Taxonomy of Lucanidae
Encyclopedia
The classification presented here follows Smith (2006), with the exception of the tribal classification within the Lucaninae.

Subfamily Aesalinae
Tribe Aesalini
Genus Aesalus (Fabricius
Johan Christian Fabricius
Johan Christian Fabricius was a Danish zoologist, specialising in "Insecta", which at that time included all arthropods: insects, arachnids, crustaceans and others...

, 1801)
  • Aesalus asiaticus Lewis, 1883
  • Aesalus himalayicus Kurosawa, 1985
  • Aesalus imanishii Inahara & Ratti, 1991
  • Aesalus meridionalis Bartolozzi, 1989
  • Aesalus neotropicalis Bates, 1886
  • Aesalus saburoi
  • Aesalus sawaii Fujita & Ichikawa, 1985
  • Aesalus scarabaeoides Panzer, 1793
  • Aesalus sichuanensis
  • Aesalus smithi Bates, 1889
  • Aesalus trogoides Albers, 1883
  • Aesalus ulanowskii Ganglbauer, 1886
Genus Cretaesalus (fossil)
Genus Echinoaesalus Zelenka, 1993
  • Echinoaesalus barriesi (Zelenka, 1993)
  • Echinoaesalus hidakai (Araya, 1993)
  • Echinoaesalus jaechi (Zelenka,1993)
  • Echinoaesalus matsuii (Araya, 1993)
  • Echinoaesalus sabahensis (Zelenka, 1994)
  • Echinoaesalus schuhi (Zelenka, 1994)
  • Echinoaesalus timidus (Krikken, 1975)
  • Echinoaesalus yongi (Araya, 1993)
Genus Lucanobium Howden & Lawrence, 1974
  • Lucanobium squamosum Howden & Lawrence, 1974
Tribe Ceratognathini
Genus Ceratognathus Westwood, 1838
  • Ceratognathus abdominalis Parry, 1870
  • Ceratognathus bitumulatus Carter, 1925
  • Ceratognathus flabellatus Boileau, 1905
  • Ceratognathus frenchi Blackburn, 1897
  • Ceratognathus froggatti Blackburn, 1894
  • Ceratognathus gilesi Blackburn, 1895
  • Ceratognathus macrognathus Boileau, 1905
  • Ceratognathus mentiferus Westwood, 1863
  • Ceratognathus minutus Lea, 1929
  • Ceratognathus niger Westwood, 1838
  • Ceratognathus ocularis Carter, 1925
  • Ceratognathus rufipennis Westwood, 1872
  • Ceratognathus tasmanus Benesh, 1943
  • Ceratognathus westwoodi Thomson, 1862
Genus Hilophyllus Paulsen & Mondaca, 2006
  • Hilophyllus argentinensis (Martinez, 1981)
  • Hilophyllus martinezi Paulsen & Mondaca, 2006
  • Hilophyllus penai (Martinez, 1976)
Genus Mitophyllus Parry, 1843
  • Mitophyllus alboguttatus (Bates, 1867)
  • Mitophyllus dispar (Sharp, 1882)
  • Mitophyllus foveolatus (Broun, 1880)
  • Mitophyllus gibbosus (Broun, 1884)
  • Mitophyllus insignis Broun, 1921
  • Mitophyllus irroratus Parry, 1842
  • Mitophyllus macrocerus (Broun, 1886)
  • Mitophyllus parrianus Westwood, 1863
  • Mitophyllus reflexus Broun, 1908
Tribe Nicagini
Genus Nicagus LeConte, 1861
  • Nicagus obscurus (LeConte, 1848)
  • Nicagus occultus Paulsen & Smith, 2005
  • Nicagus japonicus Nagel, 1928
Genus Holloceratognathus Nikolajev, 1998
  • Holloceratognathus cylindricus (Broun, 1895)
  • Holloceratognathus helotoides (Thomson, 1862)
  • Holloceratognathus passaliformis (Holloway, 1962)

Subfamily Lampriminae
Genus Dendroblax White, 1846
  • Dendroblax earlianus White, 1846
Genus Homolamprima
  • Homolamprima crenulata (MacLeay, 1885)
Genus Lamprima
  • Lamprima adolphinae
    Lamprima adolphinae
    Lamprima adolphinae is a beetle of the Family Lucanidae.-External links:*...

