Cupressus goveniana
Encyclopedia
Cupressus goveniana is a species of cypress
Cupressaceae
The Cupressaceae or cypress family is a conifer family with worldwide distribution. The family includes 27 to 30 genera , which include the junipers and redwoods, with about 130-140 species in total. They are monoecious, subdioecious or dioecious trees and shrubs from 1-116 m tall...

 endemic
Endemic (ecology)
Endemism is the ecological state of being unique to a defined geographic location, such as an island, nation or other defined zone, or habitat type; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, all species of lemur are endemic to the...

 to coastal California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 in the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

, where it is found in small, scattered populations, not in large forests.

Description

It is an evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 with a conic to ovoid-conic crown, very variable in size, with mature trees of under 1 m on some sites, to 50 m tall in ideal conditions. The foliage grows in dense sprays, dark green to somewhat yellow-green in color. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are scale-like, 2-5 mm long, and produced on rounded (not flattened) shoots. The seed cones
Conifer cone
A cone is an organ on plants in the division Pinophyta that contains the reproductive structures. The familiar woody cone is the female cone, which produces seeds. The male cones, which produce pollen, are usually herbaceous and much less conspicuous even at full maturity...

 are globose to oblong, 12-22 mm long, with 6 to 10 scales, green at first, maturing brown or gray-brown about 20-24 months after pollination. The cones remain closed for many years, only opening after the parent tree is killed in a wildfire
Wildfire
A wildfire is any uncontrolled fire in combustible vegetation that occurs in the countryside or a wilderness area. Other names such as brush fire, bushfire, forest fire, desert fire, grass fire, hill fire, squirrel fire, vegetation fire, veldfire, and wilkjjofire may be used to describe the same...

, thereby allowing the seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s to colonise the bare ground exposed by the fire. The male cones are 3-5 mm long, and release pollen in February/March; typically, cones of C. goveniana are smaller than those of C. macrocarpa.

Taxonomy

There are two or three varieties
Variety (biology)
In botanical nomenclature, variety is a taxonomic rank below that of species: as such, it gets a three-part infraspecific name....

, treated as distinct species by some botanists:
  • Cupressus goveniana var. goveniana - Gowen Cypress (Vulnerable)
Monterey
Monterey County, California
Monterey County is a county located on the Pacific coast of the U.S. state of California, its northwestern section forming the southern half of Monterey Bay. The northern half of the bay is in Santa Cruz County. As of 2010, the population was 415,057. The county seat and largest city is Salinas...

 County, strictly coastal, within 3 km of the coast and below 200 m altitude. Foliage dark green, not rough, with leaf tips not spreading; cones globose.
  • Cupressus goveniana var. pygmaea (C. pygmaea) - Mendocino Cypress (Vulnerable)
Mendocino
Mendocino County, California
Mendocino County is a county located on the north coast of the U.S. state of California, north of the greater San Francisco Bay Area and west of the Central Valley. As of the 2010 census, the population was 87,841, up from 86,265 at the 2000 census...

 and Sonoma
Sonoma County, California
Sonoma County, located on the northern coast of the U.S. state of California, is the largest and northernmost of the nine San Francisco Bay Area counties. Its population at the 2010 census was 483,878. Its largest city and county seat is Santa Rosa....

 Counties, coastal, within 10 km of the coast and below 500 m altitude. Doubtfully distinguishable from var. goveniana, with very similar foliage and cones. More modern taxonomic thinking classifies Mendocino Cypress as a separate species Cupressus pigmaea
Cupressus pigmaea
Cupressus pigmaea is a taxon of disputed status in the genus Cupressus endemic to certain coastal terraces and coastal mountain ranges of Mendocino and Sonoma Counties in northwestern California...

, and not a variety of C. goveniana.
  • Cupressus goveniana var. abramsiana (Cupressus abramsiana
    Cupressus abramsiana
    Cupressus abramsiana is a cypress taxon of disputed status, placed in either the genus Cupressus or else Callitropsis...

    ) - Santa Cruz Cypress (Endangered)
Santa Cruz
Santa Cruz County, California
Santa Cruz County is a county located on the Pacific coast of the U.S. state of California, on the California Central Coast. The county forms the northern coast of the Monterey Bay. . As of the 2010 U.S. Census, its population was 262,382. The county seat is Santa Cruz...

 and San Mateo
San Mateo County, California
San Mateo County is a county located in the San Francisco Bay Area of the U.S. state of California. It covers most of the San Francisco Peninsula just south of San Francisco, and north of Santa Clara County. San Francisco International Airport is located at the northern end of the county, and...

 Counties, in the Santa Cruz Mountains
Santa Cruz Mountains
The Santa Cruz Mountains, part of the Pacific Coast Ranges, are a mountain range in central California, United States. They form a ridge along the San Francisco Peninsula, south of San Francisco, separating the Pacific Ocean from San Francisco Bay and the Santa Clara Valley, and continuing south,...

 10-20 km inland and at 300-760 m altitude. More distinct, and could well be a valid species, with yellow-green foliage slightly rough-textured from the acute and slightly spreading leaf tips; cones often oval. It also shows similarities to Cupressus sargentii
Cupressus sargentii
Cupressus sargentii is a species of conifer in the Cupressaceae family known by the common name Sargent's cypress. It is endemic to California, where it is known from Mendocino county southwards to Santa Barbara county. This taxon is limited to the Coast Range mountains. Like Mcnab Cypress, it...

.

Ecology

Gowen Cypress occurs with Monterey Cypress, Cupressus macrocarpa
Cupressus macrocarpa
Cupressus macrocarpa, commonly known as Monterey Cypress or Macrocarpa, is a species of cypress that is endemic to the Central Coast of California. In the wild, the species is confined to two small populations, near Monterey and Carmel, California. These two small populations represent what was...

, in the only two groves where C. macrocarpa is known to occur naturally.

External links

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