Pomatiidae
Encyclopedia
The family Pomatiidae is a taxonomic family of small operculate
Operculum (gastropod)
The operculum, meaning little lid, is a corneous or calcareous anatomical structure which exists in many groups of sea snails and freshwater snails, and also in a few groups of land snails...

 land snail
Land snail
A land snail is any of the many species of snail that live on land, as opposed to those that live in salt water and fresh water. Land snails are terrestrial gastropod mollusks that have shells, It is not always an easy matter to say which species are terrestrial, because some are more or less...

s, terrestrial
Terrestrial animal
Terrestrial animals are animals that live predominantly or entirely on land , as compared with aquatic animals, which live predominantly or entirely in the water , or amphibians, which rely on a combination of aquatic and terrestrial habitats...

 gastropod mollusks that can be found over the warmer parts of the Old World
Old World
The Old World consists of those parts of the world known to classical antiquity and the European Middle Ages. It is used in the context of, and contrast with, the "New World" ....

. In the older literature, this family is designated as Pomatiasidae.

This family is a lineage closely related to the Littorinidae
Littorinidae
Littorinidae is a taxonomic family of over 200 species of sea snails, marine gastropod molluscs in the clade Littorinimorpha, commonly known as periwinkles and found world-wide.-Names:...

 (periwinkles) common in coast
Coast
A coastline or seashore is the area where land meets the sea or ocean. A precise line that can be called a coastline cannot be determined due to the dynamic nature of tides. The term "coastal zone" can be used instead, which is a spatial zone where interaction of the sea and land processes occurs...

al habitat. They have adapted to terrestrial life and are thus sometimes humorously called "winkles-come-ashore".

They are defined by a chalky operculum at the rear end of the body, the shape of their thick shell and their mouth and a characteristic spiral sculpture. The sexes are separate and can sometimes be recognised because the female shell is slightly larger than the male shell.

Taxonomy

According to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
The taxonomy of the Gastropoda as it was revised by Philippe Bouchet and Jean-Pierre Rocroi is currently the most up-to-date overall system for classifying gastropod mollusks...

, the family Pomatiidae consists of two subfamilies:
  • subfamily Pomatiinae Newton, 1891 (1828) - synonyms: Cyclostomatidae Menke, 1828; Cyclotopsinae Kobelt & Möllendorff, 1898; Ericiidae Wenz, 1915
  • subfamily Annulariinae Henderson & Bartsch, 1920
    • tribe Annulariini Henderson & Bartsch, 1920 - synonyms: Licininae Gray, 1857; Chondropomatinae Henderson & Bartsch, 1920
    • tribe Adamsiellini Henderson & Bartsch, 1920
    • tribe Choanopomatini Thiele, 1929
    • tribe Cistulopsini H. B. Baker, 1924 - synonym: Cistulinae L. Pfeiffer, 1858
    • tribe Rhytidopomatini Henderson & Bartsch, 1920

Genera

Genera within the family Pomatiidae include:

subfamily Pomatiinae
  • Cyclostoma Lamarck, 1899
  • Cyclotopsis Blanford, 1864
  • Ericia Partiot, 1848
  • Georgia Bourguignat, 1882
  • Guillainia Crosse, 1884
  • Leonia Gray, 1850
  • Lithidion Gray, 1850
  • Otopoma Gray, 1850
  • Pomatias
    Pomatias
    Pomatias is a genus of small operculate land snails, terrestrial gastropod mollusks gastropod in the family Pomatiidae.- Species :Species within the genus Pomatias include:* Pomatias elegans - Round-mouthed Snail...

    Studer, 1789 - type genus of the family Pomatiidae
  • Tropidophora
    Tropidophora
    Tropidophora is a genus of land snails, terrestrial gastropod mollusks in the family Pomatiidae.- Distribution :These are large, split-sole, operculate snails found in Tanzania, Madagascar, the Mascarenes, the Comoros and South Africa....

    Troschel, 1847
  • Tudorella
    Tudorella
    Tudorella is a genus of land snails, terrestrial gastropod mollusks in the family Pomatiidae.- Distribution :This genus of land snails inhabits the coasts of the western Mediterranean sea.- Species :Species in the genus Tudorella include:...

    Fischer, 1885


subfamily Annulariinae

tribe Annulariini
  • Annularia Schumacher, 1817 - type genus of the subfamily Annulariinae
  • Chondropoma
    Chondropoma
    Chondropoma is a genus of small operculate land snails, terrestrial gastropod mollusks gastropod in the family Pomatiidae.- Distribution :Distribution of Chondropoma include Cuba, Nicaragua, ...63 species of Chondropoma lives in Cuba....

