Bat detector
Encyclopedia
A bat detector is a device used to detect the presence of bat
Bat
Bats are mammals of the order Chiroptera "hand" and pteron "wing") whose forelimbs form webbed wings, making them the only mammals naturally capable of true and sustained flight. By contrast, other mammals said to fly, such as flying squirrels, gliding possums, and colugos, glide rather than fly,...

s by converting their echolocation
Animal echolocation
Echolocation, also called biosonar, is the biological sonar used by several kinds of animals.Echolocating animals emit calls out to the environment and listen to the echoes of those calls that return from various objects near them. They use these echoes to locate and identify the objects...

 ultrasound
Ultrasound
Ultrasound is cyclic sound pressure with a frequency greater than the upper limit of human hearing. Ultrasound is thus not separated from "normal" sound based on differences in physical properties, only the fact that humans cannot hear it. Although this limit varies from person to person, it is...

 signals to audible frequencies as they are emitted by the bats. There are other types of detector which record bat calls so that they can be analysed afterwards, but these are more commonly referred to by their particular function.

To use a bat detector, it is useful to have some understanding of frequencies in hertz
Hertz
The hertz is the SI unit of frequency defined as the number of cycles per second of a periodic phenomenon. One of its most common uses is the description of the sine wave, particularly those used in radio and audio applications....

 (Hz). 1 Hz is 1 complete cycle per second. 1 kilohertz (kHz) is 1000 Hz. Human ears are normally limited to the range from 20 Hz to 16 kHz, whereas bat calls range from about 12 kHz - 160 kHz; so for the most part bat calls are too high in frequency to be heard by most humans. Bat detectors work by converting the bat call sound to a lower, audible frequency, usually about 300 Hz to 5 kHz.

Bats emit calls from about 12 kHz to 160 kHz, but the upper frequencies in this range are rapidly absorbed in air and many bat detectors are limited to around 15 kHz to 125 kHz at best. Bat detectors are available commercially and also can be self-built.

Using bat detectors

Bat detectors are used to detect the presence of bats and also help form conclusions about their species. Some bat calls are distinct and easy to recognise such as the Horseshoe bat
Horseshoe bat
Horseshoe bats are a family of bats. In addition to the single living genus, Rhinolophus, there is one extinct genus, Palaeonycteris. The closely related Hipposideridae are sometimes included within the horseshoe bats as a subfamily, Hipposiderinae...

s; other calls are less distinct between similar species. While bats can vary their calls as they fly and hunt, the ear can be trained to recognise species according to the frequency ranges and repetition rates of the echolocation calls. Bats also emit social calls (non-echolocation calls) at ultrasound frequencies.

A major limitation of acoustic bat detectors is their range, which is limited by the absorption of ultrasound in air. At mid range frequencies around 50 kHz, the maximum range is only about 25 to 30 metres in average atmospheric conditions when bats fly. This decreases with increasing frequency. Some bat calls have components around 20 kHz or even lower and sometimes these can be detected at 2 or 3 times the usual range. However, only the lower frequency components will be detected at a distance. The usable range of bat detectors decreases with humidity and in misty conditions the maximum range can be very low.

It is important to recognise three types of bat echolocation call: frequency modulation
Frequency modulation
In telecommunications and signal processing, frequency modulation conveys information over a carrier wave by varying its instantaneous frequency. This contrasts with amplitude modulation, in which the amplitude of the carrier is varied while its frequency remains constant...

 (FM), constant frequency (CF) (sometimes called amplitude modulation
Amplitude modulation
Amplitude modulation is a technique used in electronic communication, most commonly for transmitting information via a radio carrier wave. AM works by varying the strength of the transmitted signal in relation to the information being sent...

), and composite calls with both FM and CF components. The following illustrates a bat making an FM type call followed by a bat which uses a CF type call:
The FM call is heard as rapid dry clicks and the CF call as peeps. These vary in frequency due to the Doppler effect
Doppler effect
The Doppler effect , named after Austrian physicist Christian Doppler who proposed it in 1842 in Prague, is the change in frequency of a wave for an observer moving relative to the source of the wave. It is commonly heard when a vehicle sounding a siren or horn approaches, passes, and recedes from...

 as the bat flies past. A heterodyne bat detector exaggerates the Doppler effect. As the bat making the CF calls flies toward the detector, the pitch falls.

