Coastal Giant Salamander
Encyclopedia
The Coastal Giant Salamander (Dicamptodon tenebrosus) is a species of salamander
Salamander
Salamander is a common name of approximately 500 species of amphibians. They are typically characterized by a superficially lizard-like appearance, with their slender bodies, short noses, and long tails. All known fossils and extinct species fall under the order Caudata, while sometimes the extant...

 in the Dicamptodontidae family (Pacific giant salamanders). It is endemic to the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 and Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

. There are three closely related species to this taxon: D. ensatus (California Giant Salamander
California Giant Salamander
The California Giant Salamander is a species of salamander in the Dicamptodontidae family.It is endemic to the United States. The species name once additionally included individuals now belonging to the species D. aterrimus and D...

), D. copei (Cope's Giant Salamander
Cope's Giant Salamander
Cope's Giant Salamander is a species of salamander in the Dicamptodontidae family. It reaches between 12.4–19.1 cm . The salamander resembles Pacific Giant Salamander larvae, but it never transforms to a terrestrial stage. It is smaller overall with a narrower head and shorter limbs...

), and D. aterrimus (Idaho Giant Salamander
Idaho Giant Salamander
The Idaho Giant Salamander, Dicamptodon aterrimus, is a species of salamander. There are three closely related species to this taxon: D. ensatus, , D. copei and D...

).

Its natural habitat
Habitat
* Habitat , a place where a species lives and grows*Human habitat, a place where humans live, work or play** Space habitat, a space station intended as a permanent settlement...

s are temperate forest
Forest
A forest, also referred to as a wood or the woods, is an area with a high density of trees. As with cities, depending where you are in the world, what is considered a forest may vary significantly in size and have various classification according to how and what of the forest is composed...

s, river
River
A river is a natural watercourse, usually freshwater, flowing towards an ocean, a lake, a sea, or another river. In a few cases, a river simply flows into the ground or dries up completely before reaching another body of water. Small rivers may also be called by several other names, including...

s, freshwater lake
Lake
A lake is a body of relatively still fresh or salt water of considerable size, localized in a basin, that is surrounded by land. Lakes are inland and not part of the ocean and therefore are distinct from lagoons, and are larger and deeper than ponds. Lakes can be contrasted with rivers or streams,...

s, and freshwater marsh
Marsh
In geography, a marsh, or morass, is a type of wetland that is subject to frequent or continuous flood. Typically the water is shallow and features grasses, rushes, reeds, typhas, sedges, other herbaceous plants, and moss....

es.

Pacific Giant Salamanders are protected from being killed or collected under the Wildlife Act in British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

.

Description

The adult Coastal Giant Salamander can reach up to 34 cm (13 inches) in total length. The Coastal Giant Salamander has four toes on the front feet and five toes on the back feet. The species tail is around 40 percent of the total length of the salamander and is laterally compressed. The head, back, and sides have a marbled or reticulate pattern of dark blotches on a light brown or brassy-colored background. The head is broad with a shovel-like snout and a fold of skin across the throat called the gular fold. The eye
Eye
Eyes are organs that detect light and convert it into electro-chemical impulses in neurons. The simplest photoreceptors in conscious vision connect light to movement...

s are medium in size and have a brass-flecked iris and a large black pupil. This species is one of the few salamanders capable of vocalizing.

Range

The Coastal Giant Salamander is endemic to the Pacific Northwest, found in Northern California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

, Oregon
Oregon
Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

, Washington, and southern British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

.

Neotenes

Some Coastal Giant Salamander larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

 continue to grow into an adult and become sexually mature without losing their external gill
Gill
A gill is a respiratory organ found in many aquatic organisms that extracts dissolved oxygen from water, afterward excreting carbon dioxide. The gills of some species such as hermit crabs have adapted to allow respiration on land provided they are kept moist...

s. This process is called neoteny
Neoteny
Neoteny , also called juvenilization , is one of the two ways by which paedomorphism can arise. Paedomorphism is the retention by adults of traits previously seen only in juveniles, and is a subject studied in the field of developmental biology. In neoteny, the physiological development of an...

. Neotony is particularly common in the British Columbia populations. Adult-sized neotenes have a uniform brown coloring on their heads, sides, and backs.
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