Gourami
Encyclopedia
Gouramis are a family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

, Osphronemidae, of freshwater
Fresh Water
Fresh Water is the debut album by Australian rock and blues singer Alison McCallum, released in 1972. Rare for an Australian artist at the time, it came in a gatefold sleeve...

 perciform fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

es. The fish are native to Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

, from Pakistan
Pakistan
Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

 and India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

 to the Malay Archipelago
Malay Archipelago
The Malay Archipelago refers to the archipelago between mainland Southeastern Asia and Australia. The name was derived from the anachronistic concept of a Malay race....

 and north-easterly towards Korea
Korea
Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

. The name "gourami" is also used for fish of the families Helostomatidae and Anabantidae. "Gouramis" is an example of a redundant plural. Gourami is already plural, in its original language.

Many gouramis have an elongated ray at the front of their pelvic fins. Many species show parental care: some are mouthbrooder
Mouthbrooder
Mouthbrooding, also known as oral incubation and buccal incubation, is the care given by some groups of animals to their offspring by holding them in the mouth of the parent for extended periods of time. Although mouthbrooding is performed by a variety of different animals, most notably Darwin's...

s, and others, like the Siamese fighting fish
Siamese fighting fish
The Siamese fighting fish , also known as the betta , is a popular species of freshwater aquarium fish. The name of the genus is derived from ikan bettah, taken from a local dialect of Malay...

 (Betta splendens), build bubble nest
Bubble nest
Bubblenests, also spelled bubble nests or bubble-nests, created by some fish species, are floating masses of bubbles blown with an oral secretion, saliva bubbles, and occasionally aquatic plants, or an area for egg deposit attached at the bottom. Fish that build and guard bubble nests are known as...

s. Currently, about 90 species are recognised, placed in 4 subfamilies and about 15 genera.

The name Polyacanthidae has also been used for this family. Some fish now classified as gouramis were previously placed in family Anabantidae. The subfamily Belontiinae was recently demoted from the family Belontiidae. As labyrinth fishes, gouramis have a lung
Lung
The lung is the essential respiration organ in many air-breathing animals, including most tetrapods, a few fish and a few snails. In mammals and the more complex life forms, the two lungs are located near the backbone on either side of the heart...

-like labyrinth organ that allows them to gulp air and use atmospheric oxygen. This organ is a vital innovation for fishes that often inhabit warm, shallow, oxygen-poor water.

As food

Giant gourami
Giant gourami
The Giant Gourami is a gourami, a freshwater fish belonging to the sub-family Osphroneminae in the family Osphronemidae. Some other larger members of this family are also occasionally or regionally referred to as "giant gouramis", including the banded gourami, Trichogaster fasciatus, and the three...

s, Osphronemus goramy, are eaten in some parts of the world. In Indonesia they are often deep-fried and served in sweet-sour sauce, chili sauce, and other spices. Canned, fried gourami are available in China, and throughout the world in oriental supermarkets. Which species the cans contain is uncertain, however; the illustration on the lid of the can of one product depicts the moonlight gourami
Moonlight gourami
Trichopodus microlepis , the moonlight gourami or moonbeam gourami, is a labyrinth fish of the family Osphronemidae. This peaceful, attractive species is a popular aquarium fish....

, Trichogaster microlepis.

In the aquarium

Gouramis, particularly the tri-spot and dwarf, are often kept in home aquariums. As labyrinth fish, they will often swim near the top of the tank. As with other tropical freshwater fish, an aquarium heater
Heater (aquarium)
An aquarium heater is a device used in the fishkeeping hobby to warm the temperature of water in aquariums. Most tropical freshwater and marine aquariums are maintained at temperatures that range from 22-30 °C . The types include glass immersion heaters and undergravel heating...

 is often used. Gouramis will eat either prepared or live foods
Fish food
Aquarium fish feed is plant or animal material intended for consumption by pet fish kept in aquariums or ponds. Fish foods normally contain macro nutrients, trace elements and vitamins necessary to keep captive fish in good health. Approximately 80% of fishkeeping hobbyists feed their fish...

. Some species can grow quite large, and are unsuitable for the general hobbyist.

Compatibility

Generally regarded as peaceful, gouramis are still capable of harassing or killing smaller or long-finned fish. Depending on the species, adult and juvenile males have been known to spar with one another; caution must be taken when keeping males together. Aggression can also occur as a result of overcrowding.