Several species of bat use a composite FM and CF call starting with a rapid falling FM call which slows to become a CF call at the end, giving a "hockey stick" shape to the graph. This makes the call sound different on a bat detector:
This gives a much wetter sound that the pure FM call. Pipistrelles generally use the hockey stick call for general echolocation, but use only the FM part at times. The end frequencies for the Common Pipistrelle and the Soprano Pipistrelle are around 45 kHz and 55 kHz respectively, but these frequencies can vary widely.

There are three types of "real time" audio bat detector in common use: the heterodyne, frequency division, and time expansion. Some bat detectors combine two or all three types.

Heterodyne

Heterodyne detectors are the most commonly used, and most self-build detectors are of this type. A heterodyne function is often also built into the other types of detector. A heterodyne bat detector simply shifts all the ultrasound frequencies downwards by a fixed amount so we can hear them.

A "heterodyne" is a beat frequency such as can be heard when two close musical notes are sounded together. A heterodyne bat detector combines the bat call with a constant internal frequency so that sum and difference frequencies are generated. For instance a bat call at 45 kHz and an internal frequency of 43 kHz produces output frequencies of 2 kHz and 88 kHz. The 88 kHz frequency is inaudible and is filtered out and the 2 kHz frequency is fed to a loudspeaker or headphones. The internal frequency is displayed on a dial or on a display.

A better quality version of a heterodyne, or direct conversion, bat detector is the super-heterodyne detector. In this case the bat signal is mixed with a high frequency oscillator, typically around 450–600 kHz. The difference frequency is then amplified and filtered in an 'intermediate frequency' or i.f. amplifier before being converted back to audible frequencies again. This design, which is based on standard radio design, gives improved frequency discrimination and avoids problems with interference from the local oscillator.

In more recent DSP-based detectors, the heterodyne conversion can be done entirely digitally.

The tuning issue can be addressed by using a scanning circuit to enable the detector to scan the spectrum automatically and stop scanning when a bat call is heard. One example of such a detector is the Bat Scanner.

It is also possible to use a 'comb spectrum' generator as the local oscillator so that the detector is effectively tuned to lots of frequencies, 10 kHz apart, all at once.

Some early bat detectors used ex Navy, low frequency radio sets, simply replacing the aerial with a microphone and pre-amplifier. It is also possible to modify a portable Long Wave radio to be a bat detector by adjusting the tuning frequencies and replacing the ferrite rod aerial with a microphone and pre-amplifier.

How it is used

The operator guesses the likely species to be present and tunes the frequency accordingly. Many users will start listening around 45 kHz. If a bat is seen or a bat-like call is heard, the frequency is tuned up and down until the clearest sound is heard.

Species like Pipistrelles which end their call with a "hockey stick" CF component can be recognised according to the lowest frequency which gives the clearest "plop" sound. Horseshoe bats give a peeping sound at a frequency depending on their species. FM calls all tend to sound like clicks, but the start and end frequencies and the call repetition pattern can give clues as to the species.

Pros and cons

The advantages of a heterodyne bat detector is that it works in real time, exaggerates the frequency changes of a bat call, is easy to use, and is the least expensive. It is easy to recognise a doppler shift
Doppler effect
The Doppler effect , named after Austrian physicist Christian Doppler who proposed it in 1842 in Prague, is the change in frequency of a wave for an observer moving relative to the source of the wave. It is commonly heard when a vehicle sounding a siren or horn approaches, passes, and recedes from...

 in CF calls of flying bats due to their speed of flight. Stereo listening and recording is possible with models such as the CSE stereo heterodyne detector, and this can help to track bats when visibility is poor.

The disadvantages of a heterodyne bat detector are that it can only convert a narrow band of frequencies, typically 5 kHz, and has to be continually retuned, and can easily miss species out of its current tuned range.

Frequency division

Frequency division (FD) bat detectors synthesise a sound which is a fraction of the bat call frequencies, typically 1/10. This is done by converting the call into a square wave
Square wave
A square wave is a kind of non-sinusoidal waveform, most typically encountered in electronics and signal processing. An ideal square wave alternates regularly and instantaneously between two levels...