    (Gestro, 1875)
  • Lamprima aurata
Genus Phalacrognathus
  • Phalacrognathus muelleri
    Phalacrognathus muelleri
    Phalacrognathus muelleri MacLeay, 1885 is a beetle of the Family Lucanidae.- List of subspecies :* Phalacrognathus muelleri muelleri Origin: Queensland, Australia* Phalacrognathus muelleri fuscomicans Origin: New Guinea-External links:*...

    MacLeay, 1885
Genus Streptocerus Dejean, 1833
  • Streptocerus speciosus Fairmaire, 1850

Subfamily Lucaninae
Lucaninae
Lucaninae is the largest subfamily of the stag beetles .Characteristics include partial to complete division of the eyes by a canthus, geniculate antennae, and distinctly separated coxae...

 Latreille
Pierre André Latreille
Pierre André Latreille was a French zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained his freedom after recognising a rare species he found in the prison, Necrobia ruficollis...

, 1804
Tribe Aegini
Genus Aegus MacLeay
Alexander Macleay
Hon. Alexander Macleay MLC FLS FRS was a leading member of the Linnean Society and a fellow of the Royal Society.Macleay was born on Ross-shire, Scotland, eldest son of William Macleay, provost of Wick...

, 1819
Tribe Allotopini
Genus Allotopus
  • Allotopus moellenkampi (Fruhstorfer, 1894)
  • Allotopus rosenbergi (Vollenhoven, 1872)
Genus Mesotopus
Tribe Chiasognathini
Genus Casignetus
Genus Chiasognathus
Chiasognathus
Chiasognathus is a genus of stag beetles found in Argentina and Chile. It includes seven species:*Chiasognathus beneshi Lacroix, 1978*Chiasognathus grantii Stephens, 1832*Chiasognathus impubis Parry, 1870...

Stephens
James Francis Stephens
James Francis Stephens was an English entomologist.-Biography:Stephens was born in Shoreham-by-Sea and studied at Christ's Hospital.He was employed in the Admiralty office, Somerset House, from 1807 to 1845...

, 1831
  • Chiasognathus beneshi Lacroix, 1978
  • Chiasognathus grantii Stephens
    James Francis Stephens
    James Francis Stephens was an English entomologist.-Biography:Stephens was born in Shoreham-by-Sea and studied at Christ's Hospital.He was employed in the Admiralty office, Somerset House, from 1807 to 1845...

    , 1831
  • Chiasognathus jousselini Reiche, 1850
  • Chiasognathus latreillei Solier, 1851
  • Chiasognathus mniszechii Kiesche 1919
Genus Protognathinus
Genus Sphaenognathus
Tribe Cladognathini
Genus Aphanognathus
  • Aphanognathus laterotarsoides (Houlbert, 1915)
Genus Capreolucanus
  • Capreolucanus sicardi Didier, 1928
Genus Cladophyllus
Genus Eligmodontus
Genus Gonometopus
Genus Macrodorcas Motschulsky, 1862
  • Macrodorcas adachii
  • Macrodorcas amamiana
  • Macrodorcas barbata
  • Macrodorcas bisignata
  • Macrodorcas bomansi
  • Macrodorcas davidi
  • Macrodorcas dierli
  • Macrodorcas elsiledis
  • Macrodorcas fulvonotata
  • Macrodorcas humilis
  • Macrodorcas intricata
  • Macrodorcas kobayashii
  • Macrodorcas koyamai
  • Macrodorcas melliana
  • Macrodorcas mochizukii
  • Macrodorcas moellenkampi
  • Macrodorcas montivaga
  • Macrodorcas pieli
  • Macrodorcas pseudaxis
  • Macrodorcas recta Motschulsky, 1858
  • Macrodorcas ruficrus
  • Macrodorcas songiana
  • Macrodorcas striatipennis Motschulsky, 1862
  • Macrodorcas ursulae
  • Macrodorcas vernicata
  • Macrodorcas yaeyamaensis
Genus Palaeognathus
Genus Prismognathus Motschulsky, 1860
  • Prismognathus alessandrae
  • Prismognathus angularis C._O._Waterhouse, 1874
  • Prismognathus branczicki
  • Prismognathus cheni
  • Prismognathus dauricus Motschulsky, 1860
  • Prismognathus davidis
  • Prismognathus delislei
  • Prismognathus formosanus
  • Prismognathus klapperichi
  • Prismognathus lucidus
  • Prismognathus morimotoi
  • Prismognathus nigerrimus
  • Prismognathus nigricolor
  • Prismognathus piluensis
  • Prismognathus platycephalus
  • Prismognathus ruficephalus
  • Prismognathus sinensis
  • Prismognathus subnitens
  • Prismognathus tokui
Genus Prosopocoilus Westwood
John Obadiah Westwood
John Obadiah Westwood was an English entomologist and archaeologist also noted for his artistic talents.Born in Sheffield, he studied to be a lawyer but abandoned that for his scientific interests....

