Littoraria
Encyclopedia
Littoraria is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of sea snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

s, marine
Marine (ocean)
Marine is an umbrella term. As an adjective it is usually applicable to things relating to the sea or ocean, such as marine biology, marine ecology and marine geology...

 gastropod mollusks in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Littorinidae
Littorinidae
Littorinidae is a taxonomic family of over 200 species of sea snails, marine gastropod molluscs in the clade Littorinimorpha, commonly known as periwinkles and found world-wide.-Names:...

, the winkles or periwinkles.

There are more than fifty species in this genus of which more than 20 species arebelieved to besynonyms of Littorina scabra, a very variable species.

Many of the species in this genus occur in the Indo-West Pacific region and in the Tropical Eastern Pacific, where they are found in large numbers on the trunks, trees and prop roots of tropical mangrove
Mangrove
Mangroves are various kinds of trees up to medium height and shrubs that grow in saline coastal sediment habitats in the tropics and subtropics – mainly between latitudes N and S...

 forests a few metres above high tide level. These snails feed on the thin film of algae
Algae
Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

, epiphyte
Epiphyte
An epiphyte is a plant that grows upon another plant non-parasitically or sometimes upon some other object , derives its moisture and nutrients from the air and rain and sometimes from debris accumulating around it, and is found in the temperate zone and in the...

s, fungi
Fungus
A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...

, diatom
Diatom
Diatoms are a major group of algae, and are one of the most common types of phytoplankton. Most diatoms are unicellular, although they can exist as colonies in the shape of filaments or ribbons , fans , zigzags , or stellate colonies . Diatoms are producers within the food chain...

s and leaf epidermis
Epidermis (botany)
The epidermis is a single-layered group of cells that covers plants' leaves, flowers, roots and stems. It forms a boundary between the plant and the external environment. The epidermis serves several functions, it protects against water loss, regulates gas exchange, secretes metabolic compounds,...

 of these mangroves. The species living on higher levels of the trees have thinner shells, and are more variable in shell colour.

Within this genus, Littoraria aberrans is the only ovoviviparous species with an intracapsular metamorphosis.

Species

Species in the genus Littoraria include:
  • Littoraria aberrans
    Littoraria aberrans
    Littoraria aberrans is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria ahenea Reid, 1986
  • Littoraria albicans
    Littoraria albicans
    Littoraria albicans is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Metcalfe, 1852
  • Littoraria angulifera Lamarck, 1822 - mangrove periwinkle
    • Littoraria angulifera hessei Boettger 1912
  • Littoraria ardouiniana
    Littoraria ardouiniana
    Littoraria ardouiniana is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Heude, 1885
  • Littoraria articulata
    Littoraria articulata
    Littoraria articulata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria bengalensis
    Littoraria bengalensis
    Littoraria bengalensis is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Reid, 2001
  • Littoraria carinifera
    Littoraria carinifera
    Littoraria carinifera is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Menke, 1830
  • Littoraria cingulata
    Littoraria cingulata
    Littoraria cingulata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria cingulifera
    Littoraria cingulifera
    Littoraria cingulifera is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Dunker, 1845
  • Littoraria coccinea
    Littoraria coccinea
    Littoraria coccinea is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Gmelin, 1791)
  • Littoraria conica
    Littoraria conica
    Littoraria conica is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria delicatula
    Littoraria delicatula
    Littoraria delicatula is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Nevill, 1885
  • Littoraria filosa
    Littoraria filosa
    Littoraria filosa is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (G.B. Sowerby I, 1832)
  • Littoraria fasciata
    Littoraria fasciata
    Littoraria fasciata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Sowerby, 1832
  • Littoraria flammea
    Littoraria flammea
    †Littoraria flammea was a species of sea snail with a gill and an operculum, a marine gastropod mollusk in the family Littorinidae, the periwinkles or winkles.This species was endemic to China. It is now extinct....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1847
  • Littoraria flava
    Littoraria flava
    Littoraria flava is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    King & Broderip, 1832
  • Littoraria flavaosa King & Broderip, 1832
  • Littoraria ianthostoma
    Littoraria ianthostoma
    Littoraria ianthostoma is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Stuckey & D.G. Reid, 2002
  • Littoraria intermedia
    Littoraria intermedia
    Littoraria intermedia is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria irrorata
    Littoraria irrorata
    Littoraria irrorata, common name the Marsh periwinkle, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae....

