Cladonematidae
Encyclopedia
Cladonematidae is a small family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of anthomedusan
Anthomedusae
Anthomedusae or Anthomedusa, the athecate hydroids, are an order of the Hydrozoa, a class of marine invertebrates belonging to the phylum Cnidaria...

 hydrozoa
Hydrozoa
Hydrozoa are a taxonomic class of very small, predatory animals which can be solitary or colonial and which mostly live in saltwater. A few genera within this class live in freshwater...

ns. They have stolonal hydroid
Hydroid
-Marine biology:Hydroids are a life stage for most animals of class Hydrozoa, small predators related to jellyfish.-Botany:In mosses, hydroids form the innermost layer of the stem of long, colourless, thin walled cells of small diameter.The cells are dead and lack protoplasm.They function as water...

 colonies, and their medusa
Medusa
In Greek mythology Medusa , " guardian, protectress") was a Gorgon, a chthonic monster, and a daughter of Phorcys and Ceto. The author Hyginus, interposes a generation and gives Medusa another chthonic pair as parents. Gazing directly upon her would turn onlookers to stone...

e are benthic and can crawl across the sediment
Sediment
Sediment is naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of fluids such as wind, water, or ice, and/or by the force of gravity acting on the particle itself....

; in many species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 they have lost the ability to swim however.

Description

The hydranths carry a single whorl of mouth tentacle
Tentacle
A tentacle or bothrium is one of usually two or more elongated flexible organs present in animals, especially invertebrates. The term may also refer to the hairs of the leaves of some insectivorous plants. Usually, tentacles are used for feeding, feeling and grasping. Anatomically, they work like...

s which each bear a little knob at the tip; in some Cladonematidae an aboral
Aboral
In biology, aboral surfaces are surfaces away from or opposite the mouth. The term is a compound of the Latin preposition ā, a, abs, meaning from or away from and the noun ōs, ōris n., meaning mouth. It is also the opposite of oral which is the end containing the mouth of a bilaterally...

 whorl of thread-like unknobbed tentacles is also present. They have a preoral chamber formed by epidermal
Epidermis (zoology)
The Epidermis is an epithelium that covers the body of an eumetazoan . Eumetazoa have a cavity lined with a similar epithelium, the gastrodermis, which forms a boundary with the epidermis at the mouth.Sponges have no epithelium, and therefore no epidermis or gastrodermis...

 gland
Gland
A gland is an organ in an animal's body that synthesizes a substance for release of substances such as hormones or breast milk, often into the bloodstream or into cavities inside the body or its outer surface .- Types :...

 cells.

The medusa
Medusa
In Greek mythology Medusa , " guardian, protectress") was a Gorgon, a chthonic monster, and a daughter of Phorcys and Ceto. The author Hyginus, interposes a generation and gives Medusa another chthonic pair as parents. Gazing directly upon her would turn onlookers to stone...

e bud off
Budding
Budding is a form of asexual reproduction in which a new organism grows on another one. The new organism remains attached as it grows, separating from the parent organism only when it is mature. Since the reproduction is asexual, the newly created organism is a clone and is genetically identical...

 at the base of the hydrants, though where aboral tentacle are present these are located between the stolons and the budding sites. Cladonematidae medusae have a various extend of nematocysts around their umbrellar margin, varying between a continuous dense ring to none at all among the species. The number of radial canals and whether they are branched or not is variable in this family. Their marginal is branched, with nematocyst clusters on the upper and adhesive organs on the lower branches. The manubrium is cylindrical and in some has perradial pouches and/or above it a brood pouch; the gonad
Gonad
The gonad is the organ that makes gametes. The gonads in males are the testes and the gonads in females are the ovaries. The product, gametes, are haploid germ cells. For example, spermatozoon and egg cells are gametes...

s either surround the manubrium entirely or are carried on manubrial extrusions or (if present) the brood pouch. The mouth is bordered either with clusters of nematocysts, or with short lips of tissue, or fringed with branching tentacles. The tentacle bases abaxially bear ocelli.

Genera

There are a mere 3 genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

, one of which might not be valid:
  • Cladonema Dujardin, 1843
  • Dendronema Haeckel, 1879
  • Stauridium (doubtfully distinct from Cladonema)
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