Rubus
Encyclopedia
Rubus is a large genus
of flowering plant
s in the rose
family, Rosaceae
, subfamily Rosoideae
. Raspberries
, blackberries
, and dewberries
are common, widely distributed members of the genus. Most of these plants have woody stems with prickles like roses; spines, bristles, and gland-tipped hairs are also common in the genus. The Rubus fruit
, sometimes called a bramble fruit, is an aggregate of drupelets.
The blackberries, as well as various other Rubus species with mounding or rambling growth habits, are often called bramble
s. However, this name is not used for those like the raspberry that grow as upright canes, or for trailing or prostrate species such as most dewberries, or various low-growing boreal, arctic, or alpine species.
The generic name means blackberry in Latin
and was derived from the word ruber, meaning "red".
The genus Rubus is believed to have existed since at least 23.7 to 36.6 million years ago.
The scientific study of brambles is known as "batology".
Examples of the hundreds, if not thousands, of species of Rubus include:
The genus also includes numerous hybrids, both natural and bred by man, such as the Loganberry
(Rubus × loganobaccus).
/dewberry
subgenus (Rubus), with polyploidy
, hybridization, and facultative apomixis
apparently all frequently occurring, making species
classification of the great variation in the subgenus one of the grand challenges of systematic botany.
Rubus species have a basic chromosome number of seven. Polyploidy from the diploid (14 chromosomes) to the tetradecaploid (98 chromosomes) is exhibited.
Some treatments have recognized dozens of species each for what other, comparably qualified botanists have considered single, more variable species. On the other hand, species in the other Rubus subgenera
(such as the raspberries) are generally distinct, or else involved in more routine one-or-a-few taxonomic debates, such as whether the European and American red raspberries are better treated as one species or two. (In this case, the two-species view is followed here, with Rubus idaeus and R. strigosus both recognized; if these species are combined, then the older name R. idaeus has priority for the broader species.)
Molecular data have backed up classifications based on geography and chromosome number, but following morphological
data such as the structure of the leaves and stems do not appear to produce a phylogenetic classification.
The classification presented below recognizes 13 subgenera
within Rubus, with the largest subgenus (Rubus) in turn divided into 12 sections
. Representative examples are presented, but there are many more species not mentioned here.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
s in the rose
Rose
A rose is a woody perennial of the genus Rosa, within the family Rosaceae. There are over 100 species. They form a group of erect shrubs, and climbing or trailing plants, with stems that are often armed with sharp prickles. Flowers are large and showy, in colours ranging from white through yellows...
family, Rosaceae
Rosaceae
Rosaceae are a medium-sized family of flowering plants, including about 2830 species in 95 genera. The name is derived from the type genus Rosa. Among the largest genera are Alchemilla , Sorbus , Crataegus , Cotoneaster , and Rubus...
, subfamily Rosoideae
Rosoideae
The rose subfamily Rosoideae consists of more than 850 species, including many shrubs, perennial herbs, and fruit plants such as strawberries and brambles...
. Raspberries
Raspberry
The raspberry or hindberry is the edible fruit of a multitude of plant species in the genus Rubus, most of which are in the subgenus Idaeobatus; the name also applies to these plants themselves...
, blackberries
Blackberry
The blackberry is an edible fruit produced by any of several species in the Rubus genus of the Rosaceae family. The fruit is not a true berry; botanically it is termed an aggregate fruit, composed of small drupelets. The plants typically have biennial canes and perennial roots. Blackberries and...
, and dewberries
Dewberry
The dewberries are a group of species in the genus Rubus, section Rubus, closely related to the blackberries. They are small trailing brambles with berries reminiscent of the raspberry, but are usually purple to black instead of red.Dewberries are common throughout most of the Northern Hemisphere,...
are common, widely distributed members of the genus. Most of these plants have woody stems with prickles like roses; spines, bristles, and gland-tipped hairs are also common in the genus. The Rubus fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...
, sometimes called a bramble fruit, is an aggregate of drupelets.
The blackberries, as well as various other Rubus species with mounding or rambling growth habits, are often called bramble
Bramble
Brambles are thorny plants of the genus Rubus, in the rose family . Bramble fruit is the fruit of any such plant, including the blackberry and raspberry. The word comes from Germanic *bram-bezi, whence also German Brombeere , Dutch Braam and French framboise...
s. However, this name is not used for those like the raspberry that grow as upright canes, or for trailing or prostrate species such as most dewberries, or various low-growing boreal, arctic, or alpine species.