Gouramis have been housed with many species, such as danio
Danio
The Danio genus comprises many of the species of danionins familiar to aquarists. The common name "danio" is used for members of the genera Danio and Devario.-Taxonomy:...

s, mollies
Poecilia
Poecilia is a genus of euryhaline brackish water fish in family Poeciliidae of order Cyprinodontiformes. The type species is P. vivipara. Live-bearers, the Poecilia species are collectively known as mollies, with the exception of Endler's livebearer and the famous guppy...

, silver dollars
Metynnis argenteus
Silver dollar is a common name given to a number of species of fishes, mostly in the genus Metynnis, tropical fish belonging to the Characidae family which are closely related to piranha and pacu...

, and plecostomus
Plecostomus
Hypostomus plecostomus is the scientific name for a type of freshwater tropical Central and South American fish belonging to the family Loricariidae. They are large algae eaters, and to differentiate them from small algae eaters, they are often referred to as plecostomus, often abbreviated as...

 catfish
Catfish
Catfishes are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the heaviest and longest, the Mekong giant catfish from Southeast Asia and the second longest, the wels catfish of Eurasia, to detritivores...

, but will often show aggression toward species with long, flowing fins like male guppies
Guppy
The guppy , also known as the millionfish, is one of the most popular freshwater aquarium fish species in the world. It is a small member of the Poeciliidae family [females long, males long] and like all other members of the family, is live-bearing....

, goldfish
Goldfish
The goldfish is a freshwater fish in the family Cyprinidae of order Cypriniformes. It was one of the earliest fish to be domesticated, and is one of the most commonly kept aquarium fish....

, and betta
Betta
Betta is a large genus of small, often colorful, freshwater ray-finned fishes in the gourami family . The type species is B. picta, the spotted betta.By far the best known Betta species, however, is B...

s.

Species

There are about 96 species in 15 genera.
  • Subfamily Belontiinae (combtail gouramis)
    • Genus Belontia
      Belontia
      Belontia is a genus of gourami. It is the only genus within the subfamily Belontiinae. The Belontia genus contains two species, the Malay combtail and the Ceylonese combtail ....

      • Malay combtail, Belontia hasselti (Cuvier
        Georges Cuvier
        Georges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...

        , 1831)
        .
      • Ceylonese combtail, Belontia signata (Günther, 1861).
  • Subfamily Macropodusinae (paradise fish)(pink and blue)
    • Genus Betta
      Betta
      Betta is a large genus of small, often colorful, freshwater ray-finned fishes in the gourami family . The type species is B. picta, the spotted betta.By far the best known Betta species, however, is B...

      • Akar betta, Betta akarensis Regan, 1910.
      • Betta albimarginata Kottelat
        Maurice Kottelat
        Dr. Maurice Kottelat is a Swiss ichthyologist.In 1976 he entered the University of Neuchâtel where he obtained his diploma in 1987. In 1980 he went to Thailand where he began his field research on Southeast Asiatic and Indonesian fresh water fish species...

         & Ng, 1994
        .
      • Giant betta, Betta anabatoides Bleeker
        Pieter Bleeker
        Pieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....

        , 1851
        .
      • Betta antoni Tan & Ng, 2006.
      • Betta balunga Herre, 1940.
      • Slim betta, Betta bellica Sauvage, 1884.
      • Betta breviobesus Tan & Kottelat, 1998.
      • Betta brownorum Witte & Schmidt, 1992.
      • Betta burdigala
        Betta burdigala
        Betta burdigala is a species of fish in the Osphronemidae family. It is endemic to Indonesia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007.*...

        Kottelat & Ng, 1994.
      • Betta channoides Kottelat & Ng, 1994.
      • Betta chini
        Betta chini
        Betta chini is a species of fish in the Osphronemidae family. It is endemic to Malaysia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Ng, 1993.
      • Betta chloropharynx
        Betta chloropharynx
        Betta chloropharynx is a species of fish in the Osphronemidae family. It is endemic to Indonesia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Kottelat & Ng, 1994.
      • Betta coccina
        Betta coccina
        Betta coccina is a species of fish in the Osphronemidae family. Its closest relatives are Betta channoides, Betta albomarginata, and the Siamese Fighting Fish ....

        Vierke, 1979.
      • Betta compuncta Tan & Ng, 2006.
      • Betta dimidiata Roberts, 1989.
      • Betta edithae Vierke, 1984.
      • Betta enisae
        Betta enisae
        Betta enisae, or the Blue band mouthbrooder, is a species of fish in the Osphronemidae family. It is endemic to Indonesia. First described in 1995 from Kalimantan, Indonesia ....

