Viola chamissoniana
Encyclopedia
Viola chamissoniana is a species of flowering plant in the violet family
Violaceae
Violaceae are a family of flowering plants consisting of about 800 species in 21 genera. It takes its name from the genus Viola, the violets and pansies.Older classifications such as the Cronquist system placed Violaceae in an order named after it, the Violales...

 known by the common name 'Olopu. It is endemic to Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

, where it is known from the islands of Kauai
Kauai
Kauai or Kauai, known as Tauai in the ancient Kaua'i dialect, is geologically the oldest of the main Hawaiian Islands. With an area of , it is the fourth largest of the main islands in the Hawaiian archipelago, and the 21st largest island in the United States. Known also as the "Garden Isle",...

, Oahu
Oahu
Oahu or Oahu , known as "The Gathering Place", is the third largest of the Hawaiian Islands and most populous of the islands in the U.S. state of Hawaii. The state capital Honolulu is located on the southeast coast...

, Molokai
Molokai
Molokai or Molokai is an island in the Hawaiian archipelago. It is 38 by 10 miles in size with a land area of , making it the fifth largest of the main Hawaiian Islands and the 27th largest island in the United States. It lies east of Oahu across the 25-mile wide Kaiwi Channel and north of...

, and Maui
Maui
The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

.

This plant is a woody shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 growing from a taproot
Taproot
A taproot is an enlarged, somewhat straight to tapering plant root that grows vertically downward. It forms a center from which other roots sprout laterally.Plants with taproots are difficult to transplant...

 and rooting where the lower stem comes in contact with the substrate. The leaves are up to 29 centimeters long by 9 wide and have toothed edges. The purple-tinged white flowers have petals measuring 1 to 2 centimeters long. The fruit is a capsule 1 to 2 centimeters in length which contains small dark-colored seeds.

There are three subspecies of this plant. Subspecies tracheliifolia is common. Subspecies chamissoniana is a federally listed endangered species
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

 of the United States which is limited to Oahu. There are three populations in the Waianae Range
Waianae Range
Waianae Range is the eroded remains of an ancient shield volcano that comprises the western half of the Hawaiian Island of Oahu...

 containing a total of about 443 individuals. Most of the plants grow on steep cliffs, but fences are in place to protect them from feral goat
Feral goat
The feral goat is the domestic goat when it has become established in the wild. Feral goats occur in Australia, New Zealand, Ireland, Great Britain, Hawaii, the Galapagos and in many other parts of the world...

s. Another threat is exotic plants
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

 such as Maui pamakani
Ageratina adenophora
Ageratina adenophora is a species of flowering plant in the daisy family known by many common names, including eupatory, sticky snakeroot, crofton weed, and Mexican devil.- Description :...

(Ageratina adenophora).
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