    L. Pfeiffer, 1847
  • Licina Gray, 1847


tribe Adamsiellini
  • Adamsiella L. Pfeiffer, 1851 - type genus of the tribe Adamsiellini


tribe Choanopomatini
  • Choanopoma L. Pfeiffer, 1847 - type genus of the tribe Choanopomatini


tribe Cistulopsini
  • Cistula Gray, 1850
  • Cistulops H. B. Baker, 1924 - type genus of the tribe Cistulopsini


tribe Rhytidopomatini
  • Rhytidopoma
    Rhytidopoma
    Rhytidopoma is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.Rhytidopoma is the type genus of the tribe Rhytidopomatini.- Species :Species within the genus Rhytidopoma include:...

    Sykes, 1901 - type genus of the tribe Rhytidopomatini


to be sorted to subfamilies:
  • Abbottella
  • Annularisca
    Annularisca
    Annularisca is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Annularisca include:* Annularisca aberrans Torre & Barsch, 1941...

    Henderson & Bartsch, 1920
  • Annularita
    Annularita
    Annularita is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Annularita include:* Annularita majuscula...

    Henderson & Bartsch, 1920
  • Annularodes
    Annularodes
    Annularodes is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Annularodes include:* Annularodes boqueronensis...

    Henderson & Bartsch, 1920
  • Annularops
    Annularops
    Annularops is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Annularops include:* Annularops attenuata...

    Henderson & Bartsch, 1920
  • Bermudezia
    Bermudezia
    Bermudezia is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Bermudezia include:* Bermudezia bermudezi...

    Torre & Bartsch, 1941
  • Blaesospira
    Blaesospira
    Blaesospira is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Blaesospira include:* Blaesospira echinus - type species...

    Crosse, 1890
  • Chondropomatus
    Chondropomatus
    Chondropomatus is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Chondropomatus include:* Chondropomatus latum...

    Henderson & Bartsch, 1920
  • Chondropometes
    Chondropometes
    Chondropometes is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Chondropometes include:* Chondropometes bellisimum Torre & Bartsch, 1938...

    Henderson & Bartsch, 1920
  • Chondrothyra
    Chondrothyra
    Chondrothyra is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Chondrothyra include:* Chondrothyra affinis...

    Henderson & Bartsch, 1920
  • Chondrothyrella
    Chondrothyrella
    Chondrothyrella is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Chondrothyrella include:* Chondrothyrella assimilis...

    Torre & Bartsch, 1938
  • Chondrothyrium
    Chondrothyrium
    Chondrothyrium is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Chondrothyrium include:* Chondrothyrium alcaldei Jaume & Sánchez de Fuentes, 1943...

    Henderson & Bartsch, 1920
  • Cubadamsiella
    Cubadamsiella
    Cubadamsiella is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Cubadamsiella include:* Cubadamsiella beneitoi Fernández-Garcés, Espinosa & Ortea, 2003...

    Torre & Bartsch, 1941
  • Dallsiphona
    Dallsiphona
    Dallsiphona is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Dallsiphona include:* Dallsiphona dalli...

    Torre & Bartsch, 1941
  • Diploma - an orth. error for Diplopoma
    Diplopoma
    Diplopoma is a genus of operculate land snails, terrestrial gastropod mollusks gastropod in the family Pomatiidae.-Species:Species within the genus Diplopoma include:* Diplopoma crenulatum...

    ? (original research)
    • Diploma arangoi (Jaume, 1984)
    • Diploma architectonica (Gundlach in Pfeiffer, 1859)
    • Diploma songoensis (Torre & Bartsch, 1941)
    • Diploma torrei (Ramsden, 1915)
    • Diploma varonai (Jaume, 1984)
    • Diploma zayasi (Jaume, 1984)
  • Diplopoma
    Diplopoma
    Diplopoma is a genus of operculate land snails, terrestrial gastropod mollusks gastropod in the family Pomatiidae.-Species:Species within the genus Diplopoma include:* Diplopoma crenulatum...

    L. Pfeiffer, 1859
  • Eutudora
    Eutudora
    Eutudora is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Eutudora include:* Eutudora agassizi...

    Henderson & Bartsch, 1920
  • Eutudorops
    Eutudorops
    Eutudorops is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Eutudorops include:* Eutudorops complanata...

    Henderson & Bartsch, 1920
  • Guajaibona
    Guajaibona
    Guajaibona is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Guajaibona include:* Guajaibona petrei...

    Torre & Bartsch, 1941
  • Gundlachtudora
    Gundlachtudora
    Gundlachtudora is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Gundlachtudora include:* Gundlachtudora decolorata...

    Torre & Bartsch, 1941
  • Hendersonina
    Hendersonina
    Hendersonina is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Hendersonina include:* Hendersonina bermudezi Torre & Bartsch, 1938...