        Kottelat, 1995.
      • Betta falx Tan & Kottelat, 1998.
      • Betta foerschi Vierke, 1979.
      • Dusky betta, Betta fusca Regan, 1910.
      • Betta hipposideros
        Betta hipposideros
        Betta hipposideros is a species of fish in the Osphronemidae family. It is endemic to Malaysia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Ng & Kottelat, 1994.
      • Betta ibanorum Tan & Ng, 2004.
      • Betta ideii Tan & Ng, 2006.
      • Crescent betta, Betta imbellis Ladiges, 1975.
      • Betta krataios Tan & Ng, 2006.
      • Betta livida
        Betta livida
        Betta livida is a species of fish in the Osphronemidae family. It is endemic to Malaysia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Ng & Kottelat, 1992.
      • Spotfin betta, Betta macrostoma
        Betta macrostoma
        Betta macrostoma is a species of fish in the Osphronemidae family. It is endemic to Brunei and some parts of Borneo.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Regan, 1910.
      • Betta mandor
        Betta mandor
        Betta mandor is a species of Betta. Prior to 2006, the trade names Betta sp. "Mandor" or Betta sp. "ninja" were used in the aquarium trade....

        Tan & Ng, 2006.
      • Betta miniopinna
        Betta miniopinna
        Betta miniopinna is a species of fish in the Osphronemidae family. It is endemic to Indonesia.-Source:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Tan & Tan, 1994.
      • Betta patoti Weber & de Beaufort, 1922.
      • Betta persephone
        Betta persephone
        Betta persephone is a species of freshwater ray-finned fish in the gourami family of order Perciformes. First described in 1986 from a population near Ayer Hitam in Johor state, Malaysia, B. persephone is a rare and elusive species....

        Schaller, 1986.
      • Elephant betta, Betta pi Tan, 1998.
      • Spotted betta
        Spotted betta
        The spotted betta or Javanese fighting fish is a freshwater ray-finned fish in the gourami family of order Perciformes. The type species of genus Betta, it is native to relatively cool, 22‒25 °C , highland streams of Java and Sumatra.In 1967, D. S...

        , Betta picta (Valenciennes
        Achille Valenciennes
        Achille Valenciennes was a French zoologist.Valenciennes was born in Paris, and studied under Georges Cuvier. Valenciennes' study of parasitic worms in humans made an important contribution to the study of parasitology...

        , 1846)
        .
      • Betta pinguis Tan & Kottelat, 1998.
      • Big head betta, Betta prima Kottelat, 1994.
      • Forest betta, Betta pugnax (Cantor
        Theodore Edward Cantor
        Theodore Edward Cantor was a Danish physician, zoologist and botanist.Cantor worked for the British East India Company. He made natural history collections in Penang and Malacca...

        , 1849)
        .
      • Betta pulchra Tan & Tan, 1996.
      • Betta renata Tan, 1998.
      • Toba betta, Betta rubra Perugia, 1893.
      • Betta rutilans Witte & Kottelat, 1991.
      • Betta schalleri Kottelat & Ng, 1994.
      • Krabi betta, Betta simplex
        Betta simplex
        Betta simplex is a species of fish in the Osphronemidae family. It is endemic to Thailand.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Kottelat, 1994.
      • Blue betta, Betta smaragdina
        Betta smaragdina
        Betta smaragdina, or the Emerald green betta is a species of Betta. They are anabantoids, and breathe air.-Importance to humans:Betta smaragdina is relatively uncommon in regular aquarium trade, however, among Betta hobbyists, is a relatively commonly kept species.-External links:**...

        Ladiges, 1972.
      • Betta spilotogena
        Betta spilotogena
        Betta spilotogena is a species of fish in the Osphronemidae family. It is endemic to Indonesia.Betta Spilotogena is a fish species first discovered in Indonesia. This species is classified as critically endangered by the IUCN Red List of Threatened species.Habitat: These fish are often found in...

        Ng & Kottelat, 1994.
      • Siamese fighting fish
        Siamese fighting fish
        The Siamese fighting fish , also known as the betta , is a popular species of freshwater aquarium fish. The name of the genus is derived from ikan bettah, taken from a local dialect of Malay...

        , Betta splendens Regan, 1910.
      • Betta strohi Schaller & Kottelat, 1989.
      • Borneo betta, Betta taeniata Regan, 1910.
      • Betta tomi
        Betta tomi
        Betta tomi is a species of fish in the Osphronemidae family. It is endemic to Malaysia.-References:* Kottelat, M. 1996. . Downloaded on 4 August 2007....

