Hillia
Encyclopedia
Hillia is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

s in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Rubiaceae
Rubiaceae
The Rubiaceae is a family of flowering plants, variously called the coffee family, madder family, or bedstraw family. The group contains many commonly known plants, including the economically important coffee , quinine , and gambier , and the horticulturally valuable madder , west indian jasmine ,...

. It has 24 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

. All are indigenous
Indigenous (ecology)
In biogeography, a species is defined as native to a given region or ecosystem if its presence in that region is the result of only natural processes, with no human intervention. Every natural organism has its own natural range of distribution in which it is regarded as native...

 to tropical America
Americas
The Americas, or America , are lands in the Western hemisphere, also known as the New World. In English, the plural form the Americas is often used to refer to the landmasses of North America and South America with their associated islands and regions, while the singular form America is primarily...

.

Most of the species are slightly succulent
Succulent plant
Succulent plants, also known as succulents or fat plants, are water-retaining plants adapted to arid climates or soil conditions. Succulent plants store water in their leaves, stems, and also in roots...

 epiphyte
Epiphyte
An epiphyte is a plant that grows upon another plant non-parasitically or sometimes upon some other object , derives its moisture and nutrients from the air and rain and sometimes from debris accumulating around it, and is found in the temperate zone and in the...

s or small tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

s. A few are subshrub
Subshrub
A subshrub or dwarf shrub is a short woody plant. Prostrate shrub is a similar term.It is distinguished from a shrub by its ground-hugging stems and lower height, with overwintering perennial woody growth typically less than 10–20 cm tall, or by being only weakly woody and/or persisting...

s or liana
Liana
A liana is any of various long-stemmed, woody vines that are rooted in the soil at ground level and use trees, as well as other means of vertical support, to climb up to the canopy to get access to well-lit areas of the forest. Lianas are especially characteristic of tropical moist deciduous...

s. Hillia rivalis is a rheophyte
Rheophyte
A rheophyte is an aquatic plant that lives in fast moving water currents in an environment where few other organisms can survive. Rheophytes tend to be found in currents that move at rates of 1 to 2 meters per second and that are up to 3 to 6 feet deep. The amount of force such fast moving currents...

. The tissues
Tissue (biology)
Tissue is a cellular organizational level intermediate between cells and a complete organism. A tissue is an ensemble of cells, not necessarily identical, but from the same origin, that together carry out a specific function. These are called tissues because of their identical functioning...

 of all the species contain raphides. The capsules
Capsule (fruit)
In botany a capsule is a type of simple, dry fruit produced by many species of flowering plants. A capsule is a structure composed of two or more carpels that in most cases is dehiscent, i.e. at maturity, it splits apart to release the seeds within. A few capsules are indehiscent, for example...

 have a beak-like appendage.

Hillia triflora is cultivated
Gardening
Gardening is the practice of growing and cultivating plants. Ornamental plants are normally grown for their flowers, foliage, or overall appearance; useful plants are grown for consumption , for their dyes, or for medicinal or cosmetic use...

 as an ornamental
Ornamental plant
Ornamental plants are plants that are grown for decorative purposes in gardens and landscape design projects, as house plants, for cut flowers and specimen display...

.

Hillia was named
Botanical name
A botanical name is a formal scientific name conforming to the International Code of Botanical Nomenclature and, if it concerns a plant cultigen, the additional cultivar and/or Group epithets must conform to the International Code of Nomenclature for Cultivated Plants...

 by Nicolaus Jacquin
Nikolaus Joseph von Jacquin
Nikolaus Joseph Freiherr von Jacquin or Baron Nikolaus von Jacquin. was a scientist who studied medicine, chemistry and botany....

 in 1760. It was named for the English
English people
The English are a nation and ethnic group native to England, who speak English. The English identity is of early mediaeval origin, when they were known in Old English as the Anglecynn. England is now a country of the United Kingdom, and the majority of English people in England are British Citizens...

 botanist  John Hill
John Hill (author)
John Hill , called because of his Swedish honours, "Sir" John Hill, was an English author and botanist. He contributed to contemporary periodicals and was awarded the title of Sir in recognition of his illustrated botanical compendium The Vegetable System.He was the son of the Rev. Theophilus Hill...

 (1716-1775). Jacquin named only one species, Hillia parasitica. It has been suggested that the specific epithet as well as the generic name might well be a reference to John Hill.

Some authors have placed five of the species in a separate
Segregate (taxonomy)
In taxonomy, a segregate, or a segregate taxon is created when a taxon is split off, from another taxon. This other taxon will be better known, usually bigger, and will continue to exist, even after the segregate taxon has been split off...

 genus, Ravnia. A cladistic analysis of morphological
Plant morphology
Plant morphology or phytomorphology is the study of the physical form and external structure of plants. This is usually considered distinct from plant anatomy, which is the study of the internal structure of plants, especially at the microscopic level...

  characters found Ravnia to be embedded
Paraphyly
A group of taxa is said to be paraphyletic if the group consists of all the descendants of a hypothetical closest common ancestor minus one or more monophyletic groups of descendants...

 within Hillia. This hypothesis
Hypothesis
A hypothesis is a proposed explanation for a phenomenon. The term derives from the Greek, ὑποτιθέναι – hypotithenai meaning "to put under" or "to suppose". For a hypothesis to be put forward as a scientific hypothesis, the scientific method requires that one can test it...

 has not been tested
Statistical hypothesis testing
A statistical hypothesis test is a method of making decisions using data, whether from a controlled experiment or an observational study . In statistics, a result is called statistically significant if it is unlikely to have occurred by chance alone, according to a pre-determined threshold...

 with molecular data.

The genera Hillia, Balmea
Balmea
Balmea is a monospecific genus of flowering plants in the Rubiaceae family. The genus contains only one species, i.e. Balmea stormae, which is found in El Salvador, Guatemala and Mexico. It is locally known as ayuque. The plant is picked for its scarlet flowers, and the whole plant is sold on...

, and Cosmibuena form a monophyletic group
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

. Some authors have designated this group as the tribe
Tribe (biology)
In biology, a tribe is a taxonomic rank between family and genus. It is sometimes subdivided into subtribes.Some examples include the tribes: Canini, Acalypheae, Hominini, Bombini, and Antidesmeae.-See also:* Biological classification* Rank...

 Hillieae, but it might be embedded within another tribe, Hamelieae.

Species

The following species list may be incomplete or contain synonyms
Synonym (taxonomy)
In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

.
  • Hillia macrophylla Standley
    Paul Carpenter Standley
    Paul Carpenter Standley was an American botanist.Standley was born in Avalon, Missouri...

  • Hillia parasitica  Jacquin
    Nikolaus Joseph von Jacquin
    Nikolaus Joseph Freiherr von Jacquin or Baron Nikolaus von Jacquin. was a scientist who studied medicine, chemistry and botany....

       
  • Hillia rivalis C.M. Taylor
  • Hillia triflora ( Oersted
    Anders Sandøe Ørsted (botanist)
    Anders Sandøe Ørsted, also written as Anders Sandoe Oersted or Anders Sandö Örsted was a Danish botanist, mycologist, zoologist and marine biologist. He was nephew of the brothers physicist H.C...

    ) C.M. Taylor

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