Eriphia verrucosa, sometimes called the warty crab or yellow crab, is a species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of crab

Crab

True crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" , or where the reduced abdomen is entirely hidden under the thorax...

 found in the Black Sea

Black Sea

The Black Sea is bounded by Europe, Anatolia and the Caucasus and is ultimately connected to the Atlantic Ocean via the Mediterranean and the Aegean seas and various straits. The Bosphorus strait connects it to the Sea of Marmara, and the strait of the Dardanelles connects that sea to the Aegean...

, Mediterranean Sea

Mediterranean Sea

The Mediterranean Sea is a sea connected to the Atlantic Ocean surrounded by the Mediterranean region and almost completely enclosed by land: on the north by Anatolia and Europe, on the south by North Africa, and on the east by the Levant...

 and eastern Atlantic Ocean

Atlantic Ocean

The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 from Brittany

Brittany

Brittany is a cultural and administrative region in the north-west of France. Previously a kingdom and then a duchy, Brittany was united to the Kingdom of France in 1532 as a province. Brittany has also been referred to as Less, Lesser or Little Britain...

 to Mauritania

Mauritania

Mauritania is a country in the Maghreb and West Africa. It is bordered by the Atlantic Ocean in the west, by Western Sahara in the north, by Algeria in the northeast, by Mali in the east and southeast, and by Senegal in the southwest...

 and the Azores

Azores

The Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...

. Individual crabs have been caught as far north as Cornwall

Cornwall

Cornwall is a unitary authority and ceremonial county of England, within the United Kingdom. It is bordered to the north and west by the Celtic Sea, to the south by the English Channel, and to the east by the county of Devon, over the River Tamar. Cornwall has a population of , and covers an area of...

. Formerly a frequent species in the Black Sea, it has decreased in numbers since the 1980s and is now listed in the Ukrainian

Ukraine

Ukraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...

 Red Data Book

Regional Red List

A Regional Red List is a report of the threatened status of species within a certain country or region. It is based on the IUCN Red List of Threatened Species, an inventory of the conservation status of species on a global scale...

 of endangered species

Endangered species

An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

.

Ecology

E. verrucosa lives among stones and seaweed

Seaweed

Seaweed is a loose, colloquial term encompassing macroscopic, multicellular, benthic marine algae. The term includes some members of the red, brown and green algae...

s in shallow water along rocky coastlines up to a depth of 15 metres (49.2 ft). It is reported to feed on bivalves, gastropods and hermit crab

Hermit crab

Hermit crabs are decapod crustaceans of the superfamily Paguroidea. Most of the 1100 species possess an asymmetrical abdomen which is concealed in an empty gastropod shell that is carried around by the hermit crab.-Description:...

s, or on molluscs and polychaete

Polychaete

The Polychaeta or polychaetes are a class of annelid worms, generally marine. Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. Indeed, polychaetes are sometimes referred to as bristle worms. More than 10,000...

s. In the Black Sea, E. verrucosa is the only native species capable of breaking into the shells of the invasive

Invasive species

"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

 snail

Snail

Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

 Rapana venosa, although it is unlikely that it will present an effective biological control of the invader. The species is threatened by eutrophication

Eutrophication

Eutrophication or more precisely hypertrophication, is the movement of a body of water′s trophic status in the direction of increasing plant biomass, by the addition of artificial or natural substances, such as nitrates and phosphates, through fertilizers or sewage, to an aquatic system...

 and pollution

Pollution

Pollution is the introduction of contaminants into a natural environment that causes instability, disorder, harm or discomfort to the ecosystem i.e. physical systems or living organisms. Pollution can take the form of chemical substances or energy, such as noise, heat or light...

.

Description

E. verrucosa may reach a width of 9 centimetres (3.5 in) and a length of 7 cm (2.8 in). The carapace

Carapace

A carapace is a dorsal section of the exoskeleton or shell in a number of animal groups, including arthropods such as crustaceans and arachnids, as well as vertebrates such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.-Crustaceans:In crustaceans, the...

 is thick and smooth, ranging in colour from brownish-red to brownish-green, with yellow spots; its front margin is armed with seven "teeth" on either side, and five or six between the eyes. The claw

Claw

A claw is a curved, pointed appendage, found at the end of a toe or finger in most mammals, birds, and some reptiles. However, the word "claw" is also often used in reference to an invertebrate. Somewhat similar fine hooked structures are found in arthropods such as beetles and spiders, at the end...

s are strong and have black fingers; one claw is generally larger than the other and is armed with rounded tubercle

Tubercle

A tubercle is generally a wart-like projection, but it has slightly different meaning depending on which family of plants or animals it is used to refer to....

s while the smaller claw bears sharper projections, arranged in lines. In the springtime, E. verrucosa migrates to shallow water, less than 1 metre deep, and reproduction begins in May or June; the species is highly fecund. There are four larval stages

Crustacean larvae

Crustaceans may pass through a number of larval and immature stages between hatching from their eggs and reaching their adult form. Each of the stages is separated by a moult, in which the hard exoskeleton is shed to allow the animal to grow...

, from zoea to megalopa.