, 1845
Genus Pseudorhaetus
Genus Rhaetulus
Genus Rhaetus
  • Rhaetus westwoodi (Parry, 1862)
Genus Tetrarthrius
Genus Weinreichius
  • Weinreichius perroti Lacroix, 1978
Tribe Colophonini
Genus Colophon
Colophon (genus)
Colophon is a small genus of beetle in the "stag beetle" family Lucanidae.These beetles are flightless, and are endemic to South Africa, each restricted to its own mountain range or peak within a range...

Tribe Cyclommatini
Genus Cyclommatus
Cyclommatus
Cyclommatus is a rare genus of the stag beetle. They are about 105mm big....

Parry
Charles Christopher Parry
thumb|150px|Parry circa 1875Charles Christopher Parry was a British-American botanist and mountaineer....

, 1863
Tribe Dendeziini
Genus Dendezia
Genus Oonotus Parry
Charles Christopher Parry
thumb|150px|Parry circa 1875Charles Christopher Parry was a British-American botanist and mountaineer....

, 1864
Genus Xiphodontus
Tribe Dorcini
Genus Bartolozziolucanus
Genus Cantharolethrus
Genus Cyclommatus
Cyclommatus
Cyclommatus is a rare genus of the stag beetle. They are about 105mm big....

Genus Dorcasoides
Genus Dorcus
Dorcus
Dorcus is a genus of the family Lucanidae of stag beetles. Of the 30-odd species, most occur in Asia and India; two are found in southern Europe, and three species are from North America...

MacLeay
Alexander Macleay
Hon. Alexander Macleay MLC FLS FRS was a leading member of the Linnean Society and a fellow of the Royal Society.Macleay was born on Ross-shire, Scotland, eldest son of William Macleay, provost of Wick...

, 1819
  • Dorcus akahorii
  • Dorcus alcides
  • Dorcus alexisi
  • Dorcus amamianus
  • Dorcus antaeus
  • Dorcus anteus
  • Dorcus arfakianus
  • Dorcus binodulosus
  • Dorcus brevis
  • Dorcus bucephalus
  • Dorcus carinulatus
  • Dorcus consentaneus
  • Dorcus curvidens
    Dorcus curvidens
    Dorcus curvidens is a beetle of the family Lucanidae.It is also known as the Ookuwagata beetle in Japan....

  • Dorcus cylindricus
  • Dorcus davidis
  • Dorcus emikoae
  • Dorcus eurycephalus
  • Dorcus formosanus
  • Dorcus gracilicornis
  • Dorcus garndis
  • Dorcus hopei
  • Dorcus hyperion
  • Dorcus japonicus
  • Dorcus kyanrauensis
  • Dorcus metacostatus
  • Dorcus miwai
    Dorcus miwai
    Dorcus miwai is a beetle of the Family Lucanidae....

  • Dorcus musimon
  • Dorcus parallelus
  • Dorcus parallelipipedus
    Dorcus parallelipipedus
    Dorcus parallelipipedus, the lesser stag beetle, is a species of stag beetle found in Europe.Both sexes resemble the female stag beetle , though they are a uniformly blackish colour rather than having the chestnut brown wing covers of the larger species...

    Linnaeus
    Carolus Linnaeus
    Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

    , 1758
  • Dorcus parryi
  • Dorcus parvulus
  • Dorcus peyronis
  • Dorcus prochazkai
  • Dorcus ratiocinativus
  • Dorcus rectus
  • Dorcus reichei
  • Dorcus rudis
  • Dorcus sawaii
  • Dorcus schenklingi
  • Dorcus sewertzowi
  • Dorcus suturalis
  • Dorcus taiwanicus
  • Dorcus tanakai
  • Dorcus taurus
  • Dorcus tenuecostatus
  • Dorcus titanus
    Dorcus titanus
    Dorcus titanus, also known as the giant stag beetle, is a beetle of the Family Lucanidae.- List of subspecies :* Dorcus titanus castanicolor* Dorcus titanus daitoensis* Dorcus titanus elegans* Dorcus titanus hachijoensis...