    Say
    Thomas Say
    Thomas Say was an American naturalist, entomologist, malacologist, herpetologist and carcinologist. A taxonomist, he is often considered to be the father of descriptive entomology in the United States. He described more than 1,000 new species of beetles and over 400 species of insects of other...

    , 1822 - marsh periwinkle
  • Littoraria kraussi Rosewater, 1970
  • Littoraria lutea
    Littoraria lutea
    Littoraria lutea is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1847
  • Littoraria luteola
    Littoraria luteola
    Littoraria luteola is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Quoy & Gaimard, 1833
  • Littoraria mauritiana
    Littoraria mauritiana
    Littoraria mauritiana is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Lamarck, 1822
  • Littoraria melanostoma
    Littoraria melanostoma
    Littoraria melanostoma is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Gray, 1839
  • Littoraria miodelicatula Oyama, 1950
  • Littoraria nebulosa
    Littoraria nebulosa
    Littoraria nebulosa is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Lamarck, 1822 - cloudy periwinkle
  • Littoraria obesa Gmelin
    Johann Friedrich Gmelin
    Johann Friedrich Gmelin was a German naturalist, botanist, entomologist, herpetologist and malacologist.- Education :Johann Friedrich Gmelin was born as the eldest son of Philipp Friedrich Gmelin in 1748 in Tübingen...

    , 1791
  • Littoraria pallescens
    Littoraria pallescens
    Littoraria pallescens is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria philippiana
    Littoraria philippiana
    Littoraria philippiana is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Reeve, 1857
  • Littoraria pintado
    Littoraria pintado
    Littoraria pintado is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Wood, 1820
    • Littoraria pintado pullata Carpenter
      Phillip Pearsall Carpenter
      Philip Pearsall Carpenter Rev. Dr. , who in 1841, was ordained Presbyterian minister in England, and a Doctorate of Philosophy in 1860, and whose field work as a malacologist or conchologist in North America is still well regarded today...

      , 1864
  • Littoraria planaxis Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria punctata Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria rosewateri
    Littoraria rosewateri
    Littoraria rosewateri is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Reid, 1999
  • Littoraria scabra
    Littoraria scabra
    Littoraria scabra, common name : the mangrove periwinkle, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-Description:The size of an adult shell varies between 15 mm and 40 mm....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1847
  • Littoraria sinensis
    Littoraria sinensis
    Littoraria sinensis is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1847
  • Littoraria strigata
    Littoraria strigata
    Littoraria strigata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria subvittata
    Littoraria subvittata
    Littoraria subvittata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Reid, 1986
  • Littoraria sulculosa
    Littoraria sulculosa
    Littoraria sulculosa is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846
  • Littoraria tessellata
    Littoraria tessellata
    Littoraria tessellata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles. The maximum recorded shell length is 23 mm.-References:...

    Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1847
  • Littoraria undulata
    Littoraria undulata
    Littoraria undulata, common name the robust shell, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Gray, 1839
  • Littoraria varia
    Littoraria varia
    Littoraria varia is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Sowerby, 1832
  • Littoraria variegata
    Littoraria variegata
    Littoraria variegata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-References:...

    Souleyet, in Eydoux & Souleyet, 1852
  • Littoraria vespacea
    Littoraria vespacea
    Littoraria vespacea is a species of small sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Reid, 1986
  • Littoraria zebra
    Littoraria zebra
    Littoraria zebra is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Donovan, 1825 - zebra periwinkle

Species brought into synonymy :
  • Littoraria glabrata Philippi
    Rodolfo Amando Philippi
    Rodolfo Amando Philippi was a German-Chilean paleontologist and zoologist....

    , 1846 : synonym of Littoraria coccinea glabrata (Philippi, 1846)

Further reading

  • REID, D G 1986. The Littorinid Molluscs of Mangrove Forests in the Indo-Pacific Region: the genus Littoraria. British Museum (Natural History), London. 228 pp.
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