The generic name means blackberry in Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...
and was derived from the word ruber, meaning "red".
The genus Rubus is believed to have existed since at least 23.7 to 36.6 million years ago.
The scientific study of brambles is known as "batology".
Examples of the hundreds, if not thousands, of species of Rubus include:
- Rubus allegheniensis – Allegheny Blackberry
- Rubus arcticus – Arctic Raspberry
- Rubus armeniacusRubus armeniacusRubus armeniacus, Armenian Blackberry or Himalayan Blackberry, is a species of Rubus in the blackberry group Rubus subgenus Rubus series Discolores Focke. It is native to Armenia in southwest Asia, and widely naturalised elsewhere...
– Himalayan Blackberry - Rubus caesius – European Dewberry
- Rubus canadensis – Canadian Blackberry
- Rubus chamaemorus – Cloudberry
- Rubus coreanusRubus coreanusRubus coreanus is a species of raspberry native to Korea, Japan, and China. It produces edible berries that are fermented into bokbunja ju, a Korean fruit wine...
- Bokbunja - Rubus cuneifolius – Sand Blackberry
- Rubus fruticosus agg. – Blackberry
- Rubus glaucifoliusRubus glaucifoliusRubus glaucifolius is a species of wild raspberry known by the common name San Diego raspberry. It is native to Oregon and California, where it grows in mountain forests. It is a tangling shrub with very slender, lightly prickly stem spreading and branching outward. The leaves are each made up of...
- San Diego Raspberry - Rubus hayata-koidzumii (R. calycinoides) – Creeping Raspberry
- Rubus hochstetterorum - Azorean Blackberry
- Rubus idaeusRubus idaeusRubus idaeus is a red-fruited species of Rubus native to Europe and northern Asia and commonly cultivated in other temperate regions. A closely related plant in North America, sometimes regarded as the variety Rubus idaeus var...
– European Red Raspberry - Rubus laciniatusRubus laciniatusRubus laciniatus, the Cutleaf Evergreen Blackberry or Evergreen Blackberry, is a species of Rubus native to Europe. it is an introduced species in North America...
- Cutleaf Evergreen Blackberry - Rubus lasiococcusRubus lasiococcusRubus lasiococcus is a species of wild blackberry known by the common names roughfruit berry and dwarf bramble. It is native to western North America from British Columbia to northern California, where it grows in mountain forests...
- Roughfruit Berry - Rubus leucodermisRubus leucodermisRubus leucodermis is a species of Rubus native to western North America, from British Columbia, Canada south to California, New Mexico and Mexico...
– Whitebark Raspberry or Western Raspberry - Rubus occidentalisRubus occidentalisRubus occidentalis is a species of Rubus native to eastern North America. Its common name black raspberry is shared with the closely related western American species Rubus leucodermis...
– Black Raspberry - Rubus odoratusRubus odoratusRubus odoratus is a species of Rubus, native to eastern North America, from Nova Scotia west to Ontario, and south to Georgia and Alabama....
– Flowering Raspberry - Rubus parviflorus – Thimbleberry
- Rubus parvifoliusRubus parvifoliusRubus parvifolius, native raspberry, or small-leaf bramble, is a scrambling shrub occurring in heathland and eucalyptus woodland native to eastern Australia....
- Small-leaf Bramble (Australia) - Rubus pensilvanicusRubus pensilvanicusRubus pensilvanicus, known commonly as Pennsylvania Blackberry, is a prickly bramble native to eastern North America. The canes are green at first but then turning dark red, usually ridged, with copious straight prickles. The leaves are compound, usually bearing 5 or 7 leaflets. The flowers are...
– Pennsylvania Blackberry - Rubus phoenicolasiusRubus phoenicolasiusRubus phoenicolasius is a species of raspberry native to northern China, Japan, and Korea. The species was introduced to Europe and North America as an ornamental plant and for its potential in breeding hybrid raspberries...
– Wineberry - Rubus saxatilisRubus saxatilisRubus saxatilis or Stone Bramble is a species of bramble found in Europe and temperate Asia to Japan. The green stems are 20–60 cm tall and covered with minute needle-like prickles, and leaves are usually compound with three leaflets...
