Lactarius subdulcis
Encyclopedia
Lactarius subdulcis, commonly known as the mild milkcap or beech milk cap, is an edible mushroom
Edible mushroom
Edible mushrooms are the fleshy and edible fruiting bodies of several species of fungi. Mushrooms belong to the macrofungi, because their fruiting structures are large enough to be seen with the naked eye. They can appear either below ground or above ground where they may be picked by hand...

 in the genus Lactarius
Lactarius
Lactarius is a genus of mushroom-producing fungi. The genus, collectively known commonly as milk-caps, are characterized by the fact that they exude a milky fluid if cut or damaged...

. It is brown in colour, with a large number of gills and a particularly thin layer of flesh in the cap. Mycorrhizal, the mushroom is found from late summer to late autumn at the base of beech trees
Beech
Beech is a genus of ten species of deciduous trees in the family Fagaceae, native to temperate Europe, Asia and North America.-Habit:...

 in small groups or individually, where it is one of the two most common species of fungi. Alternatively, it can be found in large groups in fields, sometimes with more than a hundred individual mushrooms. It is found in Europe, and, despite previous research to the contrary, is absent in North America. Although considered edible, it is not particularly useful as food due to its ivy
Ivy
Ivy, plural ivies is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to western, central and southern Europe, Macaronesia, northwestern Africa and across central-southern Asia east to Japan and Taiwan.-Description:On level ground they...

-like taste and the fact that more choice mushrooms will be easily found at the same time. L. subdulcis is known for its abundant, sweet-tasting milk that, unlike the latex of some of its relatives, does not stain fabric yellow.

Taxonomy

Lactarius subdulcis was first described as Agaricus subdulcis by mycologist Christian Hendrik Persoon
Christian Hendrik Persoon
Christiaan Hendrik Persoon was a mycologist who made additions to Linnaeus' mushroom taxonomy.-Early life:...

 in 1801, before English mycologist Samuel Frederick Gray
Samuel Frederick Gray
Samuel Frederick Gray was a British botanist, mycologist, and pharmacologist. He was the father of the zoologists John Edward Gray and George Robert Gray.-Background:...

 placed it in its current genus Lactarius
Lactarius
Lactarius is a genus of mushroom-producing fungi. The genus, collectively known commonly as milk-caps, are characterized by the fact that they exude a milky fluid if cut or damaged...

in 1821 in his The Natural Arrangement of British Plants. The specific epithet is derived from Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 words sub "under", and dulcis "sweet", after the milk's delayed sweet taste. As well as mild milk cap, beech milk cap is an alternate common name.

Description

Lactarius subdulcis has a convex cap
Pileus (mycology)
The pileus is the technical name for the cap, or cap-like part, of a basidiocarp or ascocarp that supports a spore-bearing surface, the hymenium. The hymenium may consist of lamellae, tubes, or teeth, on the underside of the pileus...

 of 3–7 cm (1.4–2.8 in) across that later develops a depression. It sometimes has a small umbo
Umbo (mycology)
thumb|right|[[Cantharellula umbonata]] has an umbo.thumb|right|The cap of [[Psilocybe makarorae]] is acutely papillate.An umbo is a raised area in the center of a mushroom cap. Caps that possess this feature are called umbonate. Umbos that are sharply pointed are called acute, while those that are...

, and in colour can be a reddish-brown, rusty or dark-cinnamon, later paling to buff, though darker in the middle. The cap can be fairly rigid to flexible, and smooth to slightly wrinkled. At first, the margin is incurved though it is sometimes slightly furrowed. The stem
Stipe (mycology)
thumb|150px|right|Diagram of a [[basidiomycete]] stipe with an [[annulus |annulus]] and [[volva |volva]]In mycology a stipe refers to the stem or stalk-like feature supporting the cap of a mushroom. Like all tissues of the mushroom other than the hymenium, the stipe is composed of sterile hyphal...

