Ornithoptera priamus
Encyclopedia
Ornithoptera priamus, commonly known as the common green birdwing, Cape York Birdwing, Priam's Birdwing or Northern Birdwing, is a widespread species of birdwing
Birdwing
Birdwings are papilionid butterflies native to the Indian Subcontinent, mainland and archipelagic Southeast Asia and Australasia, and are usually regarded as belonging to three genera: Ornithoptera, Trogonoptera and Troides. Some authorities include additional genera...

 butterfly
Butterfly
A butterfly is a mainly day-flying insect of the order Lepidoptera, which includes the butterflies and moths. Like other holometabolous insects, the butterfly's life cycle consists of four parts: egg, larva, pupa and adult. Most species are diurnal. Butterflies have large, often brightly coloured...

 found in the central and south Moluccas, New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

, Bismarck Archipelago
Bismarck Archipelago
The Bismarck Archipelago is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea.-History:...

, Solomon Islands
Solomon Islands
Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

, and northeast Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

.

Variation

Ornithoptera priamus is a variable species. There have been as many as 99 subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 described (most of which are synonyms
Synonym (taxonomy)
In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

 of O. priamus poseidon), with many more named variants and forms described for both sexes.

As in other species of the genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Ornithoptera, the females are larger and less vividly coloured than the males, them being mainly blackish or dark brown with patterns in pale brown, yellow or white. Nonetheless, females are extremely variable, and they may have colour patterns similar to females of other Ornithoptera species.

In males, iridescent areas of the wing are typically green, although some subspecies endemic to island groups east of New Guinea have blue males. These subspecies are O. priamus urvilleanus (New Ireland
New Ireland (island)
New Ireland is a large island in Papua New Guinea, approximately 7,404 km² in area. It is the largest island of the New Ireland Province, lying northeast of the island of New Britain. Both islands are part of the Bismarck Archipelago, named after Otto von Bismarck, and they are separated by...

, Bougainville
Bougainville Island
Bougainville Island is the main island of the Autonomous Region of Bougainville of Papua New Guinea. This region is also known as Bougainville Province or the North Solomons. The population of the province is 175,160 , which includes the adjacent island of Buka and assorted outlying islands...

, and Solomon Islands), O. priamus miokensis (Mioko Island), and O. priamus caelestis (Louisiade Archipelago
Louisiade Archipelago
The Louisiade Archipelago is a string of ten larger volcanic islands frequently fringed by coral reefs, and 90 smaller coral islands located 200 km southeast of New Guinea, stretching over more than and spread over an ocean area of between the Solomon Sea to the north and the Coral Sea to...

).

However, it is likely that O. priamus miokensis is actually a hybrid between O. priamus urvillenaus and O. priamus bornemanni from the neighbouring islands of New Britain
New Britain
New Britain, or Niu Briten, is the largest island in the Bismarck Archipelago of Papua New Guinea. It is separated from the island of New Guinea by the Dampier and Vitiaz Straits and from New Ireland by St. George's Channel...

 and Bougainville. Specimens of this subspecies are variable, and can be identical to specimens of either parent, or intermediates. Immigration may also explain its rarity, as Mioko is a small island and, as of 2001, its hostplants had been reduced to a few Aristolochia tagala
Aristolochia tagala
Aristolochia tagala is commonly known as Indian birthwort and locally as Dutchman's pipe.-Distribution:It is widely distributed: the distribution is from the Himalaya to Sri Lanka through South East Asia and China, to Oceania...

vines growing in a local village.

Subspecies

Ornithoptera priamus was originally described from Seram Island, Indonesia. Some list as few as five subspecies (excluding nominal), but a taxonomic review is needed and most recognise more:
  • O. p. admiralitatis (Admiralty Islands
    Admiralty Islands
    The Admiralty Islands are a group of eighteen islands in the Bismarck Archipelago, to the north of New Guinea in the south Pacific Ocean. These are also sometimes called the Manus Islands, after the largest island. These rainforest-covered islands form part of Manus Province, the smallest and...

     and nearby smaller islands, Papua New Guinea)
  • O. p. albiro (Tayandu Islands
    Tayandu Islands
    The Tayandu Islands are a group of low-lying islands just west of the larger Kai Islands of Maluku, Indonesia. The main group consists of Tayando , Walir, Heniar and several smaller isle-lets...

