Excretory system of gastropods
Encyclopedia
The excretory system of gastropods removes nitrogenous waste and maintains the internal water balance of these creatures, commonly referred to as snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

s and slug
Slug
Slug is a common name that is normally applied to any gastropod mollusc that lacks a shell, has a very reduced shell, or has a small internal shell...

s. The primary organ of excretion is a nephridium
Nephridium
A Nephridium is an invertebrate organ which occurs in pairs and function similar to kidneys. Nephridia remove metabolic wastes from an animal's body. They are present in many different invertebrate lines. There are two basic types, metanephridia and protonephridia, but there are other...

.

Structure

The most primitive gastropods retain two nephridia, but in the great majority of species, the right nephridium has been lost, leaving a single excretory organ, located in the anterior part of the visceral mass. The nephridium projects into the main venous sinus in the animal's foot. The circulatory fluid of gastropods, known as haemolymph directly bathes the tissues, where it supplies them with oxygen and absorbs carbon dioxide and nitrogenous waste, a necessary waste product of metabolism
Metabolism
Metabolism is the set of chemical reactions that happen in the cells of living organisms to sustain life. These processes allow organisms to grow and reproduce, maintain their structures, and respond to their environments. Metabolism is usually divided into two categories...

. From the arterial sinuses bathing the tissues, it drains into the venous sinus, and thus flows past the nephridium.

The main body cavity of most aquatic gastropods also includes pericardial glands, often located above the heart. These secrete waste into the haemolymph, prior to further filtration in the nephridium. Pulmonates lack these glands, so that the nephridium is the only major organ of excretion.

In some gastropods, the nephridium opens directly into the sinus, but more usually, there is a small duct, referred to as the renopericardial canal. In aquatic gastropods, the nephridium is drained by a ureter
Ureter
In human anatomy, the ureters are muscular tubes that propel urine from the kidneys to the urinary bladder. In the adult, the ureters are usually long and ~3-4 mm in diameter....

 that opens near the rear of the mantle cavity. This allows the flow of water through the cavity to flush out the excreta. Terrestrial pulmonates instead have a much longer ureter, that opens near the anus
Anus
The anus is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, unwanted semi-solid matter produced during digestion, which, depending on the type of animal, may be one or more of: matter which the animal cannot digest,...

.

In addition to the pericardial glands and nephridum, excretory cells are also present in the digestive glands opening into the stomach
Stomach
The stomach is a muscular, hollow, dilated part of the alimentary canal which functions as an important organ of the digestive tract in some animals, including vertebrates, echinoderms, insects , and molluscs. It is involved in the second phase of digestion, following mastication .The stomach is...

. These glands have a metabolic function, somewhat similar to that of the vertebrate liver
Liver
The liver is a vital organ present in vertebrates and some other animals. It has a wide range of functions, including detoxification, protein synthesis, and production of biochemicals necessary for digestion...

, and excrete waste products directly into the digestive system, where it is voided with the faeces.

Water balance

In aquatic gastropods, the waste product is invariably ammonia
Ammonia
Ammonia is a compound of nitrogen and hydrogen with the formula . It is a colourless gas with a characteristic pungent odour. Ammonia contributes significantly to the nutritional needs of terrestrial organisms by serving as a precursor to food and fertilizers. Ammonia, either directly or...

, which rapidly dissolves in the surrounding water. In the case of freshwater species, the nephridium also resorbs a significant amount of salt in order to prevent its loss through osmosis
Osmosis
Osmosis is the movement of solvent molecules through a selectively permeable membrane into a region of higher solute concentration, aiming to equalize the solute concentrations on the two sides...

 into the surrounding water.

Terrestrial species instead excrete insoluble uric acid
Uric acid
Uric acid is a heterocyclic compound of carbon, nitrogen, oxygen, and hydrogen with the formula C5H4N4O3. It forms ions and salts known as urates and acid urates such as ammonium acid urate. Uric acid is created when the body breaks down purine nucleotides. High blood concentrations of uric acid...

, which allows them to maintain their internal water balance. Even so, most species require a somewhat humid environment, and secrete a considerable amount of water in their slime trail. Those few species that dwell in arid environments typically hibernate
Hibernation
Hibernation is a state of inactivity and metabolic depression in animals, characterized by lower body temperature, slower breathing, and lower metabolic rate. Hibernating animals conserve food, especially during winter when food supplies are limited, tapping energy reserves, body fat, at a slow rate...

 or aestivate during dry periods to preserve moisture.
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