Theridion grallator
Encyclopedia
Theridion grallator, also known as the "happy face spider
Spider
Spiders are air-breathing arthropods that have eight legs, and chelicerae with fangs that inject venom. They are the largest order of arachnids and rank seventh in total species diversity among all other groups of organisms...

," is a member of the Theridiidae
Theridiidae
Theridiidae is a large family of spiders, also known as the tangle-web spiders, cobweb spiders and comb-footed spiders. The diverse family includes over 2200 species in over 100 genera) of three-dimensional space-web-builders found throughout the world...

 family.

The Hawaiian
Hawaiian language
The Hawaiian language is a Polynesian language that takes its name from Hawaii, the largest island in the tropical North Pacific archipelago where it developed. Hawaiian, along with English, is an official language of the state of Hawaii...

 name is nananana makakii (face-patterned spider). The binomial
Binomial
In algebra, a binomial is a polynomial with two terms —the sum of two monomials—often bound by parenthesis or brackets when operated upon...

 grallator is Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 for "stiltwalker", reference to the species' long spindly legs.

Appearance

The spider is about five milllimeters long overall.

Certain morphs have a pattern uncannily resembling a smiley
Smiley
A smiley, smiley face, or happy face, is a stylized representation of a smiling human face, commonly occurring in popular culture. It is commonly represented as a yellow circle with two black dots representing eyes and a black arc representing the mouth...

 face or a grinning clown face on their yellow body. Each spider has a unique pattern, and the patterns differ from island to island. Some lack markings altogether.
On the island of Maui, the happy types seem to follow simple Mendelian inheritance
Mendelian inheritance
Mendelian inheritance is a scientific description of how hereditary characteristics are passed from parent organisms to their offspring; it underlies much of genetics...

 rules, while on other Hawaiian islands the body inheritance patterns seem to be sex-limited. http://www.benecke.com/happyface.html The variation is possibly a kind of camouflage against birds, their only natural enemies of significance, to counteract pattern recognition by predators. As the pattern may change according to what food the spider has eaten (Gillespie, 1989) and as T. grallator is very small, hides during the day, and is thus not a significant prey item for any species of predator, it is more likely that the bizarre variety of patterns serves no significant adaptive
Adaptation
An adaptation in biology is a trait with a current functional role in the life history of an organism that is maintained and evolved by means of natural selection. An adaptation refers to both the current state of being adapted and to the dynamic evolutionary process that leads to the adaptation....

 purpose at all.

Habits

They live beneath the leaves of plants, where they also spin their much reduced webs. The females guard their eggs until they hatch, and catch prey for their young.
T. grallator hunts mainly during the evening.

Distribution

T. grallator is endemic to Oahu
Oahu
Oahu or Oahu , known as "The Gathering Place", is the third largest of the Hawaiian Islands and most populous of the islands in the U.S. state of Hawaii. The state capital Honolulu is located on the southeast coast...

, Molokai
Molokai
Molokai or Molokai is an island in the Hawaiian archipelago. It is 38 by 10 miles in size with a land area of , making it the fifth largest of the main Hawaiian Islands and the 27th largest island in the United States. It lies east of Oahu across the 25-mile wide Kaiwi Channel and north of...

, Maui
Maui
The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

 and the Big Island
Hawaii (island)
The Island of Hawaii, also called the Big Island or Hawaii Island , is a volcanic island in the North Pacific Ocean...

 in rainforests at elevations of 300 to 2,000 meters.

External links

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