 is 3–7 cm long and between 6 and 13 mm thick, and is generally cylindrical though can be club-shaped. The stem is sometimes furrowed lengthwise, and is generally the same colour as the cap, though paler at the top. The flesh
Trama (mycology)
In mycology trama is a term for the inner, fleshy portion of a mushroom's basidiocarp, or fruit body. It is distinct from the outer layer of tissue, known as the pileipellis or cuticle, and from the spore-bearing tissue layer known as the hymenium....

 is pale and there is only a thin layer in the cap. The crowded gills are adnate
Adnation
Adnation in plants is the "union of unlike parts; organically united or fused with another dissimilar part, e.g. an ovary to a calyx tube, or stamens to petals". This is in contrast to connation, the fusion of similar organs....

 to slightly decurrent
Decurrent
Decurrent is a term used in botany and mycology to describe plant or fungal parts that extend downward.In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petiole and extend down along the stem...

, and can be white or pink in colour. It has white, plentiful milk that does not stain fabrics yellow, differentiating it from other species of Lactarius
Lactarius
Lactarius is a genus of mushroom-producing fungi. The genus, collectively known commonly as milk-caps, are characterized by the fact that they exude a milky fluid if cut or damaged...

, such as L. decipiens. It has a faint, oily scent.

Spores

Lactarius subdulcis has cream spore print
Spore print
thumb|300px|right|Making a spore print of the mushroom Volvariella volvacea shown in composite: mushroom cap laid on white and dark paper; cap removed after 24 hours showing pinkish-tan spore print...

 with a slight salmon tinge. The spores are oval, with largish warts of around 1 micrometre
Micrometre
A micrometer , is by definition 1×10-6 of a meter .In plain English, it means one-millionth of a meter . Its unit symbol in the International System of Units is μm...

 (μm) which are joined by a well-developed network of mostly thin ridges. The spores measure 7.5–11 μm by 6.5–9 μm, and are amyloid
Amyloid
Amyloids are insoluble fibrous protein aggregates sharing specific structural traits. Abnormal accumulation of amyloid in organs may lead to amyloidosis, and may play a role in various neurodegenerative diseases.-Definition:...

 or ellipsoid in shape.

Distribution and habitat

Lactarius subdulcis is found in Europe; it does not occur in North America, although a number of similar brownish-orange species were formerly classified under this species. It is found in broad-leaved woodland, especially on the floor at the base of beech
Beech
Beech is a genus of ten species of deciduous trees in the family Fagaceae, native to temperate Europe, Asia and North America.-Habit:...

 trees. Along with L. vellereus
Lactarius vellereus
Lactarius vellereus, commonly known as the fleecy milk-cap, is a very large fungus in the milk-cap genus Lactarius. It is one of the two most common milk-caps found with beech trees, with the other being L. subdulcis.-Description:...

, L. subdulcis is the most common fungi found on beech trees. The mushrooms can be found from late summer to late autumn, and are common. They are found individually, or in small groups. They can also be found in fields, generally appearing in large batches, with groups of over a hundred mushrooms not uncommon.

Edibility

Lactarius subdulcis has a mild taste with a slightly bitter after-taste. Though considered edible
Edible mushroom
Edible mushrooms are the fleshy and edible fruiting bodies of several species of fungi. Mushrooms belong to the macrofungi, because their fruiting structures are large enough to be seen with the naked eye. They can appear either below ground or above ground where they may be picked by hand...

 after cooking, it is not recommended, as it has a taste reminiscent of ivy
Ivy
Ivy, plural ivies is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to western, central and southern Europe, Macaronesia, northwestern Africa and across central-southern Asia east to Japan and Taiwan.-Description:On level ground they...

. There are a number of other mushrooms that appear at the same time and in the same areas as L. subdulcis that are preferable to it, including L. mitissimus, meaning that L. subdulcis is not particularly useful as a foodstuff. The milk is one of distinguishing features, having a sweet taste that turns bitter in the mouth, with L. subdulcis being considered a sweet milk mushroom.

External links

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