Corrientes is a province in northeast Argentina, in the Mesopotamia region. It is surrounded by : Paraguay, the province of Misiones, Brazil, Uruguay, and the provinces of Entre Rios, Santa Fe and Chaco.-History:...

     (Argentina
    Argentina
    Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

    ).
Red bill base narrow. Male white on back. Female lacks distinct cap, streaked strongly on entire head, neck, back and underside.

Ecology

This bird is found as a year-round resident in forest edges, second growth and plantation shade trees in the pantanal
Pantanal
The Pantanal is a tropical wetland and one of the world's largest wetland of any kind. Most of it lies within the Brazilian state of Mato Grosso do Sul, but it extends into Mato Grosso and portions of Bolivia and Paraguay, sprawling over an area estimated at between and...

 and cerrado
Cerrado
The Cerrado, is a vast tropical savanna ecoregion of Brazil, particularly in the states of Gioas and Minas Gerais...

 as well as in terra firme and várzea forest, usually below 500 metres (1,640.4 ft) but occasionally as far up as 1100 metres (3,608.9 ft) ASL
Above mean sea level
The term above mean sea level refers to the elevation or altitude of any object, relative to the average sea level datum. AMSL is used extensively in radio by engineers to determine the coverage area a station will be able to reach...

. Black-tailed Tityras are most commonly seen in pairs, or, less frequently, single or in small groups; they are intolerant of other birds and will try to chase them away. They are often seen perched conspicuosly as they feed on medium-sized fruits. Food is gleaned from vegetation or picked off in mid-hover. While some large insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s are caught, these are mainly fed to young birds; adults are predominantly frugivores and locally important dispersers of such species as the Meliaceae
Meliaceae
The Meliaceae, or the Mahogany family, is a flowering plant family of mostly trees and shrubs in the order Sapindales....

 Cabralea canjerana. This species rarely attends mixed-species feeding flock
Mixed-species feeding flock
A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species, that join each other and move together while foraging...

s even when provisioning young, preferring to forage for insects on its own high up in the trees.

Its main breeding season appears to be from November to March across its range, but occasional nesting birds can be encountered almost year-round at least in some regions. For example, in the lowlands of Colombia and Ecuador, breeding activity has been reported in June and July also, suggesting either lack of a distinct breeding season and/or that two broods may be raised per year. The nest is several meters above ground in a tree hole, such as an old woodpecker
Woodpecker
Woodpeckers are near passerine birds of the order Piciformes. They are one subfamily in the family Picidae, which also includes the piculets and wrynecks. They are found worldwide and include about 180 species....

 nest or the crown of a dead palm tree. The brown-marked buff eggs are laid in a bed of dry leaves and some small twigs. Three eggs are considered likely, but exact clutch size is uncertain. Only the female incubates for almost three weeks until the young hatch, but both parents feed the chicks. Fledging is believed to take at least 3 weeks, perhaps as much as one month.

This species is one of the many hosts of the brood parasitic Shiny Cowbird
Shiny Cowbird
The Shiny Cowbird, Molothrus bonariensis, is a passerine bird in the New World family Icteridae. It breeds in most of South America apart from the most dense jungles, mountains and deserts , the coldest southernmost regions , and on Trinidad and Tobago...

 (Molothrus bonariensis). It is not uncommon across its large range and occurs in many national park
National park
A national park is a reserve of natural, semi-natural, or developed land that a sovereign state declares or owns. Although individual nations designate their own national parks differently A national park is a reserve of natural, semi-natural, or developed land that a sovereign state declares or...

s and other protected areas. Therefore it is considered a Species of Least Concern
Least Concern
Least Concern is an IUCN category assigned to extant taxon or lower taxa which have been evaluated but do not qualify for any other category. As such they do not qualify as threatened, Near Threatened, or Conservation Dependent...

 by the IUCN.

External links

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