Piper cenocladum
Encyclopedia
The tropical rainforest
Tropical rainforest
A tropical rainforest is an ecosystem type that occurs roughly within the latitudes 28 degrees north or south of the equator . This ecosystem experiences high average temperatures and a significant amount of rainfall...

 understory shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 Piper cenocladum is a member of the same genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 as kava
Kava
Kava or kava-kava is a crop of the western Pacific....

, betel
Betel
The Betel is the leaf of a vine belonging to the Piperaceae family, which includes pepper and Kava. It is valued both as a mild stimulant and for its medicinal properties...

, and black pepper
Black pepper
Black pepper is a flowering vine in the family Piperaceae, cultivated for its fruit, which is usually dried and used as a spice and seasoning. The fruit, known as a peppercorn when dried, is approximately in diameter, dark red when fully mature, and, like all drupes, contains a single seed...

. It is a myrmecophyte
Myrmecophyte
Myrmecophyte is a plant that lives in a mutualistic association with a colony of ants. There are over 100 different genera of myrmecophytes. These plants possess structural adaptations that provide ants with food and/or shelter. These specialized structures include domatia, food bodies, and...

, a plant that lives in ecological mutualism with ant
Ant
Ants are social insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. More than...

s. This plant and three or four other closely related species are known collectively as ant plants or ant pipers. This species has broad, bright green leaves and grows in dim, swampy areas deep in the rainforest of Costa Rica
Costa Rica
Costa Rica , officially the Republic of Costa Rica is a multilingual, multiethnic and multicultural country in Central America, bordered by Nicaragua to the north, Panama to the southeast, the Pacific Ocean to the west and the Caribbean Sea to the east....

 and surrounding countries.

This ant piper has hollow petiole
Petiole (botany)
In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...

s which provide a home for ants, especially of the species Pheidole bicornis
Pheidole
Pheidole is a genus of ant that belongs to the ant subfamily Myrmicinae.-The Genus:The genus Pheidole is widespread and ecologically dominant...

. The plant also produces food bodies that the ants use as their main food source. The ants in turn defend the plant against predation by herbivorous caterpillars and fungi. Adding to the complexity of this food web
Food chain
A food web depicts feeding connections in an ecological community. Ecologists can broadly lump all life forms into one of two categories called trophic levels: 1) the autotrophs, and 2) the heterotrophs...

 is the beetle
Beetle
Coleoptera is an order of insects commonly called beetles. The word "coleoptera" is from the Greek , koleos, "sheath"; and , pteron, "wing", thus "sheathed wing". Coleoptera contains more species than any other order, constituting almost 25% of all known life-forms...

Tarsobaenus letourneauae, a specialized predator which lives in the plant's petioles and feeds upon the ants and their eggs. This food web is an example of ecological top-down control, in which the top predator affects lower levels of the system. The beetle reduces the number of ants, which allows more herbivory to occur, causing foliage loss on the plant.
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