Hieroglyphus daganensis
Encyclopedia
The African rice grasshopper, Hieroglyphus daganensis is a medium-sized grasshopper
Grasshopper
The grasshopper is an insect of the suborder Caelifera in the order Orthoptera. To distinguish it from bush crickets or katydids, it is sometimes referred to as the short-horned grasshopper...

 species found in the Sahel
Sahel
The Sahel is the ecoclimatic and biogeographic zone of transition between the Sahara desert in the North and the Sudanian Savannas in the south.It stretches across the North African continent between the Atlantic Ocean and the Red Sea....

 region. Although not called a locust
Locust
Locusts are the swarming phase of short-horned grasshoppers of the family Acrididae. These are species that can breed rapidly under suitable conditions and subsequently become gregarious and migratory...

 in English, this species shows gregarious behaviour and some morphological change (phase polymorphism
Polymorphism (biology)
Polymorphism in biology occurs when two or more clearly different phenotypes exist in the same population of a species — in other words, the occurrence of more than one form or morph...

) on crowding and may become a moderately important pest species for small-holder farmers in the region.

Adults

The overall colour of adult insects is usually green, with a finely dotted shiny integument. Antennae are longer than the head and pronotum together. The pronotum is cylindrical, with three deep and wide sulci (grooves) crossing the dorsum
Dorsum
Dorsum is a Latin word. In science, it could mean:* Dorsum , the posterior side of an animal* Dorsum , a term used in astrogeology for a ridge* Theta Capricorni, a star on the back of the Goat...

.

Hoppers

The colour of immature insects may vary depending on their phase: in the illustration above, during a particularly fertile year, dense populations of H. daganensis showed a distinct blackening in certain areas of cuticle
Cuticle
A cuticle , or cuticula, is a term used for any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticles" are non-homologous; differing in their origin, structure, function, and chemical composition...

.

Outbreaks and Control

Outbreaks are frequent in Sahel
Sahel
The Sahel is the ecoclimatic and biogeographic zone of transition between the Sahara desert in the North and the Sudanian Savannas in the south.It stretches across the North African continent between the Atlantic Ocean and the Red Sea....

ian countries such as Senegal
Senegal
Senegal , officially the Republic of Senegal , is a country in western Africa. It owes its name to the Sénégal River that borders it to the east and north...

, Mauritania
Mauritania
Mauritania is a country in the Maghreb and West Africa. It is bordered by the Atlantic Ocean in the west, by Western Sahara in the north, by Algeria in the northeast, by Mali in the east and southeast, and by Senegal in the southwest...

, Burkina Faso
Burkina Faso
Burkina Faso – also known by its short-form name Burkina – is a landlocked country in west Africa. It is surrounded by six countries: Mali to the north, Niger to the east, Benin to the southeast, Togo and Ghana to the south, and Côte d'Ivoire to the southwest.Its size is with an estimated...

, Mali
Mali
Mali , officially the Republic of Mali , is a landlocked country in Western Africa. Mali borders Algeria on the north, Niger on the east, Burkina Faso and the Côte d'Ivoire on the south, Guinea on the south-west, and Senegal and Mauritania on the west. Its size is just over 1,240,000 km² with...

, Niger
Niger
Niger , officially named the Republic of Niger, is a landlocked country in Western Africa, named after the Niger River. It borders Nigeria and Benin to the south, Burkina Faso and Mali to the west, Algeria and Libya to the north and Chad to the east...

 and Benin Benin
Benin
Benin , officially the Republic of Benin, is a country in West Africa. It borders Togo to the west, Nigeria to the east and Burkina Faso and Niger to the north. Its small southern coastline on the Bight of Benin is where a majority of the population is located...

.

A biological pesticide product called 'Green Muscle' is based on the entomopathogenic fungus
Entomopathogenic fungus
An entomopathogenic fungus is a fungus that can act as a parasite of insects and kills or seriously disables them.-Typical life cycle:These fungi usually attach to the external body surface of insects in the form of microscopic spores...

 (Metarhizium acridum
Metarhizium acridum
Metarhizium acridum is the new name given to a group of fungal isolates that are known to be virulent and specific to Acrididea. Previously, this species has had variety status in Metarhizium anisopliae and before that, reference was made to M. flavoviride.- Biology :M...

) is now available (see desert locust
Desert locust
Plagues of the desert locust have threatened agricultural production in Africa, the Middle East, and Asia for centuries. The livelihood of at least one-tenth of the world’s human population can be affected by this voracious insect...

). Early demonstrations of efficacy took place in northern Benin
Benin
Benin , officially the Republic of Benin, is a country in West Africa. It borders Togo to the west, Nigeria to the east and Burkina Faso and Niger to the north. Its small southern coastline on the Bight of Benin is where a majority of the population is located...

, with field tests by the LUBILOSA
LUBILOSA
LUBILOSA was the name of a research programme that aimed at developing a biological alternative to the chemical control of locusts. This name is an acronym of the French title of the programme: Lutte Biologique contre les Locustes et les Sauteriaux...

Programme on H. daganensis.
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