, otherwise called a zero crossing signal. This square wave is then divided using an electronic counter by 10 to provide another square wave. Square waves sound harsh and contain harmonic
Harmonic
A harmonic of a wave is a component frequency of the signal that is an integer multiple of the fundamental frequency, i.e. if the fundamental frequency is f, the harmonics have frequencies 2f, 3f, 4f, . . . etc. The harmonics have the property that they are all periodic at the fundamental...

s which can cause problems in analysis so these are filtered out where possible. Some recent all-digital detectors can synthesise a sine wave instead of a square wave. One example of a detector which synthesises a sine-wave FD output is the Griffin

Some FD detectors output this constant level signal which renders background noise and bat calls at the same high level. This causes problems with both listening and analysis. More sophisticated FD detectors such as the Batbox Duet measure the incoming volume level, limiting the noise threshold, and use this to restore the output level variations. This and other sophisticated FD detectors also include a heterodyne detector and provide a jack output so that independent outputs can be recorded for later analysis.

How it is used

With dual output FD detectors, headphones can be used to monitor both outputs simultaneously, or the loudspeaker used with the heterodyne function and the FD output recorded and analysed later. Alternatively, listening to the FD output gives an audible rendering of the bat call at 1/10 frequency.

Dual FD/heterodyne detectors are useful for cross country transects especially when there is a function provided for recording voice notes such as times, locations and recognised bat calls. The output or outputs are recorded on cassette tape, Minidisc or solid state recorders, downloaded to a computer, and analysed using custom software. Calls missed by the heterodyne function, if present, can be seen and measured on the analysis.

Pros and cons

Advantages, As with a heterodyne detector, an FD detector works in real time with or without a heterodyne function. Bat calls can be heard in their entirety over their whole range rather than over a limited frequency range. Retuning with an FD detector is not required although this is done with a dual type with heterodyne. By analysing the recording later, the entire call frequency range and the call pattern can be measured.

A serious disadvantage with real time listening is that the speed of a bat call remains fast, often too fast for the species to be recognised. The frequency changes of CF calls are not exaggerated as with a heterodyne detector and so are less noticeable. Also with some species such as the Lesser Horseshoe bat with a call around 110 kHz, the resulting frequency is still quite high although it can be recorded. The synthesising of the call means that only one bat call can be reproduced at a time and a muddle is caused by simultaneous calls. Surprisingly, this is not a great disadvantage when analysing a recording later.

Time expansion

Time expansion (TE) detectors work by digitising the bat calls at a high sampling rate using an analog-to-digital converter
Analog-to-digital converter
An analog-to-digital converter is a device that converts a continuous quantity to a discrete time digital representation. An ADC may also provide an isolated measurement...

 and storing the digitised signal in an on-board memory.

TE detectors are "real time" devices in that they can be monitored at the time of recording, but there is an inevitable delay while the high speed sampled extract is slowed down and replayed.

How it is used

In real time mode, with or without an associated heterodyne or FD detector, the slowed down calls can be heard as a drawn-out bat call at audible frequencies. Therefore fast FM calls can be heard as a descending note instead of a click. Thus it is possible to hear the difference between FM calls which just sound like clicks on the other types of detector.

After downloading an audio recording to a computer, the original calls are analysed as if they were still at the original non-expanded rate.

Pros and cons

The output can be recorded with an audio recorder as with FD detectors, or with more recent units, the signal can be recorded directly to an internal digital memory such as a compact flash card. The whole waveform is recorded with the full call range being preserved, rather than 1/10 of the waveform as in a FD detector. Since both frequency and amplitude information are preserved in the recorded call, more data is available for species analysis.

Early units were equipped with small memories which limited the length of time that could be digitised. Once the memory was filled (usually only a few seconds maximum), the unit would then replay the recording at slower rate, typically between 1/10 to 1/32 of the rate of the original recording. While the recorded sample is being played back slowly, nothing is being recorded, so the bat calls are being sampled intermittently. For instance, when a 1 second call is being played back at 1/32 rate, 32 seconds of bat calls are not being recorded.

More recent time-expansion recorders use large flash-based memories (such as removable compact-flash cards) and high-bandwidth direct-to-card recording to provide continuous, full-bandwidth real-time recording. Such units can record continuously for many hours while maintaining the maximum information within the signal.

Some units are also equipped with an auto-record function and these can be left in-the-field for many days.