– Stone Bramble - Rubus spectabilis – Salmonberry
- Rubus strigosusRubus strigosusRubus strigosus, the American Red Raspberry or American Raspberry, is a species of Rubus native to much of North America. It has often been treated as a variety or subspecies of the closely related Eurasian Rubus idaeus , but currently is more commonly treated as a distinct...
– American Red Raspberry - Rubus trifidusRubus trifidusRubus trifidus is a species from Japan....
- Japanese Blackberry - Rubus ursinusRubus ursinusRubus ursinus is a species of blackberry or dewberry known by the common names California blackberry/dewberry, Douglas berry, and Pacific blackberry/dewberry. It is native to western North America. This is a wide, spreading shrub or vine-bearing bush with prickly branches that can tip layer to...
– Trailing Blackberry
The genus also includes numerous hybrids, both natural and bred by man, such as the Loganberry
Loganberry
The loganberry is an hexaploid hybrid produced from crossing an octaploid blackberry and a diploid red raspberry. The plant and the fruit resemble the blackberry more than the raspberry, but the fruit colour is a dark red, rather than black...
(Rubus × loganobaccus).
- See also: List of Lepidoptera that feed on Rubus
Scientific classification
The genus Rubus is a very complex one, particularly the blackberryBlackberry
The blackberry is an edible fruit produced by any of several species in the Rubus genus of the Rosaceae family. The fruit is not a true berry; botanically it is termed an aggregate fruit, composed of small drupelets. The plants typically have biennial canes and perennial roots. Blackberries and...
/dewberry
Dewberry
The dewberries are a group of species in the genus Rubus, section Rubus, closely related to the blackberries. They are small trailing brambles with berries reminiscent of the raspberry, but are usually purple to black instead of red.Dewberries are common throughout most of the Northern Hemisphere,...
subgenus (Rubus), with polyploidy
Polyploidy
Polyploid is a term used to describe cells and organisms containing more than two paired sets of chromosomes. Most eukaryotic species are diploid, meaning they have two sets of chromosomes — one set inherited from each parent. However polyploidy is found in some organisms and is especially common...
, hybridization, and facultative apomixis
Apomixis
In botany, apomixis was defined by Winkler as replacement of the normal sexual reproduction by asexual reproduction, without fertilization. This definition notably does not mention meiosis...
apparently all frequently occurring, making species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...
classification of the great variation in the subgenus one of the grand challenges of systematic botany.
Rubus species have a basic chromosome number of seven. Polyploidy from the diploid (14 chromosomes) to the tetradecaploid (98 chromosomes) is exhibited.
Some treatments have recognized dozens of species each for what other, comparably qualified botanists have considered single, more variable species. On the other hand, species in the other Rubus subgenera
Subgenus
In biology, a subgenus is a taxonomic rank directly below genus.In zoology, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the generic name and the specific epithet: e.g. the Tiger Cowry of the Indo-Pacific, Cypraea tigris Linnaeus, which...
(such as the raspberries) are generally distinct, or else involved in more routine one-or-a-few taxonomic debates, such as whether the European and American red raspberries are better treated as one species or two. (In this case, the two-species view is followed here, with Rubus idaeus and R. strigosus both recognized; if these species are combined, then the older name R. idaeus has priority for the broader species.)
Molecular data have backed up classifications based on geography and chromosome number, but following morphological
Plant morphology
Plant morphology or phytomorphology is the study of the physical form and external structure of plants. This is usually considered distinct from plant anatomy, which is the study of the internal structure of plants, especially at the microscopic level...
data such as the structure of the leaves and stems do not appear to produce a phylogenetic classification.
The classification presented below recognizes 13 subgenera
Subgenus
In biology, a subgenus is a taxonomic rank directly below genus.In zoology, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the generic name and the specific epithet: e.g. the Tiger Cowry of the Indo-Pacific, Cypraea tigris Linnaeus, which...
within Rubus, with the largest subgenus (Rubus) in turn divided into 12 sections
Section (botany)
In botany, a section is a taxonomic rank below the genus, but above the species. The subgenus, if present, is higher than the section, and the rank of series, if present, is below the section. Sections are typically used to help organise very large genera, which may have hundreds of species...
. Representative examples are presented, but there are many more species not mentioned here.
|
|
External links
- Rubus at the Western Kentucky UniversityWestern Kentucky UniversityWestern Kentucky University is a public university in Bowling Green, Kentucky, USA. It was formally founded by the Commonwealth of Kentucky in 1906, though its roots reach back a quarter-century earlier....