Salvia involucrata
Encyclopedia
Salvia involucrata is a herbaceous perennial belonging to the family Lamiaceae
Lamiaceae
The mints, taxonomically known as Lamiaceae or Labiatae, are a family of flowering plants. They have traditionally been considered closely related to Verbenaceae, but in the 1990s, phylogenetic studies suggested that many genera classified in Verbenaceae belong instead in Lamiaceae...

. It is native to the Mexican
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 states of Puebla
Puebla
Puebla officially Estado Libre y Soberano de Puebla is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 217 municipalities and its capital city is Puebla....

, Tamaulipas
Tamaulipas
Tamaulipas officially Estado Libre y Soberano de Tamaulipas is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 43 municipalities and its capital city is Ciudad Victoria. The capital city was named after Guadalupe Victoria, the...

, and Veracruz
Veracruz
Veracruz, formally Veracruz de Ignacio de la Llave officially Estado Libre y Soberano de Veracruz de Ignacio de la Llave , is one of the 31 states that, along with the Federal District, comprise the 32 federative entities of Mexico. It is divided in 212 municipalities and its capital city is...

, growing in shady places such as the edge of forests. Its specific epithet, "involucrata", refers to the prominent flower bract
Bract
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis, or cone scale. Bracts are often different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture...

s, which are large and colorful.

Description

Salvia involucrata grows five feet or taller before it starts blooming in late summer. The plant's flowers and bracts are a reddish, beetroot color. The bracts occur in pairs which envelop three flowers each, falling away as the flowers expand. The plant's leaves are small, flat mid-green, slightly cordate-shaped. Unusually, the leaves' petiole
Petiole (botany)
In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...

s and vein
Vein
In the circulatory system, veins are blood vessels that carry blood towards the heart. Most veins carry deoxygenated blood from the tissues back to the heart; exceptions are the pulmonary and umbilical veins, both of which carry oxygenated blood to the heart...

s share the flowers' beetroot-reddish hue. Genetically, the species has eleven (11) tetraploidal chromosome
Chromosome
A chromosome is an organized structure of DNA and protein found in cells. It is a single piece of coiled DNA containing many genes, regulatory elements and other nucleotide sequences. Chromosomes also contain DNA-bound proteins, which serve to package the DNA and control its functions.Chromosomes...

s.

Several unique cultivar
Cultivar
A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

s have been collected in the wild and named. Two that are common in France and Britain are 'Bethelii', which was introduced in 1881 for its compact habit and large ovoid leaves, and 'Deschamsiana', which was chosen in 1869 for the bright rose color of its inflated flowers. Cultivars that have found popularity in the U.S. include 'Hidalgo', 'El Butano', and 'El Cielo'—each named after the place in Mexico where it was collected.

Salvia involucrata breeds freely with other Salvia
Salvia
Salvia is the largest genus of plants in the mint family, Lamiaceae, with approximately 700-900 species of shrubs, herbaceous perennials, and annuals. It is one of several genera commonly referred to as sage. When used without modifiers, sage generally refers to Salvia officinalis ; however, it is...

species, resulting in many hybrids at University of California Botanical Garden
University of California Botanical Garden
The University of California Botanical Garden is a 34 acre botanical garden located on the University of California, Berkeley campus. The Garden is in the campus's Strawberry Canyon which overlooks the San Francisco Bay...

 that show hybrid vigor. Some of these hybrid plants are known to grow up to six feet high, with a longer blooming period. 'Mulberry Jam', a smaller hybrid with upright growth, stronger stems, and continuous blooming from summer to frost was introduced in 1995 by Betsy Clebsch.

As a garden plant, it prefers good drainage, half to three-quarter a day of sun, humus enriched soil, and deep watering once a week. Propagation is by division or cuttings, which can be rooted in late summer or early autumn. The plant will grow back from the ground after light freezes. In early spring, it should be pruned back to active nodes a few inches from the crown.

In medicine

Historically, S. involucrata along with several other members of its genus have been used as a memory-enhancer in traditional herbal medicine. Scientific studies seeking to verify this have shown that extracts of S. involucrata (along with other Salvia species) contained significant amounts of cerebral cortex
Cerebral cortex
The cerebral cortex is a sheet of neural tissue that is outermost to the cerebrum of the mammalian brain. It plays a key role in memory, attention, perceptual awareness, thought, language, and consciousness. It is constituted of up to six horizontal layers, each of which has a different...

-relevant compounds, including with those with binding affinity to acetylcholine receptors. The species has also been used at least once in the past as an indicator of carbon dioxide
Carbon dioxide
Carbon dioxide is a naturally occurring chemical compound composed of two oxygen atoms covalently bonded to a single carbon atom...

 metabolism. In a 1919 study, Salvia involucrata was used to determine the effects of ether
Ether
Ethers are a class of organic compounds that contain an ether group — an oxygen atom connected to two alkyl or aryl groups — of general formula R–O–R'. A typical example is the solvent and anesthetic diethyl ether, commonly referred to simply as "ether"...

on the rate of CO2 respiration on dying cells. The study found that petals of S. involucrata immersed in ether consumed more oxygen as the cells expired.
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