  • Dorcus tormosanus
  • Dorcus ursulus
  • Dorcus vavrai
  • Dorcus velutinus
  • Dorcus vicinus
  • Dorcus yamadai
Genus Homoderus
Genus Leptinopterus
Genus Prosopocoilus
  • Prosopocoilus astacoides
    Prosopocoilus astacoides
    Prosopocoilus astacoides is a beetle of the Family Lucanidae.-External links:*...

  • Prosopocoilus biplagiatus
    Prosopocoilus biplagiatus
    Prosopocoilus biplagiatus is a beetle of the Family Lucanidae.-External links:*...

  • Prosopocoilus bison
    Prosopocoilus bison
    Prosopocoilus bison is a beetle of the Family Lucanidae....

  • Prosopocoilus giraffa
  • Prosopocoilus inclinatus
    Prosopocoilus inclinatus
    Prosopocoilus inclinatus is a beetle of the Family Lucanidae....

  • Prosopocoilus savagei
    Prosopocoilus savagei
    Prosopocoilus savagei is a beetle of the Family Lucanidae....

Genus Rhaetulus
  • Rhaetulus crenatus
    Rhaetulus crenatus
    Rhaetulus crenatus is a beetle of the Family Lucanidae.-External links:*...

Genus Serrognathus
Genus Pycnosiphorus
Genus Sclerostomus
Tribe Figulini
Genus Cardanus
Genus Dinonigidius
Genus Figulus
Genus Ganelius
Genus Nigidionus
Genus Nigidius
Genus Novonigidius
Genus Penichrolucanus
Tribe Lissapterini
Genus Bomansius
Genus Dorculus
Genus Geodorcus
Genus Hoplogonus
Genus Lissapterus
Genus Lissotes
Genus Paralissotes
Genus Pseudodorcus
Tribe Lucanini
Genus Hexarthrius
Hexarthrius
Hexarthrius is a genus of large stag beetles. It includes the following species:*Hexarthrius aduncus*Hexarthrius andreasi*Hexarthrius bowringii*Hexarthrius buquettii*Hexarthrius davisoni*Hexarthrius forsteri...

Genus Lucanus
Lucanus (genus)
Lucanus is a genus of stag beetles .-Species:not a complete list*Lucanus capreolus*Lucanus cervus*Lucanus elaphus*Lucanus formosanus*Lucanus maculifemoratus*Lucanus mazama*Lucanus placidus...

  • Lucanus capreolus
    Lucanus capreolus
    Lucanus capreolus is a beetle of the Family Lucanidae. The species name "capreolus" is derived from Latin, meaning "Roe Deer"...

  • Lucanus cervus
    Lucanus cervus
    Lucanus cervus is the best-known species of stag beetle , and is sometimes referred to simply as the stag beetle. It lives in holes in old trees and dead trunks, in the forest as well as in groves. Forest management, in eliminating old trees and deadwood, eliminates at the same time the habitat and...

  • Lucanus elaphus
    Lucanus elaphus
    The giant stag beetle is a beetle of the family Lucanidae....

  • Lucanus formosanus
    Lucanus formosanus
    Lucanus formosanus is a stag beetle which is endemic to Taiwan, and grows to a length of ....

  • Lucanus maculifemoratus
    Lucanus maculifemoratus
    Lucanus maculifemoratus is a beetle of the Family Lucanidae....

  • Lucanus mazama
    Lucanus mazama
    Lucanus mazama is a beetle of the family Lucanidae....

  • Lucanus placidus
    Lucanus placidus
    Lucanus placidus is a beetle of the Family Lucanidae....

  • Lucanus swinhoei
    Lucanus swinhoei
    Lucanus swinhoei is a beetle of the Family Lucanidae....

  • Lucanus tetraodon
    Lucanus tetraodon
    Lucanus tetraodon is a beetle of the Family Lucanidae....

