Leiurus
Encyclopedia
Leiurus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of scorpion belonging to the family Buthidae
Buthidae
Buthidae is the largest family of scorpions, containing about 80 genera and over 800 species as of mid-2008. Its members are known as, for example, thick-tailed scorpions and bark scorpions. There are a few very large genera , but also a high number of species-poor or monotypic ones...

. The most common species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

, L. quinquestriatus, is also known under the vernacular name Deathstalker. It is distributed widely across North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

 and the Middle East
Middle East
The Middle East is a region that encompasses Western Asia and Northern Africa. It is often used as a synonym for Near East, in opposition to Far East...

, including the western and southern Arabian Peninsula
Arabian Peninsula
The Arabian Peninsula is a land mass situated north-east of Africa. Also known as Arabia or the Arabian subcontinent, it is the world's largest peninsula and covers 3,237,500 km2...

 and southeastern Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

. At least one species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 occurs in West Africa
West Africa
West Africa or Western Africa is the westernmost region of the African continent. Geopolitically, the UN definition of Western Africa includes the following 16 countries and an area of approximately 5 million square km:-Flags of West Africa:...

 (northern Cameroon
Cameroon
Cameroon, officially the Republic of Cameroon , is a country in west Central Africa. It is bordered by Nigeria to the west; Chad to the northeast; the Central African Republic to the east; and Equatorial Guinea, Gabon, and the Republic of the Congo to the south. Cameroon's coastline lies on the...

).

Taxonomy

The genus was introduced in 1828 by C.G. Ehrenberg
Christian Gottfried Ehrenberg
Christian Gottfried Ehrenberg , German naturalist, zoologist, comparative anatomist, geologist, and microscopist, was one of the most famous and productive scientists of his time.- Early collections :...

 (in Hemprich & Ehrenberg 1828), originally as a subgenus
Subgenus
In biology, a subgenus is a taxonomic rank directly below genus.In zoology, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the generic name and the specific epithet: e.g. the Tiger Cowry of the Indo-Pacific, Cypraea tigris Linnaeus, which...

 of the genus Androctonus. It was finally elevated to genus rank by M. Vachon in 1949.
The genus was long considered to be monotypic
Monotypic
In biology, a monotypic taxon is a taxonomic group with only one biological type. The term's usage differs slightly between botany and zoology. The term monotypic has a separate use in conservation biology, monotypic habitat, regarding species habitat conversion eliminating biodiversity and...

, containing a single species, L. quinquestriatus, but research since 2002 has shown that there are indeed several species.

Diversity

Currently five species are recognized within this genus, but their validity is under discussion. F. Kovařík (2007) suspected that L. jordanensis and L. savanicola are possible synonym
Synonym
Synonyms are different words with almost identical or similar meanings. Words that are synonyms are said to be synonymous, and the state of being a synonym is called synonymy. The word comes from Ancient Greek syn and onoma . The words car and automobile are synonyms...

s of L. quinquestriatus.
  • Leiurus abdullahbayrami Yagmur, Koc & Kunt, 2009
  • Leiurus jordanensis Lourenço, Modry & Amr, 2002
  • Leiurus nasheri Kovařík, 2007
  • Leiurus quinquestriatus (Ehrenberg, 1828) (type species
    Type species
    In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

    )
  • Leiurus savanicola Lourenço, Qi & Cloudsley-Thompson, 2006

General characteristics

Members of Leiurus are generally moderately sized scorpions that show a typical buthid
Buthidae
Buthidae is the largest family of scorpions, containing about 80 genera and over 800 species as of mid-2008. Its members are known as, for example, thick-tailed scorpions and bark scorpions. There are a few very large genera , but also a high number of species-poor or monotypic ones...

 habitus
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 with gracile pedipalp
Pedipalp
Pedipalps , are the second pair of appendages of the prosoma in the subphylum Chelicerata. They are traditionally thought to be homologous with mandibles in Crustacea and insects, although more recent studies Pedipalps (commonly shortened to palps or palpi), are the second pair of appendages of the...

 chelae and a slender metasoma
Metasoma
The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circulatory system, and the apical segments are typically...

. The vesicle is bulbous and proportionally large in some species. The cephalothorax
Cephalothorax
The cephalothorax is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. The word cephalothorax is derived from the Greek words for head and thorax...

 and mesosoma
Mesosoma
The mesosoma is the middle part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the metasoma. It bears the legs, and, in the case of winged insects, the wings....

 shows distinct granulation. Characteristically the tergites of the mesosoma bear five distinct, longitudinal carinae (ridges). The base color is generally yellow with brown to blackish areas extending over various parts of the animal, depending on species.

Toxicity

The venom of L. quinquestriatus is among the most potent scorpion toxin
Toxin
A toxin is a poisonous substance produced within living cells or organisms; man-made substances created by artificial processes are thus excluded...

s. It severely affects the cardiac and pulmonary systems. Human
Human
Humans are the only living species in the Homo genus...

 fatalities, often children, have been confirmed by clinical reports. The median lethal dose of venom (LD50) for this species was measured at 0.16 - 0.50 mg/kgmice.

The toxicity of the other species is also potentially high to life-threatening, but reliable data are currently not available.

Habitat

Most species live in semi-arid
Semi-arid
A semi-arid climate or steppe climate describes climatic regions that receive precipitation below potential evapotranspiration, but not extremely...

 to arid
Arid
A region is said to be arid when it is characterized by a severe lack of available water, to the extent of hindering or even preventing the growth and development of plant and animal life...

 regions, including the Sahara
Sahara
The Sahara is the world's second largest desert, after Antarctica. At over , it covers most of Northern Africa, making it almost as large as Europe or the United States. The Sahara stretches from the Red Sea, including parts of the Mediterranean coasts, to the outskirts of the Atlantic Ocean...

 and Arabian deserts. At least one species occurs in savannah
Savannah
Savannah or savanna is a type of grassland.It can also mean:-People:* Savannah King, a Canadian freestyle swimmer* Savannah Outen, a singer who gained popularity on You Tube...

environment. Sparsely vegetated and sandy or rocky substrates are preferred. The scorpions live in shallow burrows in sand or beneath rocks.

In captivity

Members of the genus Leiurus are often bred in captivity and traded. Due to their extreme toxicity, keeping these species is strictly recommendable to very experienced and/or professionally trained people.

External links

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