Sphaerechinus granularis is a species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of sea urchin

Sea urchin

Sea urchins or urchins are small, spiny, globular animals which, with their close kin, such as sand dollars, constitute the class Echinoidea of the echinoderm phylum. They inhabit all oceans. Their shell, or "test", is round and spiny, typically from across. Common colors include black and dull...

 in the family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Toxopneustidae

Toxopneustidae

Toxopneustidae is a family of globular sea urchins in the class Echinoidea.-Characteristics:All Camarodonts have imperforate tubercles and compound ambulacral plates. In addition, the Toxopneustids are characterised by the peristome, or opening through the test, having a sharp margin with the...

, commonly known as the purple sea urchin. Its range includes the Mediterranean Sea

Mediterranean Sea

The Mediterranean Sea is a sea connected to the Atlantic Ocean surrounded by the Mediterranean region and almost completely enclosed by land: on the north by Anatolia and Europe, on the south by North Africa, and on the east by the Levant...

 and eastern Atlantic Ocean

Atlantic Ocean

The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

.

Description

S. granularis is a large sea urchin, somewhat flattened dorsally and growing to fifteen centimetres in diameter. There are two distinct colour forms. The test

Test (biology)

A test is a term used to refer to the shell of sea urchins, and also the shell of certain microorganisms, such as testate foraminifera and testate amoebae....

 is purple in both but one has purple spines and the other white. The spines are short and blunt, all the same length, and arranged neatly in rows.

Distribution and habitat

S. granularis is found in the Mediterranean Sea and the eastern Atlantic Ocean, from the Channel Islands

Channel Islands

The Channel Islands are an archipelago of British Crown Dependencies in the English Channel, off the French coast of Normandy. They include two separate bailiwicks: the Bailiwick of Guernsey and the Bailiwick of Jersey...

 south to Cape Verde

Cape Verde

The Republic of Cape Verde is an island country, spanning an archipelago of 10 islands located in the central Atlantic Ocean, 570 kilometres off the coast of Western Africa...

 and the Gulf of Guinea

Gulf of Guinea

The Gulf of Guinea is the northeasternmost part of the tropical Atlantic Ocean between Cape Lopez in Gabon, north and west to Cape Palmas in Liberia. The intersection of the Equator and Prime Meridian is in the gulf....

. It favours sheltered locations and lives on rocks covered with seaweed

Seaweed

Seaweed is a loose, colloquial term encompassing macroscopic, multicellular, benthic marine algae. The term includes some members of the red, brown and green algae...

 or gravelly substrate

Substrate

Substrate may mean:*Substrate , Natural stone, masonry surface, ceramic and porcelain tiles*Substrate , the material used in the bottom of an aquarium*Substrate , the material used in the bottom of a vivarium or terrarium...

s. It is usually found in the neritic zone

Neritic zone

The neritic zone, also called coastal waters, the coastal ocean or the sublittoral zone, is the part of the ocean extending from the low tide mark to the edge of the continental shelf, with a relatively shallow depth extending to about 200 meters...

 down to about thirty metres but occasionally down to a hundred metres in more exposed locations. It is also found in meadows of the seagrass

Seagrass

Seagrasses are flowering plants from one of four plant families , all in the order Alismatales , which grow in marine, fully saline environments.-Ecology:...

 Posidonia oceanica

Posidonia oceanica

Posidonia oceanica is a seagrass species that is endemic to the Mediterranean Sea. It forms large underwater meadows that are an important part of the ecosystem. The fruit is free floating and known in Italy as 'the olive of the sea'...

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Biology

S. granularis often covers itself with morsels of algae and shell fragments, which are held in place by the tube feet

Tube feet

Tube feet are the many small tubular projections found most famously on the oral face of a sea star's arms, but are characteristic of the water vascular system of the echinoderm phylum which also includes sea urchins, sand dollars and sea cucumbers and many other sea creatures.Tube feet function in...

 and by the claw-like structures known as pedicellaria. It grazes on algae

Algae

Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

, especially encrusting coralline algae, seagrass blades and their epiphytic

Epiphyte

An epiphyte is a plant that grows upon another plant non-parasitically or sometimes upon some other object , derives its moisture and nutrients from the air and rain and sometimes from debris accumulating around it, and is found in the temperate zone and in the...

 organisms and detritus

Detritus

Detritus is a biological term used to describe dead or waste organic material.Detritus may also refer to:* Detritus , a geological term used to describe the particles of rock produced by weathering...

.

Spawning takes place at any time of year but the peak period is spring and early summer. Eggs

Egg (biology)

An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

 and sperm

Sperm

The term sperm is derived from the Greek word sperma and refers to the male reproductive cells. In the types of sexual reproduction known as anisogamy and oogamy, there is a marked difference in the size of the gametes with the smaller one being termed the "male" or sperm cell...

 are liberated into the water column, where fertilisation takes place. The larva

Larva

A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e are plankton

Plankton

Plankton are any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. That is, plankton are defined by their ecological niche rather than phylogenetic or taxonomic classification...

ic. After several moults, the echinopluteus larva settles and undergoes metamorphosis

Metamorphosis

Metamorphosis is a biological process by which an animal physically develops after birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation...

 before developing into a juvenile.

Ecology

In Tunisia, S. granularis is found living in association with two other species of sea urchins, Centrostephanus longispinus and Paracentrotus lividus

Paracentrotus lividus

Paracentrotus lividus is a species of sea urchin in the family Parechinidae commonly known as the purple sea urchin. It is the type species of the genus and occurs in the Mediterranean Sea and eastern Atlantic Ocean.-Description:...

. It is preyed

Predation

In ecology, predation describes a biological interaction where a predator feeds on its prey . Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of its prey and the eventual absorption of the prey's tissue through consumption...

 upon by the starfish Marthasterias glacialis and Luidia ciliaris

Luidia ciliaris

Luidia ciliaris or the seven armed starfish is a species of starfish in the family Luidiidae. It is found in the eastern Atlantic Ocean and the Mediterranean Sea.-Description:...

.

Use as food

The gonad

Gonad

The gonad is the organ that makes gametes. The gonads in males are the testes and the gonads in females are the ovaries. The product, gametes, are haploid germ cells. For example, spermatozoon and egg cells are gametes...

s of S. granularis are considered a delicacy in Italy

Italy

Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

 and Provence

Provence

Provence ; Provençal: Provença in classical norm or Prouvènço in Mistralian norm) is a region of south eastern France on the Mediterranean adjacent to Italy. It is part of the administrative région of Provence-Alpes-Côte d'Azur...

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