    , Indonesia) – possibly a junior synonym
    Synonym (taxonomy)
    In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

     of O. p. hecuba
  • O. p. arruana (Aru Islands
    Aru Islands
    The Aru Islands are a group of about ninety-five low-lying islands in the Maluku province of eastern Indonesia. They also form a regency of Indonesia.-Geography:...

    , Indonesia)
  • O. p. aureus (Arfak Mountains
    Arfak Mountains
    The Arfak Mountains are a mountain range found on the Bird's Head Peninsula in the Province of West Papua, Indonesia. The mountain range is rugged and remote, rising up to 7,700 feet above sea level...

    , West Papua, Indonesia) – possibly a junior synonym of O. p. poseidon
  • O. p. boisduvali (Woodlark Island
    Woodlark Island
    Woodlark Island, known to its inhabitants simply as Woodlark or Muyua, is an island in Milne Bay Province, Papua New Guinea. It is called Murua by the inhabitants of some other islands in the province...

    , Papua New Guinea)
  • O. p. bornemanni (New Britain
    New Britain
    New Britain, or Niu Briten, is the largest island in the Bismarck Archipelago of Papua New Guinea. It is separated from the island of New Guinea by the Dampier and Vitiaz Straits and from New Ireland by St. George's Channel...

     and nearby smaller islands, Papua New Guinea)
  • O. p. caelestis (Louisiade Archipelago
    Louisiade Archipelago
    The Louisiade Archipelago is a string of ten larger volcanic islands frequently fringed by coral reefs, and 90 smaller coral islands located 200 km southeast of New Guinea, stretching over more than and spread over an ocean area of between the Solomon Sea to the north and the Coral Sea to...

    , Papua New Guinea)
  • O. p. demophanes (D'Entrecasteaux
    D'Entrecasteaux Islands
    D'Entrecasteaux Islands are situated near the eastern tip of New Guinea in the Solomon Sea in Milne Bay Province of Papua New Guinea. The group spans a distance of 160 km, has a total land area of approximately 3,100 km² and is separated from the Papua New Guinea mainland by the...

     and Trobriand Islands
    Trobriand Islands
    The Trobriand Islands are a 450 km² archipelago of coral atolls off the eastern coast of New Guinea. They are situated in Milne Bay Province in Papua New Guinea. Most of the population of 12,000 indigenous inhabitants live on the main island of Kiriwina, which is also the location of the...

    , Papua New Guinea)
  • O. p. garainaensis (vicinity of Garaina, Papua New Guinea?) – possibly a junior synonym of O. p. poseidon
  • O. p. gebeensis (Gebe Island, Indonesia)
  • O. p. hecuba (Kai Islands
    Kai Islands
    The Kai Islands of Indonesia are in the south-eastern part of the Maluku Islands, in Maluku Province.-Geography:...

    , Indonesia)
  • O. p. impensis (Manipa Island, Indonesia)
  • O. p. kassandra (Yapen Island, West Papua, Indonesia)
  • O. p. macalpinei (Iron
    Iron Range
    The Iron Range is a region that makes up the northeastern section of Minnesota in the United States. "The Range", as it is known by locals, is a region with multiple distinct bands of iron ore...

     and McIlwraith Range
    McIlwraith Range
    The McIlwraith Range is a rugged, dissected granite plateau on the Cape York Peninsula, Far North Queensland, Australia. Part of the Great Dividing Range, it covers about 3000 km2 and lies about 15 km east of the town of Coen, and 550 km north of Cairns...

    s, Queensland
    Queensland
    Queensland is a state of Australia, occupying the north-eastern section of the mainland continent. It is bordered by the Northern Territory, South Australia and New South Wales to the west, south-west and south respectively. To the east, Queensland is bordered by the Coral Sea and Pacific Ocean...

