Anisophylleaceae is a small family with four genera in the order Cucurbitales

Cucurbitales

The Cucurbitales are an order of flowering plants, included in the rosid group of dicotyledons. This order mostly belongs to tropical areas, with limited presence in subtropic and temperate regions. The order includes shrubs and trees, together with many herbs and climbers...

, according to the APG II

Angiosperm Phylogeny Group

The Angiosperm Phylogeny Group, or APG, refers to an informal international group of systematic botanists who came together to try to establish a consensus on the taxonomy of flowering plants that would reflect new knowledge about plant relationships discovered through phylogenetic studies., three...

. However it is more isolated from the other suprafamilal clade

Clade

A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

s in this order, while it shows some similarities in flower morphology with the genus Ceratopetalum

Ceratopetalum

Ceratopetalum is a genus of nine species of shrub and tree in the family Cunoniaceae. They are found along the eastern coast of Australia and extend north to New Guinea. Two Australian species are among the best known, one being C. apetalum or coachwood, renowned as a timber tree, and C...

(family Cunoniaceae

Cunoniaceae

The Cunoniaceae is a family of 26 genera and about 350 species of woody plants in the Antarctic flora, with many laurifolia species with glossy leaves endemic to laurel forest habitat. The family is native to Australia, New Caledonia, New Guinea, New Zealand, southern South America, the Mascarene...

, order Oxalidales

Oxalidales

The Oxalidales are an order of flowering plants, included within the rosid subgroup of eudicots. The following families are typically placed here:* Family Brunelliaceae* Family Cephalotaceae * Family Connaraceae...

). Several wood features of this family, are more primitive than those of the other families in the order Cucurbitales.

Previously this family was categorized under its own order Anisophylleales by Takhtajan

Takhtajan system

A system of plant taxonomy, the Takhtajan system of plant classification was published by Armen Takhtajan, in several versions from the 1950s onwards. It is usually compared to the Cronquist system. Key publications:-External links:* Takhtajan system at...

 in 1997

It is a pantropical family of shrubs and medium-sized to fairly large trees, occurring in wet, tropical forests and swamps of America, Africa and Asia.

The palmately-veined leaves

Leaf

A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 have a rather leathery texture, entire margins, and are often asymmetrical at the base. They have minute stipules or simply lack them. They are alternate; spiral, or distichous, or four-ranked (such as in Anisophyllea). The paired leaves may be different in size or shape.

The small flower

Flower

A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are regular and 3-merous

Merous

Merosity is the number of component parts in each whorl of a plant structure. It is most commonly used in the context of flowers, in which case it refers to the number of sepals in the calyx, the number of petals in the corolla, and the number of stamens in each whorl of the androecium...

 to 5-merous. They are usually aggregated in axillary raceme

Raceme

A raceme is a type of inflorescence that is unbranched and indeterminate and bears pedicellate flowers — flowers having short floral stalks called pedicels — along the axis. In botany, axis means a shoot, in this case one bearing the flowers. In a raceme, the oldest flowers are borne...

s or panicle

Panicle

A panicle is a compound raceme, a loose, much-branched indeterminate inflorescence with pedicellate flowers attached along the secondary branches; in other words, a branched cluster of flowers in which the branches are racemes....

s. The flower type varies considerably, most are monoecious, except Combretocarpus; which is hermaphrodite, having perfect flowers.

The inferior, 3 to 4-locular ovary

Ovary (plants)

In the flowering plants, an ovary is a part of the female reproductive organ of the flower or gynoecium. Specifically, it is the part of the pistil which holds the ovule and is located above or below or at the point of connection with the base of the petals and sepals...

 develops into a drupe

Drupe

In botany, a drupe is a fruit in which an outer fleshy part surrounds a shell of hardened endocarp with a seed inside. These fruits develop from a single carpel, and mostly from flowers with superior ovaries...

 or a samara

Samara (fruit)

A samara is a type of fruit in which a flattened wing of fibrous, papery tissue develops from the ovary wall. A samara is a simple dry fruit and indehiscent . It is a winged achene...

(as in Combretocarpus) with usually one seed, but with 3-4 seeds in Poga.