Genus Pseudolucanus
  • Pseudolucanus barbarossa (Fabricius
    Fabricius
    Fabricius may refer to:*people from the Ancient Roman gens Fabricia:*Gaius Fabricius Luscinus, the first of the Fabricii to move to Rome* Carel Fabricius , painter...

    , 1801)
Tribe Odontolabini
Genus Calcodes
Genus Neolucanus
Genus Odontolabis
Odontolabis
Odontolabis is a genus of beetles belonging to the Lucanidae family.-Description:The species of the genus Odontolabis are large , oblong and shiny. Usually males have more or less enlarged head and jaws. The head and the pronotum usually are black, while elytra are often more or less brownish or...

  • Odontolabis castelnaudi
    Odontolabis castelnaudi
    Odontolabis castelnaudi is a beetle of the Family Lucanidae-References:* Lacroix, J.-P., 1984 - The Beetles of the World, volume 4, Odontolabini I - Genera Chalcodes, Odontolabis, Heterochtes. This is a very good revision of the genus Odontolabis...

  • Odontolabis cuvera
    Odontolabis cuvera
    Odontolabis cuvera is a beetle of the Family Lucanidae, stag beetles. The males have three morphological forms with variations in the size of their mandibles.This beetle is native to southeast Asia.- List of subspecies :...

  • Odontolabis femoralis
    Odontolabis femoralis
    Odontolabis femoralis is a beetle of the Family Lucanidae. It lives in Indonesia.- List of subspecies :* Odontolabis femoralis femoralis Waterhouse, 1887* Odontolabis femoralis kinabaluensis Möllenkamp, 1904...

  • Odontolabis siva
    Odontolabis siva
    Odontolabis siva is a beetle of the Family Lucanidae.- Monograph :* Lacroix, J.-P., 1984 - The Beetles of the World, volume 4, Odontolabini I - Genera Chalcodes, Odontolabis, Heterochtes.-External links:*...

Tribe Platycerini
Genus Platyceroides Benesh, 1946
Genus Platyceropsis Benesh, 1946
  • Platyceropsis keeni (Casey, 1895)
Genus Platycerus Geoffroy, 1762
Tribe Rhyssonotini
Genus Cacostomus
  • Cacostomus floralis (Lea, 1914)
  • Cacostomus squamosus
    Cacostomus squamosus
    Cacostomus squamosus is a beetle of the Lucanidae family. It lives in Australia.-External links:* Australian Faunal Directory...

    Newman, 1840
  • Cacostomus subvittata (Moore, 1994)
Genus Rhyssonotus
Rhyssonotus
Rhyssonotus is a genus of stag beetles .- References :* G. Cassis, W.W.K. Houston, T.A. Weir & B.P. Moore . Updated by A.A. Calder . Coleoptera: Scarabaeoida.. Australian Biological Resources Study, Canberra. Viewed 22 November 2007.-External links:* *...


Subfamily Syndesinae
Tribe Ceruchini
Genus Ceruchus MacLeay, 1819
  • Ceruchus piceus (Weber
    Friedrich Weber (entomologist)
    Friedrich Weber was a German entomologist. He was a pupil of Johan Christian Fabricius , and wrote in 1795 at the age of 14 and in 1801...

    , 1801)
  • Ceruchus punctatus LeConte, 1869
  • Ceruchus striatus LeConte, 1859
  • Ceruchus yangi Huang, Imura & Chen, 2011
Tribe Sinodendronini
Genus Sinodendron Hellwig in Schneider, 1792
  • Sinodendron cylindricum (Linnaeus, 1758)
  • Sinodendron rugosum
    Sinodendron rugosum
    Sinodendron rugosum is a member of the family Lucanidae, the stag beetles. It is commonly referred to as the rugose stag beetle, and is the only known member of the genus Sinodendron to occur in western North America.-Description:...

    Mannerheim, 1843
Tribe Syndesini
Genus Psilodon Perty, 1830
  • Psilodon aequinoctiale (Buquet, 1840)
  • Psilodon gilberti Boucher, 1993
  • Psilodon schuberti Perty, 1830
  • Psilodon seguyi (Didier, 1929)
  • Psilodon xerophilicum Martinez & Reyes-Castillo, 1985
Genus Syndesus MacLeay, 1819
  • Syndesus cancellatus Montrouzier, 1860
  • Syndesus cornutus (Fabricius, 1801)
  • Syndesus macleayi Boileau, 1905
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