    , Australia) – possibly a junior synonym of O. p. poseidon
  • O. p. miokensis (Mioko Island, Papua New Guinea)
  • O. p. poseidon (New Guinea
    New Guinea
    New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

     and northern islands of Torres Straight
    Torres Strait Islands
    The Torres Strait Islands are a group of at least 274 small islands which lie in Torres Strait, the waterway separating far northern continental Australia's Cape York Peninsula and the island of New Guinea but Torres Strait Island known and Recognize as Nyumaria.The islands are mostly part of...

    )
  • O. p. priamus (Seram, Saparua
    Saparua
    Saparua is an island east of Ambon Island in the Indonesian province of Maluku. The main port is in the south at Kota Saparua.Saparua was the location of Indonesian national hero, Pattimura's rebellion against Dutch forces.Paperu is a village on the island....

     and Ambon
    Ambon Island
    Ambon Island is part of the Maluku Islands of Indonesia. The island has an area of , and is mountainous, well watered, and fertile. Ambon Island consists of 2 territories: The main city and seaport is Ambon , which is also the capital of Maluku province and Maluku Tengah Ambon Island is part of the...

    , Indonesia)
  • O. p. pronomus (southern islands of Torres Straight, Queensland, Australia) – possibly a junior synonym of O. p. poseidon
  • O. p. sterrensis (Mt. Sterren, West Papua, Indonesia) – possibly a junior synonym of O. p. poseidon
  • O. p. teucrus (Biak
    Biak
    Biak features a tropical rainforest climate with nearly identical temperatures throughout the course of the year. The average annual temperature in the city is 27 degrees celsius, which is also generally the average temperature of each day in Biak...

     and Supiori
    Supiori
    Supiori is an island just west of Biak in the Schouten Islands in Papua province, Indonesia. It has a rugged surface with highest point at 1034 metres. Its area is 659 square kilometres....

    , West Papua, Indonesia)
  • O. p. wituensis (Vitu Islands
    Vitu Islands
    The Vitu Islands are a volcanic group with an area of 37 sq mi located in the Bismarck Archipelago of Papua New Guinea in the South West Pacific....

    , Papua New Guinea) – possibly a junior synonym of O. p. bornemanni
  • O. p. urvillianus (New Ireland
    New Ireland (island)
    New Ireland is a large island in Papua New Guinea, approximately 7,404 km² in area. It is the largest island of the New Ireland Province, lying northeast of the island of New Britain. Both islands are part of the Bismarck Archipelago, named after Otto von Bismarck, and they are separated by...

     and Bougainville
    Bougainville Island
    Bougainville Island is the main island of the Autonomous Region of Bougainville of Papua New Guinea. This region is also known as Bougainville Province or the North Solomons. The population of the province is 175,160 , which includes the adjacent island of Buka and assorted outlying islands...

    , Papua New Guinea, and Solomon Islands
    Solomon Islands
    Solomon Islands is a sovereign state in Oceania, east of Papua New Guinea, consisting of nearly one thousand islands. It covers a land mass of . The capital, Honiara, is located on the island of Guadalcanal...

    )


Several species currently recognised as distinct have previously been considered subspecies of O. priamus by different authors. These are O. aesacus
Ornithoptera aesacus
Ornithoptera aesacus is a species of birdwing butterfly endemic to Indonesia.It is only found on the heavily logged island of Obi. It is seriously endangered but commercially bred.-References:...

, O. croesus
Ornithoptera croesus
Ornithoptera croesus, commonly known as Wallace's Golden Birdwing is a species of birdwing butterfly found in northern Maluku.It is a member of the Ornithoptera priamus species-group which, including croesus, is only found east of Weber's Line . The larval foodplant is Pararistolochia...

, O. euphorion and O. richmondia
Ornithoptera richmondia
The Richmond Birdwing is a species of birdwing butterfly that is endemic to Australia. It is the second smallest of the birdwing species, the smallest being Ornithoptera meridionalis.-Distribution:...

. The last two are still regarded as subspecies of O. priamus by some.

Threats

Overall this species remains widespread, but some subspecies are threatened by habitat destruction, with those endemic to smaller islands (e.g. O. priamus miokensis and O. priamus boisduvali) of greatest conservation concern. Like all birdwing butterflies, O. priamus is listed on CITES, which restricts international trade.
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