        Ng & Kottelat, 1994.
      • Betta tussyae Schaller, 1985.
      • Betta uberis Tan & Ng, 2006.
      • Howong betta, Betta unimaculata
        Betta unimaculata
        Betta unimaculata is the largest Betta species, and is found in the Indonesian province of East Kalimantan, Borneo. The male is stockier in head form, due to that they're responsible for mouthbrooding, which is holding the eggs in the mouth...

        (Popta, 1905).
      • Betta waseri Krummenacher, 1986.
    • Genus Macropodus
      • Macropodus erythropterus Freyhof & Herder, 2002.
      • Macropodus hongkongensis Freyhof & Herder, 2002.
      • Roundtail paradisefish, Macropodus ocellatus (de Beaufort, 1933).
      • Paradise fish
        Paradise fish
        Paradise fish or paradise gouramis are small freshwater labyrinth fish found in ditches and paddy fields in East Asia, ranging from the Korean Peninsula to Northern Vietnam....

        , Macropodus opercularis (Linnaeus
        Carolus Linnaeus
        Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

        , 1758)
        .
      • Macropodus spechti Schreitmüller, 1936.
    • Genus Malpulutta
      Malpulutta
      Malpulutta is a genus of fish in the Osphronemidae family. It contains the following species:* ornate paradisefish...

      • Spotted gourami, Malpulutta kretseri Deraniyagala, 1937.
    • Genus Parosphromenus
      Parosphromenus
      Parosphromenus is a genus of fish in the Osphronemidae family.It contains 18 described species:* Parosphromenus harveyi...

      • Parosphromenus allani Brown, 1987.
      • Parosphromenus anjunganensis Kottelat, 1991.
      • Parosphromenus bintan Kottelat & Ng, 1998.
      • Licorice gourami, Parosphromenus deissneri (Bleeker
        Pieter Bleeker
        Pieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....

        , 1859)
        .
      • Spiketail gourami, Parosphromenus filamentosus (Oshima, 1919).
      • Parosphromenus linkei Kottelat, 1991.
      • Parosphromenus nagyi Schaller, 1985.
      • Parosphromenus ornaticauda Kottelat, 1991.
      • Parosphromenus paludicola Tweedie, 1952.
      • Parosphromenus parvulus Vierke, 1979.
    • Genus Pseudosphromenus
      • Spiketail paradisefish, Pseudosphromenus cupanus (Cuvier
        Georges Cuvier
        Georges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...

        , 1831)
        .
      • Pseudosphromenus dayi (Köhler, 1908).
    • Genus Trichopsis
      Trichopsis
      Trichopsis is a gourami genus.-Species:Species within the genus are:*Sparkling gourami, Trichopsis pumila*Croaking gourami, Trichopsis vittata*Threestripe gourami, Trichopsis schalleri-Importance to humans:...

      • Pygmy gourami
        Pygmy gourami
        The pygmy gourami is a tropical freshwater fish belonging to the Trichopsis genus in the family . It is native to Southeast Asia, and is found in Thailand, Laos, Peninsular Malaysia, and Indonesia....

        , Trichopsis pumila (Arnold, 1936).
      • Threestripe gourami, Trichopsis schalleri (Kottelat & Ng, 1994).
      • Croaking gourami
        Croaking gourami
        The croaking gourami, Trichopsis vittata, is a species of small freshwater labyrinth fish of the gourami family. They are native to still waters in Southeast Asia and are distributed worldwide via the aquarium trade...

        , Trichopsis vittata (Cuvier
        Georges Cuvier
        Georges Chrétien Léopold Dagobert Cuvier or Jean Léopold Nicolas Frédéric Cuvier , known as Georges Cuvier, was a French naturalist and zoologist...

        , 1831)
        .
  • Subfamily Luciocephalinae (Trichogastrinae)
    • Genus Ctenops
      • Frail gourami, Ctenops nobilis McClelland, 1845.
    • Genus Luciocephalus
      • Gold Spotted Pikehead, Luciocephalus aura (Tan & Ng, 2005)
      • Pikehead, Luciocephalus pulcher (Gray
        John Edward Gray
        John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

        , 1830)
        .
    • Genus Parasphaerichthys
      • Parasphaerichthys lineatus Britz & Kottelat, 2002.
      • Eyespot gourami, Parasphaerichthys ocellatus (de Beaufort, 1933).
    • Genus Sphaerichthys
      • Sphaerichthys acrostoma Vierke, 1979.
      • Chocolate gourami
        Chocolate gourami
        The chocolate gourami, Sphaerichthys osphromenoides, is a small species of gourami. These fish originate from the Malay Peninsula, Sumatra and Borneo.-Appearance and anatomy:...

