Posidonia Shale
Encyclopedia
The Posidonia Shale is a Lower Jurassic
Early Jurassic
The Early Jurassic epoch is the earliest of three epochs of the Jurassic period...

 geological formation famous for its exceptionally well-preserved complete skeletons of fossil marine fish and reptiles. The Posidonienschiefer as German paleontologists call it, takes its name from the ubiquitous fossils of Posidonia bronni that characterize its fauna. The formation comprises finely laminated layers of bituminous shale
Bituminous shale
Bituminous shale is an argillaceous shale impregnated with bitumen, often accompanying coal. It may contain kerogen; in that case it is classified as oil shale. When carbonaceous material is present in large amounts, bituminous shale grades into bituminous coal...

s formed of fine-grained sediments intercalated with bituminous limestones and crops out in a number of locations in southwestern Germany, although most famously near the village of Holzmaden
Holzmaden
Holzmaden is a town in Baden-Württemberg, Germany that lies between Stuttgart and Ulm.The ground in and around the city contains rich layers of well preserved fossils of the Jurassic period. The fossils are found in the 160 million year old Posidonia Shale and displayed in the local Museum Hauff...

. The Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

an bituminous shales deposited on a sea floor during the Early Toarcian
Toarcian
The Toarcian is, in the ICS' geologic timescale, an age or stage in the Early or Lower Jurassic. It spans the time between 183.0 Ma and 175.6 Ma...

 in the ancient Tethys Ocean
Tethys Ocean
The Tethys Ocean was an ocean that existed between the continents of Gondwana and Laurasia during the Mesozoic era before the opening of the Indian Ocean.-Modern theory:...

 are described as being deposited in an anoxic, or oxygen depleted, deep water environment, although the details of the depositional environment are the subject of debate by researchers of the formation.

In addition to their Posidonia bronni, the shales contain some spectacularly detailed fossils of other Jurassic sea creatures—ichthyosaur
Ichthyosaur
Ichthyosaurs were giant marine reptiles that resembled fish and dolphins...

s and plesiosaur
Plesiosaur
Plesiosauroidea is an extinct clade of carnivorous plesiosaur marine reptiles. Plesiosauroids, are known from the Jurassic and Cretaceous Periods...

s, spiral-shelled ammonite
Ammonite
Ammonite, as a zoological or paleontological term, refers to any member of the Ammonoidea an extinct subclass within the Molluscan class Cephalopoda which are more closely related to living coleoids Ammonite, as a zoological or paleontological term, refers to any member of the Ammonoidea an extinct...

s and crinoid
Crinoid
Crinoids are marine animals that make up the class Crinoidea of the echinoderms . Crinoidea comes from the Greek word krinon, "a lily", and eidos, "form". They live both in shallow water and in depths as great as 6,000 meters. Sea lilies refer to the crinoids which, in their adult form, are...

s, or sea-lilies.

Plesiosaurs

Plesiosaur
Plesiosaur
Plesiosauroidea is an extinct clade of carnivorous plesiosaur marine reptiles. Plesiosauroids, are known from the Jurassic and Cretaceous Periods...

s of the Posidonia Shale
Taxa Presence Notes Images

Genus:
  • Meyerasaurus
    Meyerasaurus
    Meyerasaurus is an extinct genus of rhomaleosaurid known from Holzmaden, Baden-Württemberg of southeastern Germany.-Description:Meyerasaurus is known from the holotype SMNS 12478, articulated and complete skeleton which preserved the skull, exposed in ventral view. The skull has a length of ,...

  1. Meyerasaurus victor
  • Found near the town of Holzmaden (Baden-Württemberg), Germany.


  • Pterosaurs

    Pterosaur
    Pterosaur
    Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

    s of the Posidonia Shale
    Taxa Presence Notes Images

    Genus:
    • Campylognathoides
      Campylognathoides
      Campylognathoides was a genus of "rhamphorhynchoid" pterosaur, discovered in the Württemberg Lias deposits, the first specimen consisting of wing fragments...

    1. C. liasicus
    2. C. zitteli
  • Found near the town of Holzmaden (Baden-Württemberg), Germany.
  • Found near the town of Holzmaden (Baden-Württemberg), Germany.


  • Genus:
    • Dorygnathus
      Dorygnathus
      Dorygnathus was a genus of pterosaur that lived in Europe during the Early Jurassic period, 180 million years ago when shallow seas flooded much of the continent. It had a short 1.5 meter wingspan, and a relatively small triangular sternum, which is where its flight muscles attached. Its skull...

    1. D. banthensis
  • Found near Banz Abbey in Bavaria, as well as Holzmaden (Baden-Württemberg),

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