Some units also include a pre-buffer feature to capture events that happened shortly before the 'record' button was pressed which can be useful for manual surveys.

TE detectors are typically used for professional and research work, as they allow a complete analysis of the bats' calls at a later time.

Sampling frequency for digital/TE detectors

Research in 2010 observed that frequencies used by bats can be has high as 250kHz.). The Nyquist–Shannon sampling theorem
Nyquist–Shannon sampling theorem
The Nyquist–Shannon sampling theorem, after Harry Nyquist and Claude Shannon, is a fundamental result in the field of information theory, in particular telecommunications and signal processing. Sampling is the process of converting a signal into a numeric sequence...

 observes that the minimum sampling frequency required to record a signal successfully must be greater than twice the bandwidth of the signal. To record a bandwidth of 250 kHz therefore requires a sampling frequency in excess of 500 kHz. Modern Time-Expansion capable units typically sample at between 300 kHz and 700 kHz. In general, faster is better, though a higher sampling frequency does use more storage space.

Zero crossing analysis

ZCA is best known as an add-on device to the Anabat bat detector. The original bat calls are digitised and the zero crossing points used to produce a data stream which is recorded on a Compact Flash card. There are sophisticated timing controls and the device can be set to respond to bat calls, so that many hours of recording are available in unmanned situations. The purpose of ZCA is to reduce the amount of data which must be recorded to the memory and may be considered as a simple form of lossy data compression. Historically, to achieve long recording times, such information reduction has been necessary due to memory capacity limitations and memory cost.

The solid state ZCA recording is analysed by custom software to produce a time/frequency plot of each call which can be examined for species recognition in a similar way to FD or TE recordings.

How it is used

The ZCA detector is usually placed in a bat roost or bat flight path and left for a number of days to collect data. Thus it is less labour intensive than using a manned bat detector in real time.

Pros and cons

While the ZCA detector can also be used in real time, its value is for remote recording over long periods. The analysis is similar to that for FD recordings, but there is no amplitude data included. However it does accurately record each zero crossing point, rather than only one in ten. As with all recording devices triggered by an input, a ZCA detector recording automatically is prone to ultrasonic interference from insects such as crickets. Filters can be written to select a characteristic frequency of certain species and ignore others; some (CF species) are more easily filtered, others are nigh on impossible.

High frequency recording

This can be done by using a high speed digitiser peripheral on a computer such as a laptop. This is not a bat detector as such, but recordings of bat calls can be analysed similarly to TE recordings. This method produces large data files and produces no means of detecting bat calls without the simultaneous use of a bat detector. There are however also more sophisticated systems such as the Avisoft-UltraSoundGate that can replace a conventional bat detector. These advanced systems additionally provide a real-time spectrographic display, automated call parameter measurement and classification tools, integrated GPS functionality and a versatile metadata input tool for documenting the recordings.

DSP detectors

DSP bat detectors aim to provide an acoustically accurate portrayal of bat calls by using a digital signal processor
Digital signal processor
A digital signal processor is a specialized microprocessor with an architecture optimized for the fast operational needs of digital signal processing.-Typical characteristics:...

 to map bats' ultrasounds signals to audible sounds; different algorithms are being used to accomplish this, and there is active development and tuning of algorithms going on.
One strategy called "frequency shifting" uses a FFT signal analysis in order to find the main frequency and signal power, then using digital simulation a new audible wave is synthesized from the original one divided by a defined value.

The processes of Frequency Division and Heterodyne conversion can also be performed digitally.

Time domain signal coding

This type of bat detector is believed to be in pre-production or experimental and is not available commercially. Research is in progress for analysing many types of ultrasound calls and sounds besides those of bats.

A TDSC detector digitises the original calls and derives a two dimensional data string by analysing the parameters of each call with respect to time. This is analysed by a neural network
Artificial neural network
An artificial neural network , usually called neural network , is a mathematical model or computational model that is inspired by the structure and/or functional aspects of biological neural networks. A neural network consists of an interconnected group of artificial neurons, and it processes...

 to provide pattern recognition for each species.

