Psammosere
Encyclopedia
A psammosere is a seral community
Seral community
A seral community is an intermediate stage found in ecological succession in an ecosystem advancing towards its climax community. In many cases more than one seral stage evolves until climax conditions are attained...

, an ecological succession
Ecological succession
Ecological succession, is the phenomenon or process by which a community progressively transforms itself until a stable community is formed. It is a fundamental concept in ecology, and refers to more or less predictable and orderly changes in the composition or structure of an ecological community...

 that began life on newly exposed coastal sand. Most common psammoseres are sand dune
Dune
In physical geography, a dune is a hill of sand built by wind. Dunes occur in different forms and sizes, formed by interaction with the wind. Most kinds of dunes are longer on the windward side where the sand is pushed up the dune and have a shorter "slip face" in the lee of the wind...

 systems.

In a psammosere, the organism
Organism
In biology, an organism is any contiguous living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homoeostasis as a stable whole.An organism may either be unicellular or, as in the case of humans, comprise...

s closest to the sea will be pioneer species
Pioneer species
Pioneer species are species which colonize previously uncolonized land, usually leading to ecological succession. They are the first organisms to start the chain of events leading to a livable biosphere or ecosystem...

: salt-tolerant species
Halophyte
A halophyte is a plant that grows where it is affected by salinity in the root area or by salt spray, such as in saline semi-deserts, mangrove swamps, marshes and sloughs, and seashores. An example of a halophyte is the salt marsh grass Spartina alterniflora . Relatively few plant species are...

 such as littoral algae
Algae
Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

 and glasswort with marram grass
Marram grass
Ammophila is a genus consisting of two or three very similar species of grasses; common names for these grasses include Marram Grass, Bent Grass, and Beachgrass...

 stabilising the dunes. Progressing inland many characteristic features change and help determine the natural succession of the dunes. For instance, the drainage slows down as the land becomes more compact and has better soils, and the pH
PH
In chemistry, pH is a measure of the acidity or basicity of an aqueous solution. Pure water is said to be neutral, with a pH close to 7.0 at . Solutions with a pH less than 7 are said to be acidic and solutions with a pH greater than 7 are basic or alkaline...

 drops as the proportion of seashell fragments reduces and the amount of humus increases. Sea purslane
Sesuvium portulacastrum
Sesuvium portulacastrum, commonly known as shoreline purslane or "sea purslane", is a sprawling perennial herb that grows in coastal areas throughout much of the world.-Description:...

, sea lavender, meadow grass and heather
Calluna
Calluna vulgaris is the sole species in the genus Calluna in the family Ericaceae. It is a low-growing perennial shrub growing to tall, or rarely to and taller, and is found widely in Europe and Asia Minor on acidic soils in open sunny situations and in moderate shade...

 eventually grade into a typical non-maritime terrestrial eco-system. The first trees (or pioneer trees) that appear are typically fast-growing trees such as birch
Birch
Birch is a tree or shrub of the genus Betula , in the family Betulaceae, closely related to the beech/oak family, Fagaceae. The Betula genus contains 30–60 known taxa...

, willow
Willow
Willows, sallows, and osiers form the genus Salix, around 400 species of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere...

 or rowan
Rowan
The rowans or mountain-ashes are shrubs or small trees in genus Sorbus of family Rosaceae. They are native throughout the cool temperate regions of the Northern Hemisphere, with the highest species diversity in the mountains of western China and the Himalaya, where numerous apomictic microspecies...

. In turn these will be replaced by slow-growing, larger trees such as ash
Ash tree
Fraxinus is a genus flowering plants in the olive and lilac family, Oleaceae. It contains 45-65 species of usually medium to large trees, mostly deciduous though a few subtropical species are evergreen. The tree's common English name, ash, goes back to the Old English æsc, while the generic name...

 and oak
Oak
An oak is a tree or shrub in the genus Quercus , of which about 600 species exist. "Oak" may also appear in the names of species in related genera, notably Lithocarpus...

. This is the climax community
Climax community
In ecology, a climax community, or climatic climax community, is a biological community of plants and animals which, through the process of ecological succession — the development of vegetation in an area over time — has reached a steady state. This equilibrium occurs because the climax community...

, defined as the point where a plant succession does not develop any further because it has reached equilibrium with the environment
Ecosystem
An ecosystem is a biological environment consisting of all the organisms living in a particular area, as well as all the nonliving , physical components of the environment with which the organisms interact, such as air, soil, water and sunlight....

, in particular the climate
Climate
Climate encompasses the statistics of temperature, humidity, atmospheric pressure, wind, rainfall, atmospheric particle count and other meteorological elemental measurements in a given region over long periods...

.

In an idealised coastal psammosere model, at the seaward edge of the sand dune the pH of the soil is typically alkaline/neutral with a pH of 7.0/8.0 particularly where shell fragments provide a significant component of the sand. Tracking inland across the dunes a podsol
Podsol
In soil science, podzols are the typical soils of coniferous, or boreal forests. They are also the typical soils of eucalypt forests and heathlands in southern Australia...

develops with a pH of 5.0/ 4.0 followed by mature podsols at the climax with a pH of 3.5 - 4.5.

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