Archaeotrogonidae
Encyclopedia
Archaeotrogon is a prehistoric bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

. Its remains have been found in the Quercy Phosphorites of France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

, a geological formation containing Late Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

 and Early Oligocene
Oligocene
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 34 million to 23 million years before the present . As with other older geologic periods, the rock beds that define the period are well identified but the exact dates of the start and end of the period are slightly...

 deposits. Archaeotrogon lived some 30-35 million years ago. Not all species described herein may be valid.

This genus has been placed in a distinct family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

, the Archaeotrogonidae. The Middle Eocene Hassiavis, a more recently described bird from the famous Messel Pit
Messel pit
The Messel Pit is a disused quarry near the village of Messel, about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of fossils, it has significant geological and scientific importance...

 in Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

, might also belong there. In addition, a somewhat enigmatic fossil in the M. Daniels collection, found in the Early Eocene London Clay
London Clay
The London Clay Formation is a marine geological formation of Ypresian age which crops out in the southeast of England. The London Clay is well known for the fossils it contains. The fossils from the Lower Eocene indicate a moderately warm climate, the flora being tropical or subtropical...

 appears to belong here too.

As the name implies, Archaeotrogon was initially taken to be a prehistoric trogon
Trogon
The trogons and quetzals are birds in the order Trogoniformes which contains only one family, the Trogonidae. The family contains 39 species in eight genera. The fossil record of the trogons dates back 49 million years to the mid-Eocene. They might constitute a member of the basal radiation of...

. However, it is nowadays generally believed that they are not very closely related to these tropical forest birds of our time, but rather convergent
Convergent evolution
Convergent evolution describes the acquisition of the same biological trait in unrelated lineages.The wing is a classic example of convergent evolution in action. Although their last common ancestor did not have wings, both birds and bats do, and are capable of powered flight. The wings are...

. The Archaeotrogonidae actually seem to be Cypselomorphae
Cypselomorphae
Cypselomorphae is a clade of birds. It includes the living families and orders Caprimulgidae , Nyctibiidae , Apodiformes , as well as the Aegotheliformes whose distinctness was only recently realized...

 and related to nightjar
Nightjar
Nightjars are medium-sized nocturnal or crepuscular birds with long wings, short legs and very short bills. They are sometimes referred to as goatsuckers from the mistaken belief that they suck milk from goats . Some New World species are named as nighthawks...

s and hummingbird
Hummingbird
Hummingbirds are birds that comprise the family Trochilidae. They are among the smallest of birds, most species measuring in the 7.5–13 cm range. Indeed, the smallest extant bird species is a hummingbird, the 5-cm Bee Hummingbird. They can hover in mid-air by rapidly flapping their wings...

s, either as a basal lineage or as a distinct but entirely extinct family. The latter might be more justified than with other indeterminate Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...

 and Paleogene
Paleogene
The Paleogene is a geologic period and system that began 65.5 ± 0.3 and ended 23.03 ± 0.05 million years ago and comprises the first part of the Cenozoic Era...

 modern birds: Archaeotrogon is known from a time when the living cypselomorph families were already distinct, yet appears as well highly autapomorphic and the archaeotrogonid lineage seems to go as far back as that of nightjars for example.
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