Non-acoustic detection

Visual observation is the obvious means of detecting bats, but of course this can only be done in daylight or crepuscular
Crepuscular
Crepuscular animals are those that are active primarily during twilight, that is during dawn and dusk. The word is derived from the Latin word crepusculum, meaning "twilight." Crepuscular is, thus, in contrast with diurnal and nocturnal behavior. Crepuscular animals may also be active on a bright...

 conditions. (dusk and dawn). Emergence counts are done visually at dusk, using a bat detector for confirmation of species. In lower light conditions a night vision device can be used but the more affordable generation 1 type has a lag time which fails to provide a suitable image of a flying bat.

Infrared
Infrared
Infrared light is electromagnetic radiation with a wavelength longer than that of visible light, measured from the nominal edge of visible red light at 0.74 micrometres , and extending conventionally to 300 µm...

 (IR) cameras
and camcorders are used with an IR illuminator to observe bat emergences and bat behaviour inside and outside roosts. The problem with this method is that deriving a count from a recording is tedious and time consuming, but camcorders can be useful as a backup in roost emergence counts to observe bats re-entering the roost. Many Sony camcorders are sensitive to infrared.

Infrared
Infrared
Infrared light is electromagnetic radiation with a wavelength longer than that of visible light, measured from the nominal edge of visible red light at 0.74 micrometres , and extending conventionally to 300 µm...

 beam devices
usually consist of a dual array of invisible IR beams. The size of the roost entrance determines the number of beams necessary and thus the power required and potential for off-mains use. Single beam DIY systems are available for bat boxes but these do not log the direction of transit. Almost all the systems in use today are non-commercial or DIY. A system in use in some mines in Wisconsin uses two arrays of beams however they are spaced quite far apart and consequently only log approximately 50% of the bats although extrapolated figures are achieved through correlation of time stamped video and beam break data. The Countryside Council for Wales
Countryside Council for Wales
The Countryside Council for Wales is an Assembly Government Sponsored Body. It is the Welsh Assembly Government's wildlife conservation authority for Wales...

 (CCW) uses two similar systems with beams spaced close enough together that every bat transiting the entrance is logged along with the temperature. These systems require either mains power or 12v deep cycle batteries. They can be used in conjunction with an Anabat Zcaim installed in a 6" soil pipe and pointed across the roost entrance to discriminate between species by correlating the time stamp data from the IR array and filtered Anabat Zcaim data for horsehoe bats (relatively easy due to their easily identifiable CF echolocation which can be filtered automatically using Anabat software).

Data from beam break systems must be carefully analysed to eliminate "light sampling behaviour" (environment sampling) where the bats repeatedly leave the roost and return immediately if the conditions are not suitable. Some systems discriminate for bat sized animals; they determine if the beams are broken by a bat sized animal and ignore all other transits. It is important that data is analysed using a methodology which takes light sampling behaviour into account. The method which seems to give the most accurate results is as follows: "out" transit assigned 1, "in" transit assigned -1. Start count is set to zero at 4 pm daily. Using a spreadsheet, the counts are added cumulatively from 4 pm each day until 9 am the next day. The maximum "positive" count can easily be found for each day. Since every transit is time stamped, the exact time of the maximum daily count is also known. Light sampling counts are eliminated from the data since an "out" 1 is cancelled by an "in" -1, resulting in a cumulative count of zero for light sampling bats.

Thermal imagers
Thermography
Infrared thermography, thermal imaging, and thermal video are examples of infrared imaging science. Thermal imaging cameras detect radiation in the infrared range of the electromagnetic spectrum and produce images of that radiation, called thermograms...

which are of a high enough definition to register bats at over 30 metres range are expensive, but have been used to assess the dangers of wind turbines to birds and bats. "Affordable" thermal imagers have a bat detecting range about the same order of acoustic bat detectors due to the small size and the low heat emissions of bats.

Passive infrared sensors are slow with a response speed of the order of a tenth of a second and will normally not detect a small fast mammal like a bat.

Radar has been used to detect bats beyond the acoustic limit, but is very costly in equipment and man hours. Bird Aircraft Strike Hazard (BASH) installations are capable of detecting bats, but are usually situated where few bats fly. There are very few suitable mobile terrestrial radars available anywhere. Hand-held doppler radar modules have been used in the field to allow researchers to compensate for the doppler shift imposed on recordings of bat signals due to their flight speed. This allows the researchers to tell whether the bats are changing the pitch of their calls in flight.

External links

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