Acromyrmex aspersus
Encyclopedia
Acromyrmex aspersus is a species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of New World ants of the subfamily Myrmicinae
Myrmicinae
Myrmicinae is a subfamily of ants. There are about 140 genera within the group, with the family being cosmopolitan. The pupae lack cocoons. Some species retain a functional sting. The petioles of Myrmicinae consist of two nodes...

 of the genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Acromyrmex. It is found in the wild naturally in southern Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

 and Peru
Peru
Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

.

Overview

Commonly known as "leaf-cutter ants" they are a species of ant from one of the two genera of advanced attines
Fungus-growing ants
The fungus-growing ants comprises all the known fungus-growing ant species in the world participating in ant-fungus mutualism. Leafcutter ants, including Atta and Acromyrmex, make up 2 of the genera.-See also:...

 within the tribe Attini.

Acromyrmex can be identified from the closely related Atta genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of leafcutter ants since they have 4 pairs of spines and a rough exoskeleton on the upper surface of the thorax
Thorax (insect anatomy)
The thorax is the mid section of the insect body. It holds the head, legs, wings and abdomen. It is also called mesosoma in other arthropods....

.

A mature Acromyrmex aspersus colony contains mostly sterile female workers. They are divided into castes
Eusociality
Eusociality is a term used for the highest level of social organization in a hierarchical classification....

, based mostly on size, that perform different functions. Acromyrmex exhibit a high degree of biological polymorphism
Polymorphism (biology)
Polymorphism in biology occurs when two or more clearly different phenotypes exist in the same population of a species — in other words, the occurrence of more than one form or morph...

, four castes being present in established colonies - minims (or "garden ants"), minors, mediae and majors. Majors are also known as soldiers or dinergates. Each caste has a specific function within the colony. Acromyrmex ants are less polymorphic than the other genera of leafcutter ants Atta, meaning that there is comparatively less differential in size from the smallest to largest types of Acromymex. The high degree of polymorphism in this genus is also suggestive of its high degree of advancement.

Like Atta, Acromyrmex subsists mostly on a particular species of fungus (genus Leucocoprinus
Leucocoprinus
Leucocoprinus is a genus of fungi in the family Agaricaceae. Its best known member is the yellow pot-plant mushroom , found worldwide. The type species is Leucocoprinus cepistipes...

) which it cultivates on a medium of masticated leaf tissue. This is the sole food of the queen
Eusociality
Eusociality is a term used for the highest level of social organization in a hierarchical classification....

 and other colony members that remain in the nest. The media workers also gain subsistence from plant sap they ingest whilst physically cutting out sections of leaf from a variety of plants.

Before leaving their parent colonies, winged females take a small section of fungus into their bucchal pouches and it is with this that the subsequently wingless queens 'seed' the fungus gardens of incipient colonies, cutting and collecting the first few sections of leaf themselves.

Acromyrmex has evolved to change foodplant constantly, preventing a colony from completely stripping off leaves and thereby killing trees, thus avoiding negative biological feedback on account of their sheer numbers. However, this does not diminish the huge quantities of foliage they harvest.

Sub-species

The species Acromyrmex aspersus contains the following subspecies
  • Acromyrmex aspersus aspersus F. Smith
    Frederick Smith (entomologist)
    Frederick Smith was a British entomologist.Smith worked in the zoology department of the British Museum from 1849, specialising in the Hymenoptera. In 1875 he was promoted to Assistant Keeper of Zoology...

    , 1858
  • Acromyrmex aspersus fuhrmanni Forel, 1914
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