The Silurian

Silurian

The Silurian is a geologic period and system that extends from the end of the Ordovician Period, about 443.7 ± 1.5 Mya , to the beginning of the Devonian Period, about 416.0 ± 2.8 Mya . As with other geologic periods, the rock beds that define the period's start and end are well identified, but the...

 lagerstätte

Lagerstätte

A Lagerstätte is a sedimentary deposit that exhibits extraordinary fossil richness or completeness.Palaeontologists distinguish two kinds....

 preserved in the limestone

Limestone

Limestone is a sedimentary rock composed largely of the minerals calcite and aragonite, which are different crystal forms of calcium carbonate . Many limestones are composed from skeletal fragments of marine organisms such as coral or foraminifera....

 Wenlock Series of Herefordshire

Herefordshire

Herefordshire is a historic and ceremonial county in the West Midlands region of England. For Eurostat purposes it is a NUTS 3 region and is one of three counties that comprise the "Herefordshire, Worcestershire and Gloucestershire" NUTS 2 region. It also forms a unitary district known as the...

, England

England

England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

, offers paleontologists a rare snapshot of a moment in time, about 420 Mya. In the formation, layers of fine-grained volcanic ash punctuate a sequence of carbonate muds that were accumulating in a marine environment on the outer continental shelf

Continental shelf

The continental shelf is the extended perimeter of each continent and associated coastal plain. Much of the shelf was exposed during glacial periods, but is now submerged under relatively shallow seas and gulfs, and was similarly submerged during other interglacial periods. The continental margin,...

. In this fine-grained matrix, soft-bodied animals

Soft-bodied organisms

The term "soft-bodied organism" is used to describe animals without skeleton, roughly corresponding to the group Vermes as proposed by Carl von Linné. All animals have muscles, but since muscles can only pull, never push, a number of animals have developed hard parts that the muscles can pull on,...

 and delicate, lightly sclerotized chitin

Chitin

Chitin n is a long-chain polymer of a N-acetylglucosamine, a derivative of glucose, and is found in many places throughout the natural world...

ous shells are often preserved in three dimensions, as calcitic fossilizations within calcareous nodules. Calcitic fossilization is an unusual feature.

The lagerstätte, discovered and first published in 1996, provides a wider representation of organisms than conventional fossilizations of shelly and bony elements. The Wenlock Series offers a diverse macrofauna that includes polychaete worms

Polychaete

The Polychaeta or polychaetes are a class of annelid worms, generally marine. Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. Indeed, polychaetes are sometimes referred to as bristle worms. More than 10,000...

 (Kenostrychus), sponge

Sea sponge

Sponges are animals of the phylum Porifera . Their bodies consist of jelly-like mesohyl sandwiched between two thin layers of cells. While all animals have unspecialized cells that can transform into specialized cells, sponges are unique in having some specialized cells, but can also have...

s, graptolite

Graptolite

Graptolithina is a class in the animal phylum Hemichordata, the members of which are known as Graptolites. Graptolites are fossil colonial animals known chiefly from the Upper Cambrian through the Lower Carboniferous...

s, a chelicerate (Offacolus) and a vermiform mollusc (Acaenoplax). On the microscopic scale the diverse microfauna includes abundant well-preserved radiolarian

Radiolarian

Radiolarians are amoeboid protozoa that produce intricate mineral skeletons, typically with a central capsule dividing the cell into inner and outer portions, called endoplasm and ectoplasm. They are found as zooplankton throughout the ocean, and their skeletal remains cover large portions of the...

s.

The delicate Wenlock fossils are difficult to separate from split sections, so Mark Sutton and his team have devised a method of serially grinding sections; from digital photographs three-dimensional "digital fossils" are reconstructed from datasets.