    Torre & Bartsch, 1938
  • Jaumeia
    Jaumeia
    Jaumeia is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Jaumeia include:* Jaumeia notata...

    Aguayo, 1943
  • Juannularia
    Juannularia
    Juannularia is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Juannularia include:* Juannularia arguta...

    Torre & Bartsch, 1941
  • Limadora
    Limadora
    Limadora is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Limadora include:* Limadora garciana...

    Torre & Bartsch, 1941
  • Limadorex
    Limadorex
    Limadorex is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Limadorex include:* Limadorex limonensis...

    Torre & Bartsch, 1941
  • Meganipha Thompson, 1978
  • Opisthocoelicum
    Opisthocoelicum
    Opisthocoelicum is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Opisthocoelicum include:* Opisthocoelicum dubium Sánchez Roig, 1949...

    Torre & Bartsch, 1941
  • Opisthosiphon
    Opisthosiphon
    Opisthosiphon is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Description :The genus Opisthosiphon has been described by American malacologist William Healey Dall in 1905...

    Dall, 1905
  • Parachondria Dall, 1905 - originally described as a subgenus
  • Ramsdenia
    Ramsdenia
    Ramsdenia is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Ramsdenia include:* Ramsdenia bufo...

    Preston, 1913
  • Rhytidothyra
    Rhytidothyra
    Rhytidothyra is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Rhytidothyra include:* Rhytidothyra bilabiata...

    Henderson & Bartsch, 1920
  • Subannularia
    Subannularia
    Subannularia is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Subannularia include:* Subannularia jeannereti...

    Torre & Bartsch, 1941
  • Torrella
    Torrella (gastropod)
    Torrella is a genus of operculate land snails, terrestrial gastropod mollusks in the family Pomatiidae.- Distribution :The distribution of the genus Torrella includes Cuba.- Species :Species in the genus Torrella include:...

    Henderson & Bartsch, 1920
  • Torrilla (spelling error?)
    • Torrilla emmae Jaume & Sánchez de Fuentes, 1943
    • Torrilla torreiana (Gundlach in Arango, 1878)
    • Torrilla trinidadensis Torre & Bartsch, 1941
  • Troschelvindex
    Troschelvindex
    Troschelvindex is a genus of land snails with an operculum: these terrestrial gastropod mollusks are in the family Pomatiidae.- Species :Species within the genus Troschelvindex include:* Troschelvindex agrestis...

    Baker, 1924
  • Tudora Baird, 1850
  • Tudorina
    Tudorina
    Tudorina is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Tudorina include:* Tudorina rangelina...

    Torre & Bartsch, 1941
  • Wrightudora
    Wrightudora
    Wrightudora is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Wrightudora include:* Wrightudora aguayoi...

    Torre & Bartsch, 1941
  • Xenopoma
    Xenopoma
    Xenopoma is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Pomatiidae.- Species :Species within the genus Xenopoma include:* Xenopoma aguayoi Torre & Bartsch, 1941...

    Crosse, 1890
  • Xenopoides
    • Xenopoides delicatulum Torre & Bartsch, 1941

Further reading

  • Bartsch P.
    Paul Bartsch
    Paul Bartsch was an American malacologist and carcinologist.Paul Bartsch emigrated to the U.S.A in 1880. He graduated from the University of Iowa with a B.S. in 1896, and M.S. in 1899, and PhD in 1905...

     (1946). "The operculate land mollusks of the family Annulariidae of the Island of Hispaniola
    Hispaniola
    Hispaniola is a major island in the Caribbean, containing the two sovereign states of the Dominican Republic and Haiti. The island is located between the islands of Cuba to the west and Puerto Rico to the east, within the hurricane belt...

     and the Bahama Archipelago". United States National Museum Bulletin No. 192, 264 pp., 38 pls.
  • Pfenninger M., Véla E., Jesse R., Arantzazu Elejalde M., Liberto F., Magnin F. & Martínez-Ortí A. (2009). "Temporal speciation pattern in the western Mediterranean genus Tudorella P. Fischer, 1885 (Gastropoda, Pomatiidae) supports the Tyrrhenian vicariance hypothesis". Molecular Phylogenetics and Evolution
    Molecular Phylogenetics and Evolution
    Molecular Phylogenetics and Evolution is a peer-reviewed scientific journal of evolutionary biology and phylogenetics. The journal is edited by D.E. Wildman.-Indexing:The journal is indexed in:*EMBiology*Journal Citation Reports*Scopus*Web of Science...

    54(2): 427-436. , doi:10.1016/j.ympev.2009.09.024.
  • Watters G. T. (2006). The Caribbean Land Snail Family Annulariidae: A revision of the higher taxa and a catalog of the species. 1-557, figs 1-10, map 1-56, Appendix A: 1-3. Backhuys Publishers, Leiden.
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