        , Sphaerichthys osphromenoides Canestrini, 1860.
      • Sphaerichthys selatanensis Vierke, 1979.
      • Sphaerichthys vaillanti Pellegrin, 1930.
    • Genus Trichogaster
      Trichogaster
      Trichopodus is a genus of tropical freshwater labyrinth fish of the gourami family found in Southeast Asia. Gouramis of the Trichopodus genus are closely related to those of Trichogaster , species of both genera have long, thread-like pelvic fins used to sense the environment...

      • Honey gourami
        Honey Gourami
        The Honey Gourami , is a tropical labyrinth fish of the suborder Anabantoidei often bred and kept for aquaria.-Classification:...

        , Trichogaster chuna (Hamilton 1822)
      • Trichogaster fasciata Bloch & Schneider 1801
      • Thick-Lipped Gourami, Trichogaster labiosa Day 1877
      • Dwarf gourami
        Dwarf gourami
        The dwarf gourami, Trichogaster lalius , has an almost translucent blue color, with vertical red to dark orange stripes. In its native range it is dried for food, and it is also kept as an aquarium fish. It has become highly popular for aquaria.Dwarf gouramis from Singapore may carry dwarf gourami...

        , Trichogaster lalius (Hamilton
        Francis Buchanan-Hamilton
        Dr Francis Buchanan, later known as Francis Hamilton but often referred to as Francis Buchanan-Hamilton was a Scottish physician who made significant contributions as a geographer, zoologist, and botanist while living in India.The standard botanical author abbreviation Buch.-Ham. is applied to...

         1822)
    • Genus Trichopodus
      • Trichopodus cantoris
        Trichopodus cantoris
        Trichopodus cantoris is a labyrinth fish of the gourami family.- Physical description :The two black spots are united by a longitudinal zigzag band running from the mouth through the eye to the caudal fin. The caudal is white-spotted. The height of the body is two-fifths or three-sevenths of the...

        (Günther 1861)
      • Trichopodus leerii (Bleeker
        Pieter Bleeker
        Pieter Bleeker was a Dutch medical doctor and ichthyologist, famous for his work on the fishes of East Asia – Atlas Ichthyologique des Orientales Neerlandaises – which was published 1862–1877....

         1852)
        ; pearl gourami or lace gourami
      • Trichopodus microlepis Günther 1861; moonlight gourami or moonbeam gourami
      • Trichopodus pectoralis Regan 1910; snakeskin gourami or Siamese gourami
      • Trichopodus trichopterus (Pallas 1770); three spot gourami, blue gourami, opaline gourami or gold gourami (depending on the colour morph)
  • Subfamily Osphroneminae (giant gouramis)
    • Genus Osphronemus
      Osphronemus
      Osphronemus is the only genus within the subfamily Osphroneminae . These fish are known as the giant gouramis.The genus Osphronemus contains four species.* Elephant ear gourami, Osphronemus exodon...

      • Elephant ear gourami, Osphronemus exodon Roberts, 1994.
      • Giant gourami
        Giant gourami
        The Giant Gourami is a gourami, a freshwater fish belonging to the sub-family Osphroneminae in the family Osphronemidae. Some other larger members of this family are also occasionally or regionally referred to as "giant gouramis", including the banded gourami, Trichogaster fasciatus, and the three...

        , Osphronemus goramy Lacépède, 1801.
      • Giant red tail gourami, Osphronemus laticlavius Roberts, 1992.
      • Osphronemus septemfasciatus Roberts, 1992.

See also

The name "gourami" is used of several other related fish that are now placed in different families:
  • Kissing gourami
    Kissing gourami
    Kissing gouramis, also known as kissing fish or kissers , are large tropical freshwater fish comprising the monotypic labyrinth fish family Helostomatidae . These fish originate from Thailand to Indonesia. They can befood fish which are farmed in their native Southeast Asia...

     (the sole species in the family Helostomatidae)
  • Climbing gourami
    Climbing gourami
    The Anabantidae are a family of perciform fish commonly called the climbing gouramies or climbing perches. As labyrinth fishes, they possess a labyrinth organ, a structure in the fish's head which allows them to breathe atmospheric oxygen...